Electric organ discharge diversity in the genus Gymnotus: anatomo-functional groups and electrogenic mechanisms

Previous studies describe six factors accounting for interspecific diversity of electric organ discharge (EOD) waveforms in Gymnotus. At the cellular level, three factors determine the locally generated waveforms: (1) electrocyte geometry and channel repertoire; (2) the localization of synaptic contacts on electrocyte surfaces; and (3) electric activity of electromotor axons preceding the discharge of electrocytes. At the organismic level, three factors determine the integration of the EOD as a behavioral unit: (4) the distribution of different types of electrocytes and specialized passive tissue forming the electric organ (EO); (5) the neural mechanisms of electrocyte discharge coordination; and (6) post-effector mechanisms. Here, we reconfirm the importance of the first five of these factors based on comparative studies of a wider diversity of Gymnotus than previously investigated. Additionally, we report a hitherto unseen aspect of EOD diversity in Gymnotus. The central region of the EO (which has the largest weight on the conspecific-received field) usually exhibits a negative-positive-negative pattern where the delay between the early negative and positive peaks (determined by neural coordination mechanisms) matches the delay between the positive and late negative peaks (determined by electrocyte responsiveness). Because delays between peaks typically determine the peak power frequency, this matching implies a co-evolution of neural and myogenic coordination mechanisms in determining the spectral specificity of the intraspecific communication channel. Finally, we define four functional species groups based on EO/EOD structure. The first three exhibit a heterogeneous EO in which doubly innervated electrocytes are responsible for a main triphasic complex. Group I species exhibit a characteristic cephalic extension of the EO. Group II species exhibit an early positive component of putative neural origin, and strong EO auto-excitability. Group III species exhibit an early, slow, negative wave of abdominal origin, and variation in EO auto-excitability. Representatives of Group IV generate a unique waveform comprising a main positive peak followed by a small, load-dependent negative component

The Gemini Deep Deep Survey: II. Metals in Star-Forming Galaxies at Redshift 1.3<z<2

The goal of the Gemini Deep Deep Survey (GDDS) is to study an unbiased sample of K<20.6 galaxies in the redshift range 0.8<z<2.0. Here we determine the statistical properties of the heavy element enrichment in the interstellar medium (ISM) of a subsample of 13 galaxies with 1.34<z<1.97 and UV absolute magnitude M_2000 < -19.65. The sample contains 38% of the total number of identified galaxies in the first two fields of the survey with z>1.3. The selected objects have colors typical of irregular and Sbc galaxies. Strong [OII] emission indicates high star formation activity in the HII regions (SFR~13-106 M_sun/yr). The high S/N composite spectrum shows strong ISM MgII and FeII absorption, together with weak MnII and MgI lines. The FeII column density, derived using the curve of growth analysis, is logN_FeII = 15.54^{+0.23}_{-0.13}. This is considerably larger than typical values found in damped Ly-alpha systems (DLAs) along QSO sight lines, where only 10 out of 87 (~11%) have logN_FeII > 15.2. High FeII column densities are observed in the z=2.72 Lyman break galaxy cB58 (logN_FeII ~ 15.25) and in gamma-ray burst host galaxies (logN_FeII ~ 14.8-15.9). Given our measured FeII column density and assuming a moderate iron dust depletion (delta_Fe ~ 1 dex), we derive an optical dust extinction A_V ~ 0.6. If the HI column density is log N(HI)<21.7 (as in 98% of DLAs), then the mean metallicity is Z/Z_sun > 0.2. The high completeness of the GDDS sample implies that these results are typical of star-forming galaxies in the 1<z<2 redshift range, an epoch which has heretofore been particularly challenging for observational programs.Comment: ApJ in press, corrected HI column density estimat

Protostars and Outflows in the NGC7538 - IRS9 Cloud Core

New high resolution observations of HCO+ J=1-0, H13CN J=1-0, SO 2,2 - 1,1, and continuum with BIMA at 3.4 mm show that the NGC7538 - IRS9 cloud core is a site of active ongoing star formation. Our observations reveal at least three young bipolar molecular outflows, all ~ 10,000 -- 20,000 years old. IRS9 drives a bipolar, extreme high velocity outflow observed nearly pole on. South of IRS9 we find a cold, protostellar condensation with a size of ~ 14" x 6" with a mass > 250 Msun. This is the center of one of the outflows and shows deep, red-shifted self absorption in HCO+, suggesting that there is a protostar embedded in the core, still in a phase of active accretion. This source is not detected in the far infrared, suggesting that the luminosity < 10^4 Lsun; yet the mass of the outflow is ~ 60 Msun. The red-shifted HCO+ self-absorption profiles observed toward the southern protostar and IRS9 predict accretion rates of a few times 10^-4 to 10^-3 Msun/yr. Deep VLA continuum observations at 3.6 cm show that IRS9 coincides with a faint thermal VLA source, but no other young star in the IRS9 region has any detectable free-free emission at a level of ~ 60 microJy at 3.6 cm. The HCO+ abundance is significantly enhanced in the hot IRS9 outflow. A direct comparison of mass estimates from HCO+ and CO for the well-characterized red-shifted IRS9 outflow predicts an HCO+ enhancement of more than a factor of 30, or [HCO+/H2] >= 6 10^-8.Comment: 40 pages, 3 tables and 10 figures included; to appear in Ap

High Mass Star Formation I: The Mass Distribution of Submillimeter Clumps in NGC 7538

We present submillimeter continuum maps at 450 and 850 microns of a 12 x 8 arcminute region of the NGC 7538 high-mass star-forming region, made using the Submillimeter Common-User Bolometer Array (SCUBA) on the James Clerk Maxwell Telescope. We used an automated clump-finding algorithm to identify 67 clumps in the 450 micron image and 77 in the 850 micron image. Contrary to previous studies, we find a positive correlation between high spectral index, alpha, and high submillimeter flux, with the difference being accounted for by different treatments of the error beam. We interpret the higher spectral index at submillimeter peaks as a reflection of elevated dust temperature, particularly when there is an embedded infrared source, though it may also reflect changing dust properties. The clump mass-radius relationship is well-fit by a power law of the form M \propto R^(-x) with x = 1.5-2.1, consistent with theories of turbulently-supported clumps. According to our most reliable analysis, the high-mass end (approx. 100-2700 M_sun) of the submillimeter clump mass function in NGC 7538 follows a Salpeter-like power law with index 2.0 +/- 0.3. This result agrees well with similar studies of lower-mass regions rho Oph and Orion B. We interpret the apparent invariance of the shape of the clump mass function over a broad range of parent cloud masses as evidence for the self-similarity of the physical processes which determine it. This result is consistent with models which suggest that turbulent fragmentation, acting at early times, is sufficient to set the clump mass function.Comment: 37 pages, 7 figures, accepted for publication in Ap

The Gemini Deep Deep Survey: VIII. When Did Early-type Galaxies Form?

We have used the Hubble Space Telescope's Advanced Camera for Surveys (Ford et al. 2003) to measure the cumulative mass density in morphologically-selected early-type galaxies over the redshift range 0.8 < z < 1.7. Our imaging data set covers four well-separated sight-lines, and is roughly intermediate (in terms of both depth and area) between the GOODS/GEMS imaging data, and the images obtained in the Hubble Deep Field campaigns. Our images contain 144 galaxies with ultra-deep spectroscopy obtained as part of the Gemini Deep Deep Survey. These images have been analyzed using a new purpose-written morphological analysis code which improves the reliability of morphological classifications by adopting a 'quasi-Petrosian' image thresholding technique. We find that at z \~ 1 about 80% of the stars living in the most massive galaxies reside in early-type systems. This fraction is similar to that seen in the local Universe. However, we detect very rapid evolution in this fraction over the range 0.8 < z < 1.7, suggesting that over this redshift range the strong morphology-mass relationship seen in the nearby Universe is beginning to fall into place. By comparing our images to published spectroscopic classifications, we show that little ambiguity exists in connecting spectral classes to morphological classes for spectroscopically quiescent systems. However, the mass density function of early-type galaxies is evolving more rapidly than that of spectroscopically quiescent systems, which we take as further evidence that we are witnessing the formation of massive early-type galaxies over the 0.8 < z < 1.7 redshift range