Predicting understory maximum shrubs cover using altitude and overstory basal area in different Mediterranean forests

In some areas of the Mediterranean basin where the understory stratum represents a critical fire hazard, managing the canopy cover to control the understory shrubby vegetation is an ecological alternative to the current mechanical management techniques. In this study, we determine the relationship between the overstory basal area and the cover of the understory shrubby vegetation for different dominant canopy species (Pinaceae and Fagaceae species) along a wide altitudinal gradient in the province of Catalonia (Spain). Analyses were conducted using data from the Spanish National Forest Inventory. At the regional scale, when all stands are analysed together, a strong negative relationship between mean shrub cover and site elevation was found. Among the Pinaceae species, we found fairly good relationships between stand basal area and the maximum development of the shrub stratum for species located at intermediate elevations (Pinus nigra, Pinus sylvestris). However, at the extremes of the elevationclimatic gradient (Pinus halepensis and Pinus uncinata stands), stand basal area explained very little of the shrub cover variation probably because microsite and topographic factors override its effect. Among the Fagaceae species, a negative relationship between basal area and the maximum development of the shrub stratum was found in Quercus humilis and Fagus sylvatica dominated stands but not in Quercus ilex. This can be due to the particular canopy structure and management history of Q. ilex stands. In conclusion, our study revealed a marked effect of the tree layer composition and the environment on the relationship between the development of the understory and overstory tree structure. More fine-grained studies are needed to provide forest managers with more detailed information about the relationship between these two forest strata

Effects of the overstory on the diversity of the herb and shrub layers of Anatolian black pine forests

Understory plants are important components of forests because they are responsible for the majority of the vascular plant diversity of forest ecosystems. The richness and composition of understory communities are closely related to the tree layer diversity, structure and composition. The aim of this study was to examine the understory diversity of Anatolian black pine (Pinus nigra Arnold subsp. pallasiana (Lamb.) Holmboe)-dominated forests on the KazdagA +/- Mountains of West Turkey. To describe the overstory structure and composition in a numerically and quantitatively well-defined manner, cumulative abundance profiles (CAPs) of the tree species were used. The resemblance of the sampling plots was classified into five stand types assessing the CAP through the Fuzzy C-Means clustering method. A permutational multivariate analysis of variance (PERMANOVA) was performed to test the variance of the community ecological distance between the five stand types, and the results showed significant differences in these clusters. Many shade-tolerant plants were associated with the mixed stands of Anatolian black pine-KazdagA +/- fir. The composition of the herb and shrub layer could not be explained by the environmental variables but by differences in the overstory structure of the stands. Pure or nearly pure Anatolian Black pine stands were more diverse than mixed oak-Anatolian black pine and KazdagA +/- fir-Anatolian black pine stands. However, although dense and young pure Anatolian black pine stands had the most diverse plant species in the shrub layer, they were ranked third in terms of the herb layer diversity. The Anatolian black pine-KazdagA +/- fir mixed stands had the lowest herb and shrub layer diversity. These results allow us to comprehend the relationship between the overstory structure and composition, and the understory diversity. Understanding this relationship is important for the conservation of understory plant diversity in the management of forest ecosystems