The Energy Benefits of the Pantograph Wing Mechanism in Flapping Flight: Case Study of a Gull

Bird wings generally contain a 4-bar pantograph mechanism in the forearm that enables the wrist joint to be actuated from the elbow joint thus reducing the number of wing muscles and hence reducing the wing inertia and inertial drag. In this paper we develop a theoretical model of inertial power for flapping flight to estimate the advantage of the 4-bar pantograph mechanism by comparing the inertial power required for the case where wrist muscles are present in the forearm with the case where wrist muscles are not present in the forearm. It is difficult to predict how wrist muscles would look when there is no pantograph mechanism. Therefore a lower bound and upper bound case are defined. The lower bound case involves redistributing the elbow muscles with no increase in wing mass. The upper bound case involves replicating the biceps-triceps muscles near the wrist joint. At minimum power speed the model estimates that the 4-bar pantograph mechanism reduces the inertial power for the gull from between 6.1%-12.3% and reduces the overall power by 0.6%-1.2%. When account is taken of the tight margins involved in the design of a flying vehicle, the energy savings produced by the pantograph mechanism are significant. A ring-billed gull was chosen for the case study and an adult specimen was obtained to gather morphometric data. Lessons for the design of flapping micro air vehicles are discussed

SIGNIFICANT DEVELOPMENTS IN SOUTHERN AGRICULTURE IN THE DECADE OF THE 1970S AND EMERGING POLICY ISSUES IN THE 1980S

Teaching/Communication/Extension/Profession,

AGRICULTURAL BANKS IN THE SOUTHEAST AND NATION: A STUDY IN CONTRASTS

Agricultural Finance,

Interregional and International Competition in the Pork Industry

International Relations/Trade, Livestock Production/Industries, Marketing,

The Circadian Clock Gene Period1 Connects the Molecular Clock to Neural Activity in the Suprachiasmatic Nucleus.

The neural activity patterns of suprachiasmatic nucleus (SCN) neurons are dynamically regulated throughout the circadian cycle with highest levels of spontaneous action potentials during the day. These rhythms in electrical activity are critical for the function of the circadian timing system and yet the mechanisms by which the molecular clockwork drives changes in the membrane are not well understood. In this study, we sought to examine how the clock gene Period1 (Per1) regulates the electrical activity in the mouse SCN by transiently and selectively decreasing levels of PER1 through use of an antisense oligodeoxynucleotide. We found that this treatment effectively reduced SCN neural activity. Direct current injection to restore the normal membrane potential partially, but not completely, returned firing rate to normal levels. The antisense treatment also reduced baseline [Ca(2+)]i levels as measured by Fura2 imaging technique. Whole cell patch clamp recording techniques were used to examine which specific potassium currents were altered by the treatment. These recordings revealed that the large conductance [Ca(2+)]i-activated potassium currents were reduced in antisense-treated neurons and that blocking this current mimicked the effects of the anti-sense on SCN firing rate. These results indicate that the circadian clock gene Per1 alters firing rate in SCN neurons and raise the possibility that the large conductance [Ca(2+)]i-activated channel is one of the targets

Class of {varphi}X174 Mutants Relatively Deficient in Synthesis of Viral RNA

Nonpermissive cells infected with {varphi}X174 gene D amber mutants synthesized some sixfold less viral RNA than permissive cells. The decrease was unaffected by increasing the multiplicity of infection and was a consequence of an overall decrease in all viral RNA species. It is suggested that the gene D product may function in replicative form DNA unwinding to expose the template for transcription

Rotifer rDNA-specific R9 retrotransposable elements generate an exceptionally long target site duplication upon insertion

Author Posting. © The Author(s), 2009. This is the author's version of the work. It is posted here by permission of Elsevier B.V. for personal use, not for redistribution. The definitive version was published in Gene 448 (2009): 145-150, doi:10.1016/j.gene.2009.08.016.Ribosomal DNA genes in many eukaryotes contain insertions of non-LTR retrotransposable elements belonging to the R2 clade. These elements persist in the host genomes by inserting site-specifically into multicopy target sites, thereby avoiding random disruption of single-copy host genes. Here we describe R9 retrotransposons from the R2 clade in the 28S RNA genes of bdelloid rotifers, small freshwater invertebrate animals best known for their long-term asexuality and for their ability to survive repeated cycles of desiccation and rehydration. While the structural organization of R9 elements is highly similar to that of other members of the R2 clade, they are characterized by two distinct features: sitespecific insertion into a previously unreported target sequence within the 28S gene, and an unusually long target site duplication of 126 bp. We discuss the implications of these findings in the context of bdelloid genome organization and the mechanisms of target-primed reverse transcription.This work was supported by the U.S. National Science Foundation grant MCB-0821956 to I.A

Bark Beetle-Fire Associations in the Greater Yellowstone Area

The large forest fires in and around Yellowstone National Park in 1988 bring up many ecological questions, including the role of bark beetles. Bark beetles may contribute to fuel buildup over the years preceding a fire, resulting in stand replacement fires. Fire is important to the survival of seral tree species and bark beetles that reproduce in them. Without fire, seral species are ultimately replaced by climax species. Following fire, bark- and wood-boring beetles respond to fire-injured trees. Because of synchrony of the fires and life cycles of the beetles, beetle infestation in 1988 was not observed in fire-injured trees. However, endemic populations of beetles, beetle infestation in 1988 was not observed in fire-injured trees. However, endemic populations of beetles, upon emergence in 1989, infested large numbers of fire-injured trees. Of the trees examined in each species, 28 to 65 percent were infested by bark beetles: Pinus contora (28 percent) by Ips pini:; Pseudotsuga menziesii (32 percent) by Dendroctonus pseudotsugae; Picea engelmannii (65 percent) by Dendroctonus rufipennis; and Abies lasiocarpa (35 percent) by Buprestidae and Cerambycidae. Most trees infested by bark beetles had 50 percent or more of their basal circumference killed by fire. Bark beetle populations probably will increase in the remaining fire-injured trees

A Photographic Study of the Effects of Beating on Fiber Structure

Because beating is the most fundamental, but highly important, process in the paper industry, some of the changes in fiber structure that occur during beating have been investigated. A literature survey is also presented concerning a number of related investigations of fiber structure. The experimental results indicate that fibrillation begins very shortly after the beating process is started. As beating progresses, the primary cell wall is removed, and the specific area of the fiber increases greatly when the watery medium encounters the secondary lamella area of the fiber. Continued beating shows that the action is a mechanical one, in that the fibers become bruised, brushed and cut into shorter lengths. The mechanical agitation of the Valley Beater unraveled the small fibrils that are wrapped spirally around the fiber. Prolonged beating eventually detached the fibrils and destroyed fiber structure entirely. After five hours beating, all that remained was a mass of fibrils that readily formed a mat on the slide made for microscopic study. These results compare favorably with published data. The investigation did not prove that mucilage formed after extensive beating is amorphous. It appears as though the mucilage consists of fibrils and micro-fibrils, yet, equipment limitations prevented further work along these lines