The Nature of the Peculiar Virgo Cluster Galaxies NGC 4064 and NGC 4424

We present a detailed study of the peculiar HI-deficient Virgo cluster spiral galaxies NGC 4064 and NGC 4424, using $^{12}$CO 1-0 interferometry, optical imaging and integral-field spectroscopic observations, in order to learn what type of environmental interactions have afected these galaxies. Optical imaging reveals that NGC 4424 has a strongly disturbed stellar disk, with banana-shaped isophotes and shells. NGC 4064, which lies in the cluster outskirts, possesses a relatively undisturbed outer stellar disk and a central bar. In both galaxies H-alpha emission is confined to the central kiloparsec. CO observations reveal bilobal molecular gas morphologies, with H-alpha emission peaking inside the CO lobes, implying a time sequence in the star formation process.Gas kinematics reveals strong bar-like non-circular motions in the molecular gas in both galaxies, suggesting that the material is radially infalling. In NGC 4064 the stellar kinematics reveal strong bar-like non-circular motions in the central 1 kpc. On the other hand, NGC 4424 has extremely modest stellar rotation velocities (Vmax ~ 30 km s-1), and stars are supported by random motions as far out as we can measure it. The observations suggest that the peculiarities of NGC 4424 are the result of an intermediate-mass merger plus ram pressure stripping. In the case of NGC 4064, the evidence suggests an already stripped "truncated/normal" galaxy that recently suffered a minor merger or tidal interaction with another galaxy. We propose that galaxies with "truncated/compact" H-alpha morphologies such as these are the result of the independent effects of ram pressure stripping, which removes gas from the outer disk, and gravitational interactions such as mergers, which heat stellar disks, drive gas to the central kpc and increase the central mass concentrations.Comment: 42 pages, 21 figure

The Trouble with Hubble Types in the Virgo Cluster

Quantitative measures of central light concentration and star formation activity are derived from R and Halpha surface photometry of 84 bright S0-Scd Virgo Cluster and isolated spiral galaxies. For isolated spirals, there is a good correlation between these two parameters and assigned Hubble types. In the Virgo Cluster, the correlation between central light concentration and star formation activity is significantly weaker. Virgo Cluster spirals have systematically reduced global star formation with respect to isolated spirals, with severe reduction in the outer disk, but normal or enhanced activity in the inner disk. Assigned Hubble types are thus inadequate to describe the range in morphologies of bright Virgo Cluster spirals. In particular, spirals with reduced global star formation activity are often assigned misleading early-type classifications, irrespective of their central light concentrations. 45+-25% of the galaxies classified as Sa in the Virgo Cluster sample have central light concentrations more characteristic of isolated Sb-Sc galaxies. The misleading classification of low concentration galaxies with low star formation rates as early-type spirals may account for part of the excess of `early-type' spiral galaxies in clusters. Thus the morphology-density relationship is not all due to a systematic increase in the bulge-to-disk ratio with environmental density.Comment: Accepted for publication in ApJ Letters. Latex aaspp4.sty, 9 pages, 2 Postscript (embedded) figures. Also available at http://www.astro.yale.edu/koopmann/preprint.htm

A two-step method for identifying photopigment opsin and rhodopsin gene sequences underlying human color vision phenotypes.

PurposeTo present a detailed, reliable long range-PCR and sequencing (LR-PCR-Seq) procedure to identify human opsin gene sequences for variations in the long wavelength-sensitive (OPN1LW), medium wavelength-sensitive (OPN1MW), short wavelength-sensitive (OPN1SW), and rhodopsin (RHO) genes.MethodsColor vision was assessed for nine subjects using the Farnsworth-Munsell 100 hue test, Ishihara pseudoisochromatic plates, and the Rabin cone-contrast threshold procedure (ColorDX, Konan Medical). The color vision phenotypes were normal trichromacy (n = 3), potential tetrachromacy (n = 3), dichromacy (n = 2), and unexplained low color vision (n = 1). DNA was isolated from blood or saliva and LR-PCR amplified into individual products: OPN1LW (4,045 bp), OPN1MW (4,045 bp), OPN1SW (3,326 bp), and RHO (6,715 bp). Each product was sequenced using specific internal primer sets. Analysis was performed with Mutation Surveyor software.ResultsThe LR-PCR-Seq technique identified known single nucleotide polymorphisms (SNPs) in OPN1LW and OPN1MW gene codons (180, 230, 233, 277, and 285), as well as those for lesser studied codons (174, 178, 236, 274, 279, 298 and 309) in the OPN1LW and OPN1MW genes. Additionally, six SNP variants in the OPN1MW and OPN1LW genes not previously reported in the NCBI dbSNP database were identified. An unreported poly-T region within intron 5(c.36+126) of the rhodopsin gene was also found, and analysis showed it to be highly conserved in mammalian species.ConclusionsThis LR-PCR-Seq procedure (single PCR reaction per gene followed by sequencing) can identify exonic and intronic SNP variants in OPN1LW, OPN1MW, OPN1SW, and rhodopsin genes. There is no need for restriction enzyme digestion or multiple PCR steps that can introduce errors. Future studies will combine the LR-PCR-Seq with perceptual behavior measures, allowing for accurate correlations between opsin genotypes, retinal photopigment phenotypes, and color perception behaviors

A dynamical model for the heavily ram pressure stripped Virgo spiral galaxy NGC 4522

A dynamical model including ram pressure stripping is applied to the strongly HI deficient Virgo spiral galaxy NGC 4522. A carefully chosen model snapshot is compared with existing VLA HI observations. The model successfully reproduces the large-scale gas distribution and the velocity field. However it fails to reproduce the large observed HI linewidths in the extraplanar component, for which we give possible explanations. In a second step, we solve the induction equation on the velocity fields of the dynamical model and calculate the large scale magnetic field. Assuming a Gaussian distribution of relativistic electrons we obtain the distribution of polarized radio continuum emission which is also compared with our VLA observations at 6 cm. The observed maximum of the polarized radio continuum emission is successfully reproduced. Our model suggests that the ram pressure maximum occurred only ~50 Myr ago. Since NGC 4522 is located far away from the cluster center (~1 Mpc) where the intracluster medium density is too low to cause the observed stripping if the intracluster medium is static and smooth, two scenarios are envisaged: (i) the galaxy moves very rapidly within the intracluster medium and is not even bound to the cluster; in this case the galaxy has just passed the region of highest intracluster medium density; (ii) the intracluster medium is not static but moving due to the infall of the M49 group of galaxies. In this case the galaxy has just passed the region of highest intracluster medium velocity. This study shows the strength of combining high resolution HI and polarized radio continuum emission with detailed numerical modeling of the evolution of the gas and the large-scale magnetic field.Comment: 15 pages, 11 figures, accepted for publication in A&

Ongoing Gas Stripping in the Virgo Cluster Spiral NGC 4522

The Virgo cluster galaxy NGC 4522 is one of the best spiral candidates for ICM-ISM stripping in action. Optical broadband and H-alpha images from the WIYN telescope of the highly inclined galaxy reveal a relatively undisturbed stellar disk and a peculiar distribution of H-alpha emission. Ten percent of the H-alpha emission arises from extraplanar HII regions which appear to lie within filamentary structures >3 kpc long above one side of the disk. The filaments emerge from the outer edge of a disk of bright H-alpha emission which is abruptly truncated beyond 0.35R(25). Together the truncated H-alpha disk and extraplanar H-alpha filaments are reminiscent of a bow shock morphology, which strongly suggests that the interstellar medium (ISM) of NGC 4522 is being stripped by the gas pressure of the intracluster medium (ICM). The galaxy has a line-of-sight velocity of 1300 km/sec with respect to the mean Virgo cluster velocity, and thus is expected to experience a strong interaction with the intracluster gas. The existence of HII regions apparently located above the disk plane suggests that star formation is occuring in the stripped gas, and that newly formed stars will enter the galaxy halo and/or intracluster space. The absence of HII regions in the disk beyond 0.35R(25), and the existence of HII regions in the stripped gas suggest that even molecular gas has been effectively removed from the disk of the galaxy.Comment: to appear in The Astronomical Journal, 16 pages, 5 figures, 1 tabl

Large-scale radio continuum properties of 19 Virgo cluster galaxies The influence of tidal interactions, ram pressure stripping, and accreting gas envelopes

Deep scaled array VLA 20 and 6cm observations including polarization of 19 Virgo spirals are presented. This sample contains 6 galaxies with a global minimum of 20cm polarized emission at the receding side of the galactic disk and quadrupolar type large-scale magnetic fields. In the new sample no additional case of a ram-pressure stripped spiral galaxy with an asymmetric ridge of polarized radio continuum emission was found. In the absence of a close companion, a truncated HI disk, together with a ridge of polarized radio continuum emission at the outer edge of the HI disk, is a signpost of ram pressure stripping. 6 out of the 19 observed galaxies display asymmetric 6cm polarized emission distributions. Three galaxies belong to tidally interacting pairs, two galaxies host huge accreting HI envelopes, and one galaxy had a recent minor merger. Tidal interactions and accreting gas envelopes can lead to compression and shear motions which enhance the polarized radio continuum emission. In addition, galaxies with low average star formation rate per unit area have a low average degree of polarization. Shear or compression motions can enhance the degree of polarization. The average degree of polarization of tidally interacting galaxies is generally lower than expected for a given rotation velocity and star formation activity. This low average degree of polarization is at least partly due to the absence of polarized emission from the thin disk. Ram pressure stripping can decrease whereas tidal interactions most frequently decreases the average degree of polarization of Virgo spiral galaxies. We found that moderate active ram pressure stripping has no influence on the spectral index, but enhances the global radio continuum emission with respect to the FIR emission, while an accreting gas envelope can but not necessarily enhances the radio continuum emission with respect to the FIR emission.Comment: 37 pages, 26 figures, accepted for publication in A&

Deciphering the radio-star formation correlation on kpc-scales III. Radio-dim and bright regions in spiral galaxies

The relation between the resolved star formation rate per unit area and the non-thermal radio continuum emission is studied in 21 Virgo cluster galaxies and the two nearby spiral galaxies, NGC6946 and M51. For the interpretation and understanding of our results we used a 3D model where star formation, 2D cosmic ray (CR) propagation, and the physics of synchrotron emission are included. Based on the linear correlation between the star formation rate per unit area and the synchrotron emission and its scatter radio-bright and radio-dim regions can be robustly defined for our sample of spiral galaxies. We identified CR diffusion or streaming as the physical causes of radio-bright regions of unperturbed symmetric spiral galaxies as NGC6946. We identified the probable causes of radio-bright regions in several galaxies as CR transport, via either gravitational tides (M51) or galactic winds (NGC4532) or ram pressure stripping (NGC4330 and NGC4522). Three galaxies are overall radio-dim: NGC4298, NGC4535, and NGC4567. Based on our model of synchrotron-emitting disks we suggest that the overall radio-dim galaxies have a significantly lower magnetic field than expected by equipartition between the magnetic and turbulent energy densities. Radio-bright regions frequently coincide with asymmetric ridges of polarized radio continuum emission, and we found a clear albeit moderate correlation between the polarized radio continuum emission and the radio/SFR ratio. When compression or shear motions of the interstellar medium (ISM) are present in the galactic disk, the radio-bright regions are linked to the commonly observed asymmetric ridges of polarized radio continuum emission and represent a useful tool for the interaction diagnostics. Based on our results, we propose a scenario for the interplay between star formation, CR electrons, and magnetic fields in spiral galaxies.Comment: 36 pages, accepted for publication in A&