140 research outputs found

    Time-of-day/time-of-year response functions of planetary geomagnetic indices

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    Aims: To elucidate differences between commonly-used mid-latitude geomagnetic indices and study quantitatively the differences in their responses to solar forcing as a function of Universal Time (UT), time-of-year (F), and solar-terrestrial activity level. To identify the strengths, weaknesses and applicability of each index and investigate ways to correct for any weaknesses without damaging their strengths. Methods: We model how the location of a geomagnetic observatory influences its sensitivity to solar forcing. This modelling for a single station can then be applied to indices that employ analytic algorithms to combine data from different stations and thereby we derive the patterns of response of the indices as a function of UT, F and activity level. The model allows for effects of solar zenith angle on ionospheric conductivity and of the station’s proximity to the midnight-sector auroral oval: it employs coefficients that are derived iteratively by comparing data from the current aa index stations (Hartland and Canberra) to simultaneous values of the am index, constructed from chains of stations in both hemispheres. This is done separately for 8 overlapping bands of activity level, as quantified by the am index. Initial estimates were obtained by assuming the am response is independent of both F and UT and the coefficients so derived were then used to compute a corrected F-UT response pattern for am. This cycle was repeated until it resulted in changes in predicted values that were below the adopted uncertainty level (0.001%). The ideal response pattern of an index would be uniform and linear (i.e., independent of both UT and F and the same at all activity levels). We quantify the response uniformity using the percentage variation at any activity level, V = 100(S/), where S is the index’s sensitivity at that activity level and S is the standard deviation of S: both S and S were computed using the 8 UT ranges of the 3-hourly indices and 20 equal-width ranges of F. As an overall metric of index performance, we take an occurrence-weighted mean of V, Vav, over the 8 activity-level bins. This metric would ideally be zero and a large value shows that the index compilation is introducing large spurious UT and/or F variations into the data. We also study index performance by comparisons with the SME and SML indices, compiled from a very large number of stations, and with an optimum solar wind “coupling function”, derived from simultaneous interplanetary observations. Results: It is shown that a station’s response patterns depend strongly on the level of geomagnetic activity because at low activity levels the effect of solar zenith angle on ionospheric conductivity dominates over the effect of station proximity to the midnight-sector auroral oval, whereas the converse applies at high activity levels. The metric Vav for the two-station aa index is modelled to be 8.95%, whereas for the multi-station am index it is 0.65%. The ap (and hence Kp) index cannot be analyzed directly this way because its construction employs tabular conversions, but the very low Vav for am allows us to use / to evaluate the UT-F response patterns for ap. This yields Vav = 11.20% for ap. The same empirical test applied to the classical aa index and the new “homogenous” aa index, aaH (derived from aa using the station sensitivity model), yields Vav of, respectively, 10.62% (i.e., slightly higher than the modelled value) and 5.54%. The ap index value of Vav is shown to be high because it exaggerates the average semi-annual variation and has an annual variation giving a lower average response in northern hemisphere winter. It also contains a strong artefact UT variation. We derive an algorithm for correcting for this uneven response which gives a corrected ap value, apC, for which Vav is reduced to 1.78%. The unevenness of the ap response arises from the dominance of European stations in the network used and the fact that all data are referred to a European station (Niemegk). However, in other contexts, this is a strength of ap, because averaging similar data gives increased sensitivity and more accurate values on annual timescales, for which the UT-F response pattern is averaged out

    RARÎł is critical for maintaining a balance between hematopoietic stem cell self-renewal and differentiation

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    Hematopoietic stem cells (HSCs) sustain lifelong production of all blood cell types through finely balanced divisions leading to self-renewal and differentiation. Although several genes influencing HSC self-renewal have been identified, to date no gene has been described that, when activated, enhances HSC self-renewal and, when activated, promotes HSC differentiation. We observe that the retinoic acid receptor (RAR)γ is selectively expressed in primitive hematopoietic precursors and that the bone marrow of RARγ knockout mice exhibit markedly reduced numbers of HSCs associated with increased numbers of more mature progenitor cells compared with wild-type mice. In contrast, RARα is widely expressed in hematopoietic cells, but RARα knockout mice do not exhibit any HSC or progenitor abnormalities. Primitive hematopoietic precursors overexpressing RARα differentiate predominantly to granulocytes in short-term culture, whereas those overexpressing RARγ exhibit a much more undifferentiated phenotype. Furthermore, loss of RARγ abrogated the potentiating effects of all-trans retinoic acid on the maintenance of HSCs in ex vivo culture. Finally, pharmacological activation of RARγ ex vivo promotes HSC self-renewal, as demonstrated by serial transplant studies. We conclude that the RARs have distinct roles in hematopoiesis and that RARγ is a critical physiological and pharmacological regulator of the balance between HSC self-renewal and differentiation

    Diversification of an Endemic Southeast Asian Genus: Phylogenetic Relationships of the Spiderhunters (Nectariniidae: Arachnothera)

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    This is the publisher's version, also available at http://www.bioone.org/doi/abs/10.1525/auk.2011.11019The phylogeny of spiderhunters (Nectariniidae: Arachnothera) was reconstructed by comparing mitochondrial and nuclear DNA sequences of all currently recognized species and with broad geographic sampling of two particularly variable species complexes, the Little Spiderhunter (Arachnothera longirostra) and the streaky spiderhunters (A. modesta and A. affinis). It appears to be a relatively old group, whose diversification was not caused by recent sea-level changes. However, the modern, highly sympatric distribution of the large species in the Sunda lowlands was probably a result of dispersal via recent land bridges. Within the highly variable A. longirostra group, there are substantially diverged taxa in the Philippines that should be considered different species. Within the A. affinis—modesta complex, there are three distinct species and a closely related fourth, which describe a clear allopatric distribution: A. affinis in Java, A. modesta in the rest of the Sunda lowlands (except Sabah), A. magna in the Malayan highlands and mainland Southeast Asia, and A. everetti in the Bornean highlands and Sabah. Depending on whether mitochondrial or nuclear genes were compared, monophyly of the genus was disrupted by a single outgroup sunbird (Hypogramma hypogrammicum) or by all outgroup sunbirds included in the study. The discrepancy between nuclear and mitochondrial results is probably a case of deep coalescence and will require additional markers for resolution

    The Lantern Vol. 12, No. 1, January 1944

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    • In the Den of the Titans • Lass of Dimoch • For One Gone West • Tropical New Year • Nightfall • Hope From the Blue • Christmas Shopping, a Retrospective Glance • What Makes a Wing Lift? • 201 Main Street • Paradox • Stabilizing Americans • Sky-Islands • With Timbrel and Dance • Jazz • The Grave, a Translationhttps://digitalcommons.ursinus.edu/lantern/1031/thumbnail.jp

    Semi-annual, annual and Universal Time variations in the magnetosphere and in geomagnetic activity: 4. Polar Cap motions and origins of the Universal Time effect

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    We use the am, an, as and the a-sigma geomagnetic indices to the explore a previously overlooked factor in magnetospheric electrodynamics, namely the inductive effect of diurnal motions of the Earth’s magnetic poles toward and away from the Sun caused by Earth’s rotation. Because the offset of the (eccentric dipole) geomagnetic pole from the rotational axis is roughly twice as large in the southern hemisphere compared to the northern, the effects there are predicted to be roughly twice the amplitude of those in the northern hemisphere. Hemispheric differences have previously been discussed in terms of polar ionospheric conductivities generated by solar photoionization, effects which we allow for by looking at the dipole tilt effect on the time-of-year variations of the indices. The electric field induced in a geocentric frame is shown to also be a significant factor and gives a modulation of the voltage applied by the solar wind flow in the southern hemisphere that is typically a 30% diurnal modulation for disturbed intervals rising to about 76% in quiet times. For the northern hemisphere these are 15% and 38% modulations. Motion towards/away from the Sun reduces/enhances the directly-driven ionospheric voltages and reduces/enhances the magnetic energy stored in the tail and we estimate that approximately 10% of the effect appears in directly driven ionospheric voltages and 90% in changes of the rate of energy storage or release in the near-Earth tail. The hemispheric asymmetry in the geomagnetic pole offsets from the rotational axis is shown to be the dominant factor in driving Universal Time (UT) variations and hemispheric differences in geomagnetic activity. Combined with the effect of solar wind dynamic pressure and dipole tilt on the pressure balance in the near-Earth tail, the effect provides an excellent explanation of how the observed Russell-McPherron pattern with time-of-year F and UT in the driving power input into the magnetosphere is converted into the equinoctial F - UT pattern in average geomagnetic activity (after correction is made for dipole tilt effects on ionospheric conductivity), added to a pronounced UT variation with minimum at 02-10UT. In addition, we show that the predicted and observed UT variations in average geomagnetic activity has implications for the occurrence of the largest events that also show the nett UT variation
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