Ground beetles and rove beetles be associated with temporary ponds in England

To date, research on the ecology and conservation of wetland invertebrates has concentrated overwhelmingly on fully aquatic organisms. Many of these spend part of their life-cycle in adjacent terrestrial habitats, either as pupae (water beetles) or as adults (mayflies, dragonflies, stoneflies, caddisflies and Diptera or true-flies). However, wetland specialist species also occur among several families of terrestrial insects (Williams & Feltmate 1992) that complete their whole life-cycle in the riparian zone or on emergent vegetation. There are 441 terrestrial invertebrate species which characteristically occur in riparian habitats along British rivers. Most of these species belong to two families of predatory beetles: the ground beetles (Carabidae) and the rove beetles (Staphylinidae). This paper describes the diversity of ground and rove beetles around ponds, summarises life-histories, hibernation strategies, and morphological and behavioural adaptions

The semi-aquatic habitats of terrestial Coleoptera in a lowland river floodplain

PhD Thesis281 species of terrestrial ground-living beetles were recorded from 69 riparian and wetland sites in the floodplain of the lowland River Soar, England. Differences in species composition between pitfall trapped and timed hand-collected samples were smaller than those attributable to environmental and seasonal factors. Detrended Correspondence Analysis consistently ranked all sites against seasonal variations between April and June and floodplain sites against annual variations. DCA axis 1 scores were slightly better correlated with important environmental variables at the ecohabitat (<5Om) scale rather than the microhabitat scale. Canonical Correspondence Analysis detected assemblage responses to flooding disturbance and grazing pressure along the main channel as well as to water level stability in the floodplain. A conceptual model of floodplain land-use and river management postulated a dynamic equilibrium between flooding disturbances and vegetational succession, producing geomorphic and vegetational structures which serve as semi-aquatic habitats for terrestrial beetle assemblages with appropriate species traits. Impoundment for navigation affects assemblages by modifying the severity of flooding disturbance. The effects of grazing pressure resemble flooding disturbance. The short-term (<5yr) impact of bank regrading was explained by differences in severity, predictability and frequency compared to the beetles' generation length. Evenness and species richness were affected only by flooding and grazing disturbance. This response was not predicted by the intermediate disturbance hypothesis because the frequencies of flooding and grazing disturbances in the Soar valley are not appropriate to the hypothesis, which more closely relates to disturbance by bank regrading. In comparison to diversity indices, a rarity index was much less sensitive to environmental factors than species diversity indices and more robust against seasonal and yearly fluctuations. Consequently, it has more potential for use in site quality assessment

The impact of surgical delay on resectability of colorectal cancer: An international prospective cohort study

AIM: The SARS-CoV-2 pandemic has provided a unique opportunity to explore the impact of surgical delays on cancer resectability. This study aimed to compare resectability for colorectal cancer patients undergoing delayed versus non-delayed surgery. METHODS: This was an international prospective cohort study of consecutive colorectal cancer patients with a decision for curative surgery (January-April 2020). Surgical delay was defined as an operation taking place more than 4 weeks after treatment decision, in a patient who did not receive neoadjuvant therapy. A subgroup analysis explored the effects of delay in elective patients only. The impact of longer delays was explored in a sensitivity analysis. The primary outcome was complete resection, defined as curative resection with an R0 margin. RESULTS: Overall, 5453 patients from 304 hospitals in 47 countries were included, of whom 6.6% (358/5453) did not receive their planned operation. Of the 4304 operated patients without neoadjuvant therapy, 40.5% (1744/4304) were delayed beyond 4 weeks. Delayed patients were more likely to be older, men, more comorbid, have higher body mass index and have rectal cancer and early stage disease. Delayed patients had higher unadjusted rates of complete resection (93.7% vs. 91.9%, P = 0.032) and lower rates of emergency surgery (4.5% vs. 22.5%, P < 0.001). After adjustment, delay was not associated with a lower rate of complete resection (OR 1.18, 95% CI 0.90-1.55, P = 0.224), which was consistent in elective patients only (OR 0.94, 95% CI 0.69-1.27, P = 0.672). Longer delays were not associated with poorer outcomes. CONCLUSION: One in 15 colorectal cancer patients did not receive their planned operation during the first wave of COVID-19. Surgical delay did not appear to compromise resectability, raising the hypothesis that any reduction in long-term survival attributable to delays is likely to be due to micro-metastatic disease

Proceedings of the 3rd Biennial Conference of the Society for Implementation Research Collaboration (SIRC) 2015: advancing efficient methodologies through community partnerships and team science

It is well documented that the majority of adults, children and families in need of evidence-based behavioral health interventionsi do not receive them [1, 2] and that few robust empirically supported methods for implementing evidence-based practices (EBPs) exist. The Society for Implementation Research Collaboration (SIRC) represents a burgeoning effort to advance the innovation and rigor of implementation research and is uniquely focused on bringing together researchers and stakeholders committed to evaluating the implementation of complex evidence-based behavioral health interventions. Through its diverse activities and membership, SIRC aims to foster the promise of implementation research to better serve the behavioral health needs of the population by identifying rigorous, relevant, and efficient strategies that successfully transfer scientific evidence to clinical knowledge for use in real world settings [3]. SIRC began as a National Institute of Mental Health (NIMH)-funded conference series in 2010 (previously titled the “Seattle Implementation Research Conference”; $150,000 USD for 3 conferences in 2011, 2013, and 2015) with the recognition that there were multiple researchers and stakeholdersi working in parallel on innovative implementation science projects in behavioral health, but that formal channels for communicating and collaborating with one another were relatively unavailable. There was a significant need for a forum within which implementation researchers and stakeholders could learn from one another, refine approaches to science and practice, and develop an implementation research agenda using common measures, methods, and research principles to improve both the frequency and quality with which behavioral health treatment implementation is evaluated. SIRC’s membership growth is a testament to this identified need with more than 1000 members from 2011 to the present.ii SIRC’s primary objectives are to: (1) foster communication and collaboration across diverse groups, including implementation researchers, intermediariesi, as well as community stakeholders (SIRC uses the term “EBP champions” for these groups) – and to do so across multiple career levels (e.g., students, early career faculty, established investigators); and (2) enhance and disseminate rigorous measures and methodologies for implementing EBPs and evaluating EBP implementation efforts. These objectives are well aligned with Glasgow and colleagues’ [4] five core tenets deemed critical for advancing implementation science: collaboration, efficiency and speed, rigor and relevance, improved capacity, and cumulative knowledge. SIRC advances these objectives and tenets through in-person conferences, which bring together multidisciplinary implementation researchers and those implementing evidence-based behavioral health interventions in the community to share their work and create professional connections and collaborations

Acylophorus acufer Lott, new species

Acylophorus acufer Lott, new species (Figs 23–28) Description. Length 5.5 –6.5mm. Body black except for dark chestnut margins of pronotum and the apical abdominal tergites, which are paler at both base and apex. Abdomen iridescent to varying degrees. Legs red. Antennae pale or dark with segment I always pale at base. Maxillary palpi usually all pale. Head small (pronotum 2.1x wider than head), more or less as long as wide with wide neck and temples barely suggested (Fig. 23). Antennae inserted right on front margin with no pigmented area in front. No micro-punctures visible at 80 x magnification. Short pubescence behind eyes localised and sparse. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other and five postocular setae visible from above on each side. No extra seta by hind margin of eye. Mandibles short with wide basal flange, a vestigial medial tooth discernible on the left mandible (Fig. 24). Maxillary palpi with terminal segment densely pubescent, narrow and more or less symmetric, longer than penultimate segment, which is short, triangular and glabrous (Fig. 25). First segment of antenna as long as next four (Fig. 26). Segments I to V elongate, VII to XI transverse. Pronotum slightly transverse (1.2x wider than long) with rounded sides and widest in basal half. Shining with no micro-punctures. One pair of dorsal setae. One pair of lateral setae. Marginal setae long. Elytra strongly transverse (1.7x wider than long) with pubescence arising from fine, asperate punctures. Fringe of bristles on hind margin longer than the hairs on the rest of the elytra. Empodial setae between claws on mid- and hind tarsi very short. Asperate punctures on abdominal tergites denser at base, pubescence longer than on elytra. Male with apex of sternite IX entire. Paramere bilobed, each lobe long and pointed, needle-shaped, much longer than median lobe of aedeagus, pegs confusedly arranged around or slightly in front of mid-point of each lobe (Figs 27 and 28). Median lobe barely expanded at apex which is rounded. Type material. Holotype 3: “ Namibia 30.iii. 1999 18 O 14 ’S / 21 O 43 ’E Mahango GR, Kwetche piknik site, banks of Okavango, shore washing, lg M. + B. Uhlig / HOLOTYPE Acylophorus acufer sp. n. 3 det. DA Lott, 2010 ” (ZMHB). Paratypes 536 Ƥ same data as holotype (ZMHB, cJanak); 3: “NAMIBIA-Exp. ZMB 1992 Mahango Game Reserve, Seeufer, Ufervegetation gesiebt, 18 O 17 ’S / 21 O 43 ’E 28.II. 92, leg M. Uhlig” (ZMHB); 1 Ƥ: “ NAMIBIA 24.xi. 1993 18 O 14 ’S / 21 O 43 ’E Kavango: Mahango Game Reserve: Okavango Papyrus sievings, leg Uhlig” (ZMHB); 8: “ NAMIBIA 1 + 4.iii. 1994 Kavango: Piknik site, Okavango banks, sievings, flood refuse, reed leaf litter, grass leg M. Uhlig” (ZMHB); 131 Ƥ: “ NAMIBIA 1 + 4.iii. 1994 18 O 14 ’S / 21 O 43 ’E Kavango: Mahango Game Reserve, Baobab, Okavango banks, sievings, flood refuse, leg M. Uhlig” (ZMHB); 6: “NAMIBIA-Exp. ZMB 1992 Buffalo Camp, Kavango-Ufer, Ufervegetation gesiebt, 18 O09’S / 21 O 42 ’E 28.II. 92, leg M. Uhlig” (ZMHB); 6: “NAMIBIA-Exp. ZMB 1992 Popa Falls, Kavango-Ufer, Ufervegetation gesiebt, 18 O07’S / 21 O 35 ’E 27.II. 92, leg M. Uhlig” (ZMHB); 97: “NAMIBIA-Exp. ZMB 1992 Popa Falls, Kavango-Ufer, Schilf – Papyrus - Ufervegetation gesiebt, 18 O07’S / 21 O 35 ’E 2.III. 92, leg M. Uhlig” (ZMHB); 16: “NAMIBIA-Exp. ZMB 1992 Popa Falls, 18 O07’S / 21 O 35 ’E Kavango-Ufer, Schilf – Papyrus - Ufervegetation gesiebt, 13.III. 92, leg M. Uhlig” (ZMHB); 8: “ NAMIBIA 1.iv. 1993 18 O07’ 16 ”S / 21 O 34 ’ 51 ”E Popa Falls, island banks of Okavango banks, reed - papyrus sievings, leg Uhlig” (ZMHB); 7: “ NAMIBIA 17.iv. 1993 18 O07’ 16 ”S / 21 O 34 ’ 51 ”E Popa Falls, island banks of Okavango banks, sievings: reed + papyrus, leg M. Uhlig” (ZMHB); 5: “ NAMIBIA 28.ii – 6. iii. 1994 18 O07’ 16 ”S / 21 O 34 ’ 51 ”E Popa Falls, Okavango banks, sievings, Papyrus, reed, grass, leaf litter, leg M. Uhlig” (ZMHB); 5: “ BOTSWANA 3.iii. 1994 18 O 48 ’S / 22 O08’E Sepupa, Okavango-Delta, sievings: grass + flood refuse, leg M. Uhlig” (ZMHB); 1 Ƥ: “ BOTSWANA 6.iv. 1998 Shakawe Fishing Camp 18 O 27 ’S / 21 O 56 ’E leg J. Deckert, gesiebt” (ZMHB). Distribution and bionomics. All records to date of this species come from the Okavango River system in Botswana and Namibia, where it appears to be the most frequently collected species of Acylophorus. Specimens were mainly obtained by sieving marsh litter, but a few specimens were collected by splashing riverbanks. Comparative notes. This species can be distinguished from both A. salifi and A. lomaensis by the less transverse pronotum, the more slender last segment of the maxillary palpi and, above all, by the form of the male genitalia. In addition, it is larger and darker than A. lomaensis. Populations of A. salifi occurring along the Okavango appear to be of the typical colour form (mainly black with reddish elytra), so colour may be an additional character that is useful for distinguishing A. acufer and A. salifi in the field. Etymology. This specific name is a noun in apposition meaning “bearer of needles”, a reference to the extraordinary shape of the parameres.Published as part of Lott, Derek A., 2012, Further studies of African Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae), pp. 39-52 in Zootaxa 3168 on page 49, DOI: 10.5281/zenodo.27978

Acylophorus makhoreae Lott, new species

Acylophorus makhoreae Lott, new species (Figs 8, 38, 58, 84, 116) Description. Length 7mm. Body colour very variable ranging from pale brown to black. Pronotum yellow to red-brown with a dark, diffuse mark covering the disc and sometimes suffused almost to the edges. Abdomen iridescent when dark. Appendages similarly variable in colour, but the terminal segment of the maxillary palpi is always darker than the penultimate segment. Head of average size (pronotum 1.75x wider than head), more or less as long as wide with rounded temples not very evident (Fig. 8). Pigmented area of head extending well in front of antennal insertion. Micropunctures extending over much of head, but very sparse away from the front of the head and next to the eyes. Dense short pubescence behind eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. Only four postocular setae visible from above on each side. Underside of head sparsely pubescent, depressed at base with gular sutures separate, but very proximate toward base. Right mandible with one sharp median tooth; left mandible lacking sharp tooth (Fig. 38). Maxillary palpi with terminal segment densely pubescent, with rounded angle on outer margin, asymmetric and less elongate than A. orientalis, longer than glabrous penultimate segment which is slightly elongate (Fig. 58). First segment of antenna as long as next five. Segments I to IV elongate, VII to XI transverse (Fig. 84). Pronotum only slightly transverse (1.1x wider than long) with rounded sides and widest in basal half. Shining with no micro-punctures. Dorsal, lateral and marginal setae shorter than in A. orientalis. Elytra transverse (1.7x wider than long) with pubescence arising from asperate punctures. Apical fringe of bristles longer than the hairs on the rest of the elytra. Abdominal tergites with evenly spaced, relatively sparse asperate punctures. Paramere bilobed, each lobe fairly flat, at least toward apex, pegs concentrated in dense mass at apex (Fig. 116). Median lobe longer than paramere with truncate apex. Type material. Holotype 3: “Under plants at a stream edge / ETHIOPIA: Kaffa 28km. S. of Jimma 2,000 m’ xi. 1971 / R. O.S. Clark B.M. 1973 - 450 / HOLOTYPE Acylophorus makhoreae sp. n. 3 det. DA Lott, 2009 ” (BMNH). Paratypes 13 1 Ƥ: “ ETHIOPIA: ILUBADOR 10km. W. Bedelle 0825N 3618 E x. 1972 1,800 m. / R. O.S. Clark B.M. 1973 - 450 / PARATYPE Acylophorus makhoreae sp. n. det. DA Lott, 2009 ” (BMNH); 4: “Under plants & stones at edge stream / ETHIOPIA: Kaffa Belleta F. 2,100 m 40km. S.W. Jimma 1971 / R. O.S. Clark B.M. 1973 - 450 / PARATYPE Acylophorus makhoreae sp. n. det. DA Lott, 2009 ” (BMNH); 1 Ƥ: “ih red ref. / ETHIOPIA: Kaffa Jimma to Addis Rd. 0749- 370 iii. 1972. / R. O.S. Clark B.M. 1973 - 450 / PARATYPE Acylophorus makhoreae sp. n. det. DA Lott, 2009 ” (BMNH). Distribution and bionomics. All material seen so far comes from the south-west of Ethiopia (Fig. 142). I have also seen a female specimen collected at “ 8,000 ” feet from the Djem-Djem Forest to the west of Addis Ababa that may belong to this species. Most of the specimens were collected along streams. Comparative notes. Fairly distinct within the A. orientalis species group by virtue of the position of the antennal insertion, the broader, more asymmetric terminal segment of the maxillary palpi and the more transverse medial antennal segments. The form of the aedeagus and the arrangement of medial teeth on the mandibles are also useful characters. Etymology. Makhore was a legendary queen, who founded the kingdom of Jimma in the area where many of the type specimens were collected. The species name is the genitive case of a Latinised version of her name. Discussion. A. makhoreae has been placed in the A. orientalis group, because it lacks dense micropunctures on the head and pronotum. However, the form of the maxillary palpi, the mandibles and the aedeagus and the position of the antennal insertion point all suggest that it may be more closely related to some of the species in the A. densipennis group.Published as part of Lott, Derek A., 2010, The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa, pp. 1-51 in Zootaxa 2402 on page 15, DOI: 10.5281/zenodo.27590

Acylophorus capensis Cameron

Acylophorus capensis Cameron (Figs 5, 36, 55, 82, 113) Acylophorus capensis Cameron, 1945: 716; Herman, 2001: 3028. Redescription. Length 7mm. Body black with iridescent abdomen. Abdominal tergite and sternite VIII black, occasionally pale at extreme base and apex. Antennae dark brown with segment XI and base of segment I occasionally red-brown. Legs pale with darkened meso- and meta-femora and tibiae. Palpi completely pale. Head large (pronotum 1.4–1.6x wider than head), as long as wide with temples well developed (Fig. 5). Mostly shining with sparse micro-punctures confined to front of head, but with dense short pubescence behind eyes. Antennae inserted close to front margin of head, but separated from it by a small pigmented strip. Temples long and pronounced behind relatively small eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. Four postocular setae visible from above on each side with a further seta out of line on the hind-border of the eye. Mandibles with two medial teeth on each side (Fig. 36). Maxillary palpi with terminal segment elongate and sparsely pubescent, almost symmetric with attenuated apex, longer than glabrous penultimate segment which is slightly elongate (Fig. 55). First segment of antenna longer than next three. Segments I to VI elongate, IX to XI 1 transverse (Fig. 81). Pronotum weakly transverse (1.025 –1.1x wider than long) with rounded sides, widest in basal half. One pair of dorsal setae (The type specimen has two proximate pairs of dorsal punctures, but this is probably an aberration.) One pair of lateral setae. Marginal setae long. Elytra transverse (1.5x wider than long) with long, close pubescence and asperate punctures which are sparser, but stronger than in A. orientalis. Fringe of bristles on hind margins longer than the general pubescence. Abdominal tergites also with long pubescence and strong asperate punctures, but not quite as dense as on elytra. Punctures sparser on apical half of each tergite than on basal half. Aedeagus with the paramere divided into two slightly converging, proximate lobes, each lobe being ridged dorsally, though much less so than in A. orientalis (Fig. 113). The internal pegs are confusedly distributed toward the apex. The median lobe is appreciably longer than the paramere and markedly expanded at the apex. Type material. Cameron described the species from a unique type in his collection. This specimen is the holotype by monotypy. Holotype Ƥ: “ Type / Marsh / Simon’s Town iv-vi- 15 MC / A. capensis Cam. TYPE ” (BMNH). The name capensis was also used for this species by Fauvel, but in manuscript form on a specimen label. He never published any description. Further material examined. SOUTH AFRICA: Eastern Cape: Hogsback, 970–1300m, 32 O 35 ’S 26 O 56-57 ’E, J Janák, 5–7. xii. 2006, 23 (cJanák); KwaZulu-Natal: Pietermaritzburg, Queen Elizabeth Park, 29 O 34.157 ’S 30 O 19.299 ’E, J Janák, 22.xi. 2006, 233Ƥ (cJanák & cLott); Western Cape: Cape Town, ex. Coll. Fauvel, 1 Ƥ (IRSNB); Swellendam, R. Kmeco, 28.xi– 1. xii. 1997, 13 (cJanák); Harkerville S.F., Kranshoek Picnic Area, 145m, 34 O 04.7 ’S 23 O 13.6 ’E, 1. ii. 2004, 13 (FMNH); Grootvadersbosch N.R., Bushbuck Trail (Bosbockrand), 360m, 33 O 58.9 ’S 20 O 49.1 ’E, 26. i. 2004, 1 Ƥ (FMNH). Distribution and bionomics. This species is only known so far from South Africa (Fig. 141). All recent specimens have been taken from stream margins. In two localities these were in “afromontane forest”. Comparative notes. Distinctive within the species group by virtue of its large head. Most s imilar to A. rossii but has even smaller eyes. The distinctive shape of the terminal segment of the maxillary palp, the arrangement of medial teeth on the mandibles, the short paramere in comparison to the median lobe of the aedeagus and the sparse, but strong punctures on the elytra are also useful diagnostic characters.Published as part of Lott, Derek A., 2010, The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa, pp. 1-51 in Zootaxa 2402 on pages 12-13, DOI: 10.5281/zenodo.27590

Acylophorus collarti Cameron

Acylophorus collarti Cameron (Figs 11, 41, 61, 87, 119) Acylophorus collarti Cameron, 1935: 375; Herman, 2001: 3027. Redescription. Length 6.5– 7mm. Body generally black, sometimes with dark brown abdomen or more generally dark brown. Leg colour very variable from all pale to black with red tarsi. Maxillary palpi pale with last segment darker. Antennae pale with middle segments slightly darkened to all dark. Head large (pronotum 1.6x wider than head), 1.1x longer than wide with evident temples (Fig. 11). Pigmented area of head produced in front of antennal insertion. Forehead somewhat arched. Eyes relatively small. Covered with dense micro-punctures. Dense pale pubescence behind eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. Four postocular setae visible on each side, additional seta on hind margin of eye absent. Underside of head strongly depressed at base. Mandibles with one well developed medial tooth in front of a vestigial tooth on the right and just a flange on the left (Fig. 41). Maxillary palpi with terminal segment pubescent, more rounded on outer margin than inner margin and asymmetric, sometimes slightly attenuated at apex, longer than glabrous penultimate segment which is triangular (Fig. 61). First segment of antenna as long as next five. Segments I to IV elongate, IX and X transverse (Fig. 87). Pronotum relatively narrow with sides less rounded, almost quadrate (1.05x wider than long) and widest toward basal half, covered with dense micro-punctures. One pair of dorsal setae and one pair of lateral setae. Marginal setae short. Elytra only slightly transverse (1.4x wider than long) with close, bright yellow pubescence. Asperate punctures strong as in A. orientalis. Fringe of apical bristles slightly longer than the pubescence on the rest of the elytra. Abdominal tergites with similar pubescence. Punctation on abdominal tergites finer than on elytra and becoming sparser on apical segments. Sternite IX of male with apex entire. Paramere of aedeagus bilobed, lobes slightly divergent, pegs confusedly arranged in apical inner half of each lobe, base lipped as in A. densipennis (Fig. 119). Median lobe not longer than paramere, barely expanded at apex, which is truncate. Type material. Cameron described the species from specimens collected by A. Collart on 26 th January 1929 at Blukwa (Nizi) in what is now the Democratic Republic of Congo. Two specimens from the type series have been located and one of these is here designated as the lectotype in order to fix the identity of the species. Lectotype 3: “ Paratype / Nizi: Blukwa; 26 -I- 29; A. COLLART / TYPE / Acylophorus Collarti Cam. TYPE / LECTOTYPE Acylophorus collarti Cameron 3 det. DA Lott, 2009 ” (IRSNB); Paralectotype 1 Ƥ: “ Paratype / Nizi: Blukwa; 26 -I- 29; A. COLLART / M. Cameron Bequest B.M. 1955 - 147 / Acyl. Collarti Cam. COTYPE / PARALECTOTYPE Acylophorus collarti Cameron 3 det. DA Lott, 2009 ” (BMNH). Further material examined. CAMEROON: Northwest Province: 3km SW of Bamenda, 1300m, RL Aalbu, 5. ii. 1980, 13 (FMNH). R.D. CONGO: Nizi: Blukwa, A Collart, 10.xii. 1928 & 25.i. 1929, 132Ƥ (IRSNB). Distribution and bionomics. Only recorded from the Democratic Republic of Congo and Cameroon (Fig. 144). In Cameroon it was recorded in a stream-side thicket in afromontane forest. Comparative notes. Similar to A. densipennis by virtue of the bright yellow pubescence on the elytra, but generally larger and with smaller eyes. The apical bristles on the elytra are longer and the aedeagus is completely different. Much more similar to A. antennalis, from which it is best distinguished by the aedeagus. There may also be differences in the form of the mandibles and the maxillary palpi are narrower, but the extent of intraspecific variation in these characters is not yet understood.Published as part of Lott, Derek A., 2010, The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa, pp. 1-51 in Zootaxa 2402 on pages 18-19, DOI: 10.5281/zenodo.27590

Acylophorus methneri Bernhauer

Acylophorus methneri Bernhauer (Figs 17, 93) Acylophorus methneri Bernhauer, 1917: 48; Scheerpeltz, 1933: 1467; Bernhauer & Chapman, 1939: 74; Scheerpeltz, 1974: 25; Herman, 2001: 3034. Description. Length 6.5mm. Body relatively wide and flat, pitchy with paler pronotum. Abdomen with weak iridescence. Legs and palpi pale, antennae mostly infuscate with pale terminal segments. Head somewhat small (pronotum 1.8x wider than head) as long as wide, pigmented area produced in front of antennal insertion (Fig. 17). Side of head rounded with temples not very prominent. Covered with dense micro-punctures. Dense pale pubescence behind eyes. Four rather short postocular setae visible from above. Two pairs of short and pale interocular setae arising from shallow punctures less than twice as close to eyes as to each other. Maxillary palpi missing on material available for study. First segment of antenna longer than next four. Segments I to V elongate, VII to XI transverse (Fig. 93). Pronotum slightly transverse (1.1x wider than long) with rounded sides and widest in basal half, micropunctures much sparser than on head. Dorsal and lateral setae short. Marginal setae short and reduced in number. Elytra transverse (1.7x wider than long) with short pubescence. Apical fringe of bristles short, slightly longer than hairs on the rest of the elytra. Punctures sparse and less obviously asperate than in other species. Abdominal tergites with short pubescence on basal segments, not overlapping on tergites III, IV and V, becoming longer on apical segments, punctures fine and dense throughout. Form of the aedeagus unknown. Type material. Bernhauer described this species from a single specimen, which has been located. This specimen is the holotype by monotypy. Holotype Ƥ: “D. Ostafrika; Methner; Kaguru. v.09. Bach U. / Methneri Bernh. Typus unicus / Chicago NHMus M. Bernhauer Collection / HOLOTYPE Acylophorus methneri Bernhauer Ƥ det. DA Lott, 2009 ” (FMNH). Distribution and bionomics. The type was collected from the banks of a stream at Kaguru, Tanzania (Fig. 143). There are further records in the literature from Kenya (Bernhauer & Chapman, 1939) and Sudan (Scheerpeltz, 1974), but these must be open to question given the frequency of misidentifications in collections. Comparative notes. Easily recognised by the extremely short pubescence on the elytra and abdomen and the short setae on the head and pronotum.Published as part of Lott, Derek A., 2010, The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa, pp. 1-51 in Zootaxa 2402 on pages 23-25, DOI: 10.5281/zenodo.27590

Acylophorus micans Lott 2010

&lt;i&gt;Acylophorus micans&lt;/i&gt; Lott, 2010 &lt;p&gt; &lt;b&gt;Material examined. CAMEROON&lt;/b&gt;: Soppo, SG von Rothkirch, xi.1913, 13 (ZMBH). &lt;b&gt;Discussion.&lt;/b&gt; This record comes from within the known distributional range of this species, which extends from Cote d&rsquo;Ivoire to Gabon.&lt;/p&gt;Published as part of &lt;i&gt;Lott, Derek A., 2012, Further studies of African Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae), pp. 39-52 in Zootaxa 3168&lt;/i&gt; on page 43, DOI: &lt;a href="http://zenodo.org/record/279788"&gt;10.5281/zenodo.279788&lt;/a&gt