Further evidence for low-energy protonium production in vacuum

We describe an experiment performed in the ATHENA apparatus in which there is evidence that the antiproton-proton bound state, protonium, has been produced at very low energies in vacuum following the interaction of cold antiprotons with a trapped cloud of molecular hydrogen ions. The latter were confined in a centrifugally separated belt outside a positron plasma used for antihydrogen formation. Studies have been performed at low positron plasma temperatures in which the protonium annihilation signal has been identified along with that from antihydrogen, and we discuss how their contributions can be disentangled. With the positron plasma heated to around 10000K the ions become distributed in the positrons, and the majority of the annihilation signal can be explained in terms of protonium formation, as antihydrogen creation is heavily suppressed. In this case we compare the observed protonium formation ratewith expectations from theory and find reasonable accord, when experimental systematics are taken into account. The effect on the annihilation signals of the passage of an electron current through a pre-loaded positron plasma has been studied in detail, and the results are presented here for the first time

Development of predator defences in fishes

A variety of development characteristics, morphological, behavioural, and experiential, contribute to the extreme vulnerability of young fishes to predation. The influence of these characteristics is complicated by the fact that the larval period is one of substantial and rapid change. Yet survival is the ultimate goal;-it is only by reaching maturity that individual fish have the opportunity to reproduce. With such high stakes it is not surprising that predator defences are of major importance during all phases of life. Developmental constraints may limit the defensive options for young fishes. Avoidance behaviours, which reduce the likelihood of encountering a predator or of being attacked by it, are particulaly evident in the youngest stages. Here size, coloration and dispersal are used to help elude the predator's attention. As fishes grow and acquire greater morphological and behavioural sophistication, there is more scope for predator evasion when avoidance fails. Older fishes are increasingly able to respond to external stimuli and can detect and react to predators or join conspecifics in common defence (schooling). Behavioural development is not simply a consequence of growth and the concomitant physical alterations of the body; it is also mediated by experience that comes through interaction with the physical and biotic environment. Predispositions to respond to experience may be a product of evolutionary history. Although mortality rates decline markedly with development and maturity, changes in size or behaviour can render fishes vulnerable to new suites of predators. Effective predator avoidance can compromise other activities, such as foraging, and individuals may be forced to reconcile conflicting demands. Developmental niche shifts that occur, for example, when certain size classes take refuge in less profitable feeding habitats, represent one such trade-off. Niche shifts may also be mediated by the influence of the programme for morphological development on sensory or behavioural capabilities. In addition to all of these developmental consderations, natural variations in environmental conditions - such as temperature, photoperiod, predator density and variety, and presence of alternative prey - represent additional challenges to predator defences during the rite of passage from birth to reproduction.</p