Philibaetis Gen. Nov., A New Genus from the Philippines (Ephemeroptera, Baetidae)

Investigations of type material and new material from the Philippines (Luzon) revealed that Baetis luzonensis Müller-Liebenau, 1982 and B. realonae Müller-Liebenau, 1982 do not belong to Baetis Leach, 1815. A new genus, Philibaetis gen. nov., is described to accommodate both species and both are re-described based on larvae. The new genus is characterised by having a rectangular labrum with a submarginal row of long, simple setae on the dorsal surface and ventrally on lateral margins long, simple, spine-like setae, on anterolateral margins long, feathered setae and medially long, bifid setae and a partial, submarginal row of lanceolate setae. Both mandibles have blade-like incisors and dorsally, a mediolateral patch of long, spine-like setae; additionally, the left mandible has a tuft of long, partly branched setae at the base of the subtriangular process. Philibaetis gen. nov. is further characterised by a hypopharynx with a medial tuft of stout setae and anterolaterally, two smaller tufts of stout setae, a galea-lacinia with the distal denti-seta tooth-like and directed against canines, a fore femur apically with stout setae, both on anterior and posterior side and without a femoral patch and a claw with one row of denticles and two or three subapical setae. The protogonostyli under the cuticle of male last instar larvae are folded in the Labiobaetis type, excluding their affiliation to the genus Baetis. COI sequences were obtained from both species. The genetic distance (Kimura 2-parameter) between them is 17.5% on average. Very limited genetic distances of 0% to 3% (0.75% on average) were found between specimens of P. luzonensis comb. nov

A new genus of Baetidae (Insecta, Ephemeroptera) from Southeast Asia

A new genus of Baetidae is described from Southeast Asia, Procerobaetis gen. nov. It has a wide distribution reaching from Indonesia (Sumatra) to the Philippines. Two new species are described from Indonesia, P. leptobranchius gen. et sp. nov. and P. petersorum gen. et sp. nov., and one new species from the Philippines, P. freitagi gen. et sp. nov. Procerobaetis gen. nov. is characterized by having seven pairs of elongate, apically pointed gills. At least gills I and II are very slender with strongly extended points, which is unique in Baetidae. Similar gills were described from Leptophlebiidae. Procerobaetis gen. nov. is further characterized by having long, slender legs with extended, slender and slightly bent claws. The antennae posess remarkable spines at the outer, lateral margin, which are maximally developed on segments IX–XI of the flagellum. No spines are present on the posterior margins of abdominal tergites I–VI. COI sequences were obtained from all three of the new species. The genetic distances (Kimura 2-parameter) between these species are between 13% and 20%. Very limited genetic distances of 0% to 1% were found between specimens of the same species. The occurrence of two different species in the same area of Sumatra is discussed

First contribution to Labiobaetis Novikova & Kluge in Cambodia (Ephemeroptera, Baetidae), with description of two new species

Material collected in 2018 in Cambodia gives us first insights into the diversity of Labiobaetis Novikova & Kluge, 1987 in this country. No species has been reported so far. We identified two new species using a combination of morphology and genetic distance (COI, Kimura 2-parameter). They are described and illustrated based on their larvae. A key to all Labiobaetis species of continental Southeast Asia is provided. The interspecific K2P distance between the two new species is 20–21%, the intraspecific distance of one of them is 1%. The total number of Labiobaetis species worldwide is augmented to 156

A Rapid and Accurate MinION-Based Workflow for Tracking Species Biodiversity in the Field

Genetic markers (DNA barcodes) are often used to support and confirm species identification. Barcode sequences can be generated in the field using portable systems based on the Oxford Nanopore Technologies (ONT) MinION sequencer. However, to achieve a broader application, current proof-of-principle workflows for on-site barcoding analysis must be standardized to ensure a reliable and robust performance under suboptimal field conditions without increasing costs. Here, we demonstrate the implementation of a new on-site workflow for DNA extraction, PCR-based barcoding, and the generation of consensus sequences. The portable laboratory features inexpensive instruments that can be carried as hand luggage and uses standard molecular biology protocols and reagents that tolerate adverse environmental conditions. Barcodes are sequenced using MinION technology and analyzed with ONTrack, an original de novo assembly pipeline that requires as few as 1000 reads per sample. ONTrack-derived consensus barcodes have a high accuracy, ranging from 99.8 to 100%, despite the presence of homopolymer runs. The ONTrack pipeline has a user-friendly interface and returns consensus sequences in minutes. The remarkable accuracy and low computational demand of the ONTrack pipeline, together with the inexpensive equipment and simple protocols, make the proposed workflow particularly suitable for tracking species under field conditions

Neoephemeridae (Insecta, Ephemeroptera) of the Philippines and Insights on Southeast Asian Biodiversity

A new mayfly species, Potamanthellus panayensis sp. n. (Neoephemeridae) is described from Panay Island, Philippines, based on nymphs. Supplementary nymphal description of P. caenoides (Ulmer, 1939) and a provisional description of another Potamanthellus species from Mindanao Island, Philippines are provided. The potential underestimated diversity of Potamanthellus in Southeast Asia is discussed in line with the species delimitation in this study based on COI sequences

Integrative taxonomy of the genus Dudgeodes Sartori, 2008 (Insecta, Ephemeroptera, Teloganodidae) from the Philippines with description of new species and supplementary descriptions of Southeast Asian species

COI sequences were used as an initial clustering method to delimit putative species of the genus Dudgeodes in the Philippines. An overview of the diagnostic characters of Philippine species and characters with high intraspecific variability are given. Six new species of Dudgeodes are described and illustrated: D. bauernfeindi Garces & Sartori, sp. nov., D. freitagi Garces & Sartori, sp. nov., D. luntian Garces & Sartori, sp. nov., D. pangantihoni Garces & Sartori, sp. nov., D. tabang Garces & Sartori, sp. nov., and D. vonrinteleni Garces & Sartori, sp. nov., all known from the nymphal stage. Supplementary descriptions are provided for D. pescadori Sartori, 2008, D. hutanis Sartori, 2008, D. stephani Sartori, 2008, D. ulmeri Sartori, 2008, and D. celebensis Sartori, 2008. A key to the nymphs of Philippine Dudgeodes species is proposed

Procerobaetis petersorum Kaltenbach & Garces & Gattolliat 2020, gen. et sp. nov.

<i>Procerobaetis petersorum</i> Kaltenbach & Gattolliat gen. et sp. nov. <p>urn:lsid:zoobank.org:act: 8E7DA59C-2AB6-4518-BE84-5C71C5A5BD03</p> <p>Figs 5–7, 12 C–D, 14B</p> Diagnosis: larva <p>The diagnostic characters of all species are summarized in Table 3.</p> Etymology <p> <i>P. petersorum</i> gen. et sp. nov. is dedicated to the well-known mayfly researchers Janice Peters and her late husband William L. Peters for their contribution to our knowledge of Leptophlebiidae in Southeast Asia. Specific reference is given to the remarkable gills of <i>Procerobaetis</i> gen. nov., which are similar to those of some species of Leptophlebiidae.</p> Material examined <p> <b>Holotype</b></p> <p>INDONESIA • larva; Sumatra, volcano Sago, Kobun River; 00°22.55′ S, 100°39.55′ E; 1095 m a.s.l.; 19 Mar. 2014; M. Gueuning leg.; on slide; voucher no.: GBIFCH 00421980; MZL.</p> <p> <b>Paratypes</b></p> <p>INDONESIA • 1 larva; same collection data as for holotype; on slide; voucher: GBIFCH 00421975; MZL • 3 larvae; same collection data as for holotype; in alcohol; vouchers: GBIFCH 00422930, GBIFCH 00422044, GBIFCH 00422751; MZL • 1 larva; Sumatra, volcano Sago, Simbukan River, 00°17.40′ S, 100°40.84′ E; 955 m a.s.l.; 18 Mar. 2014; M. Gueuning leg.; on slide; voucher: GBIFCH 00422039; MZL • 1 larva; Sumatra, volcano Sago, Simbukan River; 00°17.13′ S, 100°41.23′ E; 880 m a.s.l.; 18 Mar. 2014; M. Gueuning leg.; in alcohol; voucher: GBIFCH 00422000; MZL • 1 larva; Sumatra, volcano Sago, Tampo River; 00°22.33′ S, 100°39.55′ E; 960 m a.s.l.; 20 Mar. 2014; M. Gueuning leg.; on slide; voucher: GBIFCH 00421968; MZL • 4 larvae; same collection data as for preceding; in alcohol; vouchers: GBIFCH 00422045, GBIFCH 00421964, GBIFCH 00422002, GBIFCH 00422705; MZL • 2 larvae; Sumatra, volcano Sago, Kaligain River; 00°18.02′ S, 100°40.13′ E; 1040 m a.s.l.; 5 Apr. 2014; M. Gueuning leg.; on slides; vouchers: GBIFCH 00421988, GBIFCH 00422006; MZL • 3 larvae; same collection data as for preceding; in alcohol; vouchers: GBIFCH 00422613, GBIFCH 00422043, GBIFCH 00422003; MZL.</p> <p> <b>Description: larva</b> (Figs 5–7, 12 C–D)</p> <p>BODY LENGTH. 3.4–4.5 mm.</p> <p>COLOURATION (Fig. 12B). Head, thorax and abdomen dorsally brown, head and thorax with bright median dorsal suture. Head and thorax ventrally light brown, abdomen ventrally brown. Legs light brown, caudal filaments light brown without darker band or pattern.</p> <p> <b>Head</b></p> <p>ANTENNA (Fig. 7A). Flagellum with lanceolate spines at apex of each segment, longer at inner lateral margin, increasing in length distally until segment IX and decreasing thereafter.</p> <p>LABRUM (Fig. 5A). Rectangular, length 0.6× maximum width. Distal margin with medial emargination and a small process. Dorsally with long, fine, simple setae scattered over surface in proximal area; many long, stout, simple setae in anteromedial area, erratically distributed, not arranged in an arc. Ventrally with marginal row of setae composed of lateral and anterolateral long, feathered setae and medial long, bifid, pectinate setae; ventral surface with about five short, spine-like setae near lateral and anterolateral margin.</p> <p>RIGHT MANDIBLE (Fig. 5 B–C). Outer and inner set of denticles with 4+3 denticles. Margin between prostheca and mola straight, with a row of long, stout setae. Tuft of setae at apex of mola present.</p> <p>LEFT MANDIBLE (Fig. 5 D–E). Outer and inner set of denticles with 4+3 denticles. Subtriangular process long and slender, above level of area between prostheca and mola. Denticles of mola apically constricted. Setae at apex of mola absent. Both mandibles with lateral margins almost straight. Basal half with fine, simple setae scattered over dorsal surface.</p> <p>HYPOPHARYNX (Fig. 5F). Lingua shorter than superlingua, longer than broad, with medial tuft of long, stout setae. Superlingua distally almost straight, lateral margin rounded, with fine, long, simple setae along distal margin.</p> <p>MAXILLA (Fig. 5G). Galea-lacinia with two simple, robust apical setae under crown. Medially with one pectinate, spine-like seta and a row of 5–6 medium to long, simple setae. Maxillary palp 2× as long as length of galea-lacinia; palp segment II 0.7× length of segment I, palp segment III 0.7× length of segment II; setae on maxillary palp fine, simple, scattered over surface of segments I, II and III; apex of last segment rounded.</p> <p>LABIUM (Fig. 5H). Glossae basally broad, narrowing toward apex, slightly shorter than paraglossae; inner margin with 9–10 spine-like setae; apex with two long and one medium-length, robust, pectinate setae; outer margin with 5–7 spine-like setae, increasing in length distally; ventral surface with medium, fine, simple, scattered setae. Paraglossae sub-rectangular, apically curved inward; apex rounded, with three rows of long, robust setae; ventrally 1–3 medium, simple setae in anteromedial area; dorsally with a row of 5 long, spine-like setae near inner margin and several medium, fine, simple setae at outer margin. Labial palp with segment I 0.6× length of segments II and III combined, ventrally with scattered short, fine, simple setae; segment II with small distolateral expansion, ventrally with scattered short, fine, simple setae, dorsally with a row of 3–4 long, spine-like setae; segment III subquadrangular, apex rounded, ventrally covered with short, spine-like, simple setae and short, fine, simple setae. Mentum distally with scattered fine, simple setae.</p> <p> <b>Thorax</b></p> <p> FORELEG (Figs 6 A–H, 7B). Ratio of foreleg segments 1.4:1.0:1.0:0.3. <i>Femur</i>. Length 3.1–3.8× maximum width; dorsal margin with a row of 7–9 curved, spine-like setae; length of setae 0.26× maximum width of femur; apex rounded, with one pair of laterally slightly pectinate, spine-like setae; many stout, lanceolate, laterally pectinate setae scattered along ventral margin; femoral patch absent. <i>Tibia.</i> Dorsal margin bare; ventral margin with a row of curved, laterally pectinate, spine-like setae, at apex some longer, laterally pectinate, spine-like setae; anterior surface with scattered stout, lanceolate, laterally pectinate setae; patellotibial suture present on basal ⅓. <i>Tarsus.</i> Dorsal margin bare; ventral margin with a row of curved, laterally pectinate, spine-like setae; tarsal claw elongate, slender, apically pointed, with one row of 4–7 larger denticles and many minute denticles, ventral margin at apex undulated, with many stripes.</p> <p>MIDDLE LEG (Fig. 7C). As foreleg, but dorsal margin of femur almost straight, tibia at dorsal margin with a row of stout setae, and tarsal claw with one row of 3–6 larger denticles and many minute denticles.</p> <p>HIND LEG (Fig. 7D). As foreleg, but dorsal margin of femur almost straight, tarsal claw with one row of 4–7 larger denticles and many minute denticles.</p> <p> <b>Abdomen</b></p> <p>TERGITES (Fig. 6I). Surface with scattered scales, U-shaped scale bases and micropores. Posterior margin of tergites I–VIII without spines, tergite IX with triangular spines.</p> <p>GILLS (Fig. 7 E–F). Present on segments I–VII; elongate, with pronounced points; gills I and II extremely slender and with very long, extended points; margin with very small denticles intercalating fine, simple setae; tracheae limited to main trunk. Gill II as long as length of segments III and ½ of IV combined, gill III as long as length of segments IV and ½ of V combined, gill IV as long as length of segments V and ½ of VI combined, gill V as long as length of segments VI and ⅔ of VII combined, gill VI as long as length of segments VII and VIII combined, gill VII as long as length of segments VIII and ⅔ of IX combined.</p> <p>PARAPROCT (Fig. 6 J–K). Posterior margin with 8–10 stout spines; surface with scattered U-shaped scale bases and micropores; posterolateral extension (cercotractor) with numerous small, marginal spines.</p> <p>CAUDAL FILAMENTS (Fig. 12C). Cerci ca 0.5× body length.</p> Distribution <p>Indonesia: Sumatra (Fig. 14B).</p> Remarks <p>The specimens were collected at altitudes from 800 to 1100 m a.s.l. in medium to large, mainly shallow and slow to moderate fast streams (width 2–10 m, depth 20–150 cm, velocity 0.28–0.5 m /s, temperature 20–21°C, pH 7–7.5). The streams were mostly in forests with a partly closed canopy, the substrate was dominated by bedrock, boulders and stones with few patches of leaf litter or dead wood. One stream was in a more disturbed area (rice fields, waste and soap).</p>Published as part of <i>Kaltenbach, Thomas, Garces, Jhoana M. & Gattolliat, Jean-Luc, 2020, A new genus of Baetidae (Insecta, Ephemeroptera) from Southeast Asia, pp. 1-32 in European Journal of Taxonomy 612</i> on pages 12-17, DOI: 10.5852/ejt.2020.612, <a href="http://zenodo.org/record/3702176">http://zenodo.org/record/3702176</a&gt

Fig. 12. Habitus, larvae. A in A new genus of Baetidae (Insecta, Ephemeroptera) from Southeast Asia

Fig. 12. Habitus, larvae. A. Procerobaetis leptobranchius gen. et sp. nov., dorsal view. B. P. leptobranchius gen. et sp. nov., lateral view. C. P. petersorum gen. et sp. nov., dorsal view. D. P. petersorum gen. et sp. nov., lateral view. Scale bars: 1 mm.Published as part of Kaltenbach, Thomas, Garces, Jhoana M. & Gattolliat, Jean-Luc, 2020, A new genus of Baetidae (Insecta, Ephemeroptera) from Southeast Asia, pp. 1-32 in European Journal of Taxonomy 612 on page 26, DOI: 10.5852/ejt.2020.612, http://zenodo.org/record/370217

Procerobaetis leptobranchius Kaltenbach & Garces & Gattolliat 2020, gen. et sp. nov.

<i>Procerobaetis leptobranchius</i> Kaltenbach & Gattolliat gen. et sp. nov. <p>urn:lsid:zoobank.org:act: B05740B0-8BC1-4F5B-9F32-6852C54B9EB1</p> <p>Figs 1–4, 12 A–B, 14B</p> Diagnosis: larva <p>The diagnostic characters of all species are summarized in Table 3.</p> Etymology <p> <i>Procerobaetis leptobranchius</i> gen. et sp. nov. is a combination of <i>lepto</i> – derived from the Greek word ʻleptosʼ, meaning ʻfineʼ, ʻthinʼ, ʻslenderʼ – and <i>branchius</i>, derived from the Latin word branchiae, meaning ʻgillsʼ, in reference to the remarkably elongate and long-pointed gills I and II, which are typical for the genus. It is also in reference to Leptophlebiidae, where this type of gill partly exists as well.</p> Material examined <p> <b>Holotype</b></p> <p>INDONESIA • larva; Sumatra, volcano Singgalang, Caruak River; 00°23.06′ S, 100°21.41′ E; 1640 m a.s.l.; 23 Mar. 2014; M. Gueuning leg.; on slide; voucher: GBIFCH 00421984; MZL.</p> <p> <b>Paratypes</b></p> <p>INDONESIA • 4 larvae; same collection data as for holotype; on slides; vouchers: GBIFCH 00422019, GBIFCH 00422023, GBIFCH 00421977, GBIFCH 00421989; MZL • 5 larvae; same collection data as for holotype; in alcohol; vouchers: GBIFCH 00421983, GBIFCH 00422974 (3 larvae), GBIFCH 00421986; MZL • 1 larva; Sumatra, volcano Singgalang, Pagu Pagu River; 00°23.54′ S, 100°21.46′ E; 1785 m a.s.l.; 22 Mar. 2014; M. Gueuning leg.; on slide; voucher: GBIFCH 00421957; MZL.</p> <p> <b>Description: larva</b> (Figs 1–4, 12 A–B)</p> <p>BODY LENGTH. 4.6–5.5 mm.</p> <p>COLOURATION (Fig. 12A). Head, thorax and abdomen dorsally dark brown, head and thorax with bright median dorsal suture. Head and thorax ventrally brown, abdomen ventrally dark brown. Legs light brown, caudal filaments light brown without darker band or pattern.</p> <p> <b>Head</b></p> <p>ANTENNA (Fig. 3A). Approximately 2× as long as head length; flagellum with lanceolate spines at apex of each segment, longer at inner lateral margin, increasing in length distally until segment IX and decreasing thereafter.</p> <p>LABRUM (Fig. 1A). Rectangular, length 0.7× maximum width. Distal margin with medial emargination and a small process. Dorsally with long, fine, simple setae scattered over surface in proximal area; many long, stout, simple setae in anteromedial area, erratically distributed, not arranged in an arc. Ventrally with marginal row of setae composed of lateral and anterolateral long, feathered setae and medial long, bifid setae; ventral surface with about ten short, spine-like setae near lateral and anterolateral margins.</p> <p>RIGHT MANDIBLE (Fig. 1 B–C). Outer and inner set of denticles with 4+3 denticles. Margin between prostheca and mola straight, with a row of long, stout setae. Tuft of setae at apex of mola present.</p> <p>LEFT MANDIBLE (Fig. 1 D–E). Outer and inner set of denticles with 4+3 denticles. Subtriangular process long and slender, above level of area between prostheca and mola. Denticles of mola apically constricted. Setae at apex of mola absent. Both mandibles with lateral margins slightly convex. Basal half with fine, simple setae scattered over dorsal surface.</p> <p>HYPOPHARYNX (Fig. 1F). Lingua shorter than superlingua, about as broad as long, with medial tuft of long, stout setae. Superlingua distally almost straight, lateral margin rounded, with fine, long, simple setae along distal and distolateral margin.</p> <p>MAXILLA (Fig. 1G). Galea-lacinia with two simple, robust apical setae under crown. Medially with one pectinate, spine-like seta and a row of 6–8 medium to long, simple setae. Maxillary palp 2× as long as length of galea-lacinia; palp segment II 0.7× length of segment I, palp segment III 0.6× length of segment II; setae on maxillary palp fine, simple, scattered over surface of segments I, II and III; apex of last segment rounded.</p> <p>LABIUM (Fig. 1H). Glossae basally broad, narrowing toward apex, slightly shorter than paraglossae; inner margin with 13–15 spine-like setae; apex with two long, robust, pectinate setae and one medium, robust, pectinate seta; outer margin with eight spine-like setae, increasing in length distally; ventral surface with medium, fine, simple, scattered setae. Paraglossae sub-rectangular, apically curved inward; apex rounded, with three rows of long, robust setae; ventrally four medium, simple setae in anteromedial area; dorsally with a row of 6–7 long, spine-like setae near inner margin and several medium, fine, simple setae at outer margin. Labial palp with segment I 0.9× length of segments II and III combined, ventrally with scattered short, fine, simple setae; segment II with small distolateral expansion, ventrally with scattered short, fine, simple setae, dorsally with a row of 4–5 long, spine-like setae; segment III subquadrangular, apex rounded, ventrally covered with short and medium spine-like, simple setae and short, fine, simple setae. Mentum medially and distally with scattered fine, simple setae.</p> <p> <b>Thorax</b></p> <p> FORELEG (Figs 2 A–F, 4A). Ratio of foreleg segments 1.3:1.0:0.9:0.3. <i>Femur</i>. Length 4.2–4.7× maximum width; dorsal margin with a row of 7–9 curved, spine-like setae; length of setae 0.3× maximum width of femur; apex rounded, with one pair of laterally slightly pectinate, spine-like setae; many stout, lanceolate, laterally pectinate setae scattered along ventral margin; femoral patch absent. <i>Tibia.</i> Dorsal margin bare; ventral margin with a row of curved, laterally pectinate, spine-like setae, on apex some longer, laterally pectinate, spine-like setae; anterior surface with scattered stout, lanceolate, laterally pectinate setae; patellotibial suture present on basal ⅓. <i>Tarsus.</i> Dorsal margin bare; ventral margin with a row of curved, laterally pectinate, spine-like setae; tarsal claw elongate, slender, apically pointed, with one row of 7–8 larger denticles and many minute denticles, ventral margin at apex undulated, with many stripes.</p> <p>MIDDLE LEG (Fig. 4B). As foreleg, but dorsal margin of femur slightly concave, tibia at dorsal margin with a row of stout setae, and tarsal claw with one row of 5–11 larger denticles and many minute denticles.</p> <p>HIND LEG (Fig. 4C). As foreleg, but dorsal margin of femur slightly concave, and tarsal claw with one row of 7–11 larger denticles and many minute denticles.</p> <p> <b>Abdomen</b></p> <p>TERGITES (Fig. 2G). Surface with scattered scales and U-shaped scale bases. Posterior margin of tergites I–VIII without spines, tergite IX with triangular spines.</p> <p>GILLS (Fig. 3 B–H). Present on segments I–VII; elongate with pronounced points; gills I and II extremely slender and with very long, extended points; margin with very small denticles intercalating fine, simple setae; tracheae limited to main trunk. Gill I as long as length of segments II and ¾ of III combined, gill II as long as length of segments III and IV combined, gill III as long as length of segments IV and ¾ of V combined, gill IV as long as length of segments V and VI combined, gill V as long as length of segments VI and VII combined, gill VI as long as length of segments VII to ½ of IX combined, gill VII as long as length of segments VIII to X combined.</p> <p>PARAPROCT (Fig. 2H). Posterior margin with 5–9 stout spines; surface with scattered U-shaped scale bases, micropores and a few fine, simple setae; posterolateral extension (cercotractor) with numerous small, marginal spines.</p> <p>CAUDAL FILAMENTS (Fig. 12A). Cerci ca 0.4× body length.</p> Distribution <p>Indonesia: Sumatra (Fig. 14B).</p> Remarks <p>The specimens were collected at altitudes from 1600 to 1800 m a.s.l. in two small, shallow, slow and rather cool forest streams (width ca 1 m, depth 5–30 cm, velocity 0.15–0.2 m /s, temperature 15.5–17°C, pH 6–7) with partly to completely closed canopy. The substrate in both streams was dominated by boulders, stones and gravel with some sandy patches and with or without patches of leaf litter or dead wood.</p>Published as part of <i>Kaltenbach, Thomas, Garces, Jhoana M. & Gattolliat, Jean-Luc, 2020, A new genus of Baetidae (Insecta, Ephemeroptera) from Southeast Asia, pp. 1-32 in European Journal of Taxonomy 612</i> on pages 6-11, DOI: 10.5852/ejt.2020.612, <a href="http://zenodo.org/record/3702176">http://zenodo.org/record/3702176</a&gt