Spreading recruitment over time to cope with environmental variability

10 paginas, 3 figuras, 2 tablas.-- El PDF es la versión de autor.Seedling establishment is one of the most vulnerable life cycle stages, and a key component for the population dynamics in short-lived plants. In unpredictable environments, timing of emergence is critical for the success of plant performance, and different adaptive bet-hedging strategies have evolved to reduce the risk of failure in recruitment. In this study we describe the spatio-temporal pattern of seedling emergence (overall rate and timing) and survival in four contrasting Mediterranean habitats for Plantago coronopus, a small herb with dimorphic seeds. We then explore the importance of spreading germination within years, as well as the role of the two types of seeds from a broader temporal perspective. Populations strongly differed for all recruitment components analyzed in a given year, but this spatial differentiation diluted when a longer period was considered. Apical (smaller) seeds germinated later and in a significantly lower proportion than basal (larger) seeds. Both late emergents and seedlings from apical seeds had lower survival probability in a rainy year. However, our results suggest that in a population having the lowest production of apical seeds, late emergents coming from apical seeds may constitute a large fraction of yearly recruitment and that their performance was non-significantly different from that of early emergents over the 4-year study period. This study provides evidence of the importance of two related traits (spreading seedling emergence through time by producing dimorphic seeds) as bet-hedging strategies to cope with environmental unpredictability. This is at least partly accomplished by increasing the potential of recruitment in favourable years, instead of buffering such important process in extremely bad years.This study was funded by the Spanish Ministry of Science, under projects BOS2002-01162 and CGL2006-08507 to MBG.Peer reviewe

Associating Turkey with the Transatlantic Trade and Investment Partnership: A costly (re‐) engagement?

Policy debate on the implications of the Transatlantic Trade and Investment Partnership (TTIP) for Turkey has focused almost exclusively on “how” Turkey can/will take part in a forthcoming transatlantic deal. Turkey's association with a TTIP has largely been conceived as an inevitable and beneficial policy choice to re‐engage Ankara with the Atlantic alliance and emerging transatlantic trade framework. The arguments for extending TTIP to Turkey have mostly been built upon a conventional understanding of preferential trade agreements. The debate has not provided a comprehensive assessment of costs and benefits for Turkey's exclusion from or joining TTIP as it dismissed multiple dimensions of the “deep integration” agenda which underpinned the transatlantic talks. This paper intends to contribute to the “why” debate with a thorough analysis of critical issues on the transatlantic agenda by evaluating economic and policy implications of TTIP both for exclusion and association scenarios together with associated compliance and adjustment costs

Un accord entre l'UE et les États-Unis sur le commerce et l'investissement affectera inévitablement la Suisse

Die gegenwärtigen Bemühungen für ein umfassendes Handels- und Investitionsschutzabkommen zwischen der EU und den USA werden nicht ohne Auswirkungen auf die Schweiz bleiben. Eine Studie des World Trade Institute der Universität Bern im Auftrag des Staatssekretariats für Wirtschaft analysiert die möglichen Auswirkungen eines Abkommens auf die Schweizer Wirtschaft und auf die Handelsbeziehungen der Schweiz. Unterschiedliche Szenarien führen dabei zu unterschiedlichen Auswirkungen

Numerical Investigation on Windback Seals Used in Aero Engines

Seals are considered one of the most important flow elements in turbomachinery applications. The most traditional and widely known seal is the labyrinth seal but in recent years other types like the brush or carbon seals were introduced since they considerably reduce the sealing air consumption. When seals are used for sealing of aero engine bearing chambers they are subjected to high “bombardment” through oil particles which are present in the bearing chamber. These particles mainly result from the bearings as a consequence of the high rotational speeds. Particularly when carbon or brush seals are used, problems with carbon formation (coking) may arise when oil gets trapped in the very tight gap of these seals. In order to prevent oil migration into the turbomachinery, particularly when the pressure difference over a seal is small or even negligible, significant improvement can be achieved through the introduction of so called windback seals. This seal has a row of static helical teeth (thread) and below this thread a scalloped or smooth shaft section is rotating. Depending on the application, a windback seal can be used alone or as a combination with another seal (carbon, brush or labyrinth seal). A CFD analysis carried out with ANSYS CFX version 11 is presented in this paper with the aim to investigate this seal type. The simulations were performed by assuming a two-phase flow of air and oil in the bearing compartment. Design parameters like seal clearance, thread size, scallop width, were investigated at different operating conditions

Thermal and flow phenomena associated with the behavior of brush seals in aero engine bearing chambers

Due to the increasing fuel cost and environmental targets, the demand for more efficient gas turbines has risen considerably in the last decade. One of the most important systems in a gas turbine is the secondary air system, which provides cooling air to the disks and to the blades. It also provides air for sealing of the bearing chambers. The amount of secondary air that is extracted from the compressor is a performance penalty for the engine. In aero engines, bearing chambers are in most cases sealed by the most traditional type of seal, the labyrinth seal. Bearing chambers contain the oil lubricated components like bearings and gears. In order to avoid oil migration from the bearing chamber into the turbomachinery, the seals are pressurized by secondary air; thus, a pressure difference is setup across the seal, which retains the lubricant into the bearing chamber. Oil loss can lead to a number of problems like oil fire or coking with the probability of an uncontained destruction of the aero engine. Oil fumes can also cause contamination of the air conditioning system of the aircraft thus cause discomfort to the passengers. Beside labyrinth seals, other types of seals such as brush seals and carbon seals are used. Both the latter are contact type seals, that is, they may be installed with zero gap and lift during operation when they get pressurized. Brush seals particularly may be installed having an overlap with the rotating part. An original aero engine bearing chamber was modified by MTU Aero Engines to run with brush seals in a simulating rig in Munich. Two types of brush seals were used for testing: (a) a brush seal with bristles made of Kevlar fibers and (b) a brush seal with bristles made of steel. Both types were installed with an overlap to the rotor. The targets set were twofold: (a) to measure the transient temperatures in the rotor and particularly in the contact zone between the bristles and the rotor and (b) to calculate the heat generation by the seals which could enable predictions of the heat generation in future applications (i.e. scaling to bigger rotor diameters). For the heat transfer calculations, numerical models using ANSYS CFX were created. Additionally, a coupled computational fluid dynamics (CFD) and finite element analysis (FEA) approach was applied to simulate flow and bristle's behavior. In order to obtain the transient temperature measurements with high fidelity, a new pyrometric technique was developed and was applied for the first time in brush seals as reported by Flouros et al. (2013, "Transient Temperature Measurements in the Contact Zone Between Brush Seals of Kevlar and Metallic Type for Bearing Chamber Sealing Using a Pyrometric Technique," ASME J. Gas Turbines Power, 135(8), p. 081603) and Flouros et al. (2012, "Transient Temperature Measurements in the Contact Zone Between Brush Seals of Kevlar and Metallic Type for Bearing Chamber Sealing Using a Pyrometric Technique," ASME Turbo Expo 2012, Copenhagen, Paper No. GT2012-68354). This technique has enabled positioning of the pyrometer (SensorthermGmbH, www.sensortherm.com) into the bristles pack of the seal adjacent to the rotating surface. The pyrometer could record the frictional temperature evolution in the bristles/rotor contact zone during accelerations or decelerations of the rotor. The sealing air demand can be reduced up to 97% with brush seals compared to traditional three fin labyrinth. It has been estimated that this can result in a reduction in fuel burned up to 1%. Further, the reduction in air flow has additional potential benefits such as a possible simplification of the bearing chamber architecture (vent less chamber). Even though the rotor was accelerated up to 19,500 rpm, the temperature induced overshoots in the seal/rotor contact zone have caused no deterioration in either the materials or the oil.SCOPUS: ar.jinfo:eu-repo/semantics/publishe