Lichenometric dating (lichenometry) and the biology of the lichen genus rhizocarpon:challenges and future directions

Lichenometric dating (lichenometry) involves the use of lichen measurements to estimate the age of exposure of various substrata. Because of low radial growth rates and considerable longevity, species of the crustose lichen genus Rhizocarpon have been the most useful in lichenometry. The primary assumption of lichenometry is that colonization, growth and mortality of Rhizocarpon are similar on surfaces of known and unknown age so that the largest thalli present on the respective faces are of comparable age. This review describes the current state of knowledge regarding the biology of Rhizocarpon and considers two main questions: (1) to what extent does existing knowledge support this assumption; and (2) what further biological observations would be useful both to test its validity and to improve the accuracy of lichenometric dates? A review of the Rhizocarpon literature identified gaps in knowledge regarding early development, the growth rate/size curve, mortality, regeneration, competitive effects, colonization, and succession on rock surfaces. The data suggest that these processes may not be comparable on different rock surfaces, especially in regions where growth rates and thallus turnover are high. In addition, several variables could differ between rock surfaces and influence maximum thallus size, including rate and timing of colonization, radial growth rates, environmental differences, thallus fusion, allelopathy, thallus mortality, colonization and competition. Comparative measurements of these variables on surfaces of known and unknown age may help to determine whether the basic assumptions of lichenometry are valid. Ultimately, it may be possible to take these differences into account when interpreting estimated dates

Governing multisectoral action for health in low- and middle-income countries

Kumanan Rasanathan and colleagues argue that the potential of multisectoral collaboration for improving health remains untapped in many low- and middle-income countries

Phegopteris connectilis

Pteridophyte

Physical Characterization, Microstructural Evaluation, and Condition Assessment of Ancient Ahlat Tombstones in the Seljukian Cemetery of Ahlat (Turkey)

A typical Seljukian town of Ahlat, located between the north-western shores of the Lake Van and the Nemrud and Suphan volcanoes of the Eastern Turkey is hosting rich and colorful cultural heritage sites. Among these, famous Seljukian Cemetery is a major archeological district with monumental tombstones (stelae). Excessive deterioration, erosion and lichen colonization can be observed in these cultural artifacts. The main objective of this study is the investigation of stones' physical characterization and evaluation of the microstructural features. A degradation model was outlined starting with the capillary water uptake from the bottom section and lichen colonization starting from the top and covering these tombstones upto their mid sections. This article provides some information about the historical town of Ahlat and its tombstones. Some physical and microstructural characterization of the gravestones and the results of chemical and physical analysis are also presented along with some recommendations