The origins and spread of domestic horses from the Western Eurasian steppes

Analysis of 273 ancient horse genomes reveals that modern domestic horses originated in the Western Eurasian steppes, especially the lower Volga-Don region.Domestication of horses fundamentally transformed long-range mobility and warfare(1). However, modern domesticated breeds do not descend from the earliest domestic horse lineage associated with archaeological evidence of bridling, milking and corralling(2-4) at Botai, Central Asia around 3500 bc(3). Other longstanding candidate regions for horse domestication, such as Iberia(5) and Anatolia(6), have also recently been challenged. Thus, the genetic, geographic and temporal origins of modern domestic horses have remained unknown. Here we pinpoint the Western Eurasian steppes, especially the lower Volga-Don region, as the homeland of modern domestic horses. Furthermore, we map the population changes accompanying domestication from 273 ancient horse genomes. This reveals that modern domestic horses ultimately replaced almost all other local populations as they expanded rapidly across Eurasia from about 2000 bc, synchronously with equestrian material culture, including Sintashta spoke-wheeled chariots. We find that equestrianism involved strong selection for critical locomotor and behavioural adaptations at the GSDMC and ZFPM1 genes. Our results reject the commonly held association(7) between horseback riding and the massive expansion of Yamnaya steppe pastoralists into Europe around 3000 bc(8,9) driving the spread of Indo-European languages(10). This contrasts with the scenario in Asia where Indo-Iranian languages, chariots and horses spread together, following the early second millennium bc Sintashta culture(11,12).Descriptive and Comparative Linguistic

Reconstructing Age Distribution, Season of Capture and Growth Rate of Fish from Archaeological Sites Based on Otoliths and Vertebrae

The growth increments of otoliths and vertebrae of plaice (Pleuronectes platessa) derived from a 15th century single depositional event at Raversijde (Belgium) are analysed with the aim of reconstructing (a) the age distribution of the population, (b) the season of capture, and (c) the growth rate. Otoliths and vertebrae give slightly different age distributions but it is possible to arrive at similar seasonality estimations in both structures when information from the literature and our own date from monthly captures of plaice from the North Sea are taken into account. These modern data show that the timing of annulus formation in otoliths and vertebrae is more or less similar. Back-calculations on vertebrae and otoliths yield similar growth curves. The age distribution, the edge condition of both vertebrae and otoliths, and the growth rate obtained on the material from Raversijde all show that the plaice from the studied assemblage were captured during spring in the southern part of the North Sea. Vertebrae are commonly preserved in archaeological sites whereas otoliths rarely survive. Although they are more difficult to read than otoliths, vertebrae of plaice can be used for growth increment analyses, and the growth rates obtained from vertebrae from archaeological sites can, therefore, be compared in the future to growth data from modern otoliths studied in sea fisheries research. Archaeozoological material predating industrialized fishing since the 19th century can hence serve as a reference in the study of the compensatory response of commercially important species to heavy exploitation. Copyrigh

History of the reindeer (Rangifer tarandus) in the eastern Baltic region and its implications for the origin and immigration routes of the recent northern European wild reindeer populations

A total of 45 subfossil reindeer (Rangifer tarandus) antlers and bones - artefacts excluded - have been found over the years in the Baltic countries of Estonia, Latvia and Lithuania. The relatively high number of specimens suggests a stable residence of the species in the eastern Baltic region. For the first time, 12 of these finds were radiocarbon-dated. The ages of the samples range between 12085 and 9970 C-14 yr BP (14180-11280 cal. yr BP), and cover the Lateglacial and early Holocene, a time period during which climatic conditions shifted from periglacial to temperate. The dates suggest a rapid colonization of the area during the deglaciation period and a local extinction around the Pleistocene-Holocene boundary. The results of the study do not support the theory that the recent wild reindeer populations of northern Europe had their origin in the Late Weichselian reindeer populations of the eastern Baltic region

An Arctic seal in temperate waters: history of the ringed seal (Pusa hispida) in the Baltic Sea and its adaptation to the changing environment

The ringed seal (Pusa hispida) is an early immigrant in the Baltic Basin and has since its arrival experienced substantial changes in the climate, salinity and productivity of the Basin. In this paper, we discuss the dispersal and distribution of the ringed seal during different stages of the Baltic Sea in relation to past and ongoing environmental changes. Subfossil ringed seal remains around the Baltic Sea and the Danish Straits were radiocarbon dated in order to map the distribution of the species in different time periods. The δ13C data were used in evaluating the changes in the marine character of the Baltic Basin. The sequence of the dates indicates a continuous presence of the species in the Baltic Basin. The earliest ringed seal finds come from the Skagerrak/Kattegat area (Denmark, Swedish west coast) and date to the full glacial period and Baltic Ice Lake. In the Baltic Basin, the species appears in the subfossil record during the Ancylus period, but the main part of the remains date to the Littorina stage. During the Littorina stage, the distribution of the species was at least periodically wider than today, covering also southern parts of the Baltic. The presence of breeding populations in southern parts of the Baltic during the Holocene Thermal Maximum (HTM) indicates that the winters were at least periodically cold enough for winter ice. The changes in the marine influence in the Baltic Basin can be seen in the seal collagen δ13C values, which serve as a proxy for qualitative changes in water mass salinity

Detecting the medieval cod trade: a new method and first results

This paper explores the potential of stable isotope analysis to identify the approximate region of catch of cod by analysing bones from medieval settlements in northern and western Europe. It measures the d13C and d15N values of cod bone collagen from medieval control samples collected from sites around Arctic Norway, the North Sea, the Kattegat and the Baltic Sea. These data were considered likely to differ by region due to, for example, variation in the length of the food chain, water temperature and salinity. We find that geographical structuring is indeed evident, making it possible to identify bones from cod caught in distant waters. These results provide a new methodology for studying the growth of long-range trade in dried cod and the related expansion of fishing effort—important aspects of the development of commercialisation in medieval Europe. As a first test of the method, we analyse three collections of cod bones tentatively interpreted as imported dried fish based on a priori zooarchaeological criteria. The results tentatively suggest that cod were being transported or traded over very long distances since the end of the first millennium AD