The Karyology of Tisbe Reluctans (Copepoda, Harpacticoida)

SUMMARYThe diploid chromosome number, 2 n = 24, has been established, in both sexes of the harpacticoid copepod Tisbe reluctans. The chromosomes are metacentric or submetacentric and differentiated sex chromosomes are not present. In the male chiasmatic bivalents form at metaphase I. In the female, during the first meiotic division and before the diffuse stage, homologous chromosomes are closely associated so that chiasmata may occur. After the diffuse stage, the homologues separate but remain parallel and juxtaposed to each other till metaphase I. The chromosomes seem to have doubled into two chromatids during the post-diffuse stage

Chromosomes of Thaliaceans

SUMMARYThe chromosomes of six species of thaliaceans have been examined. The haploid number 11 was determined for Pyrosoma atlanticum and Doliolum denticolatum; the haploid number 12 was found for Pegea confoederata, Salpa fusiformis, Thalia democratica; the haploid number 13 was found in Salpa maxima.Salpa fusiformis and Thalia democratica are strikingly akin with respect to both number and morphology of diakinetic chromosomes whereas the other species of Salpida all possess distinguishing peculiarities. Pegea confoederata takes a place apart among the thaliaceans by virtue of the minute dimension of its chromosomes.The karyology of thaliaceans appears to be well differenciated from that of other tunicates and, although ascidians and thaliaceans are the most akin to each other, it is not possible to establish a meaningful probable relationship between their chromosomes

A Squash Method for Chromosomes of Ascidians (Tunicata)

SUMMARYA squash method for chromosomes from living and preserved Ascidians (Tunicata) has been established

The lithofacies organization of fluvial channel deposits: a meta-analysis of modern rivers

Environmental interpretations of subsurface fluvial successions are commonly based on facies observations from core and are often attempted by generalist geologists by reference to classic facies models. However, for fluvial channel deposits, the value of observations on lithofacies proportions for interpretations of depositional environment has yet to be assessed quantitatively. Here, a test is presented that is based on a comparative study of facies data from 77 reaches of 46 modern rivers. The analysis is undertaken on datasets from published case studies stored in a sedimentological database, with consideration of causes for observational bias, and with particular attention paid to sandy lithofacies. The observed variability in the proportion of facies assemblages in the channel deposits of sandy river systems is quantified for classes of environments categorized according to channel pattern (braided, low sinuosity, meandering), climatic setting (arid to perhumid), and discharge regime (ephemeral to perennial). By capturing the variability in facies organization within fluvial systems of certain types, these outputs serve as facies models that provide a measure of uncertainty to sedimentological interpretations. Concurrently, the statistical analysis presented enables a test of the significance of relationships between the relative proportions of channel lithofacies and parameters that either represent controlling factors (e.g., water-discharge characteristics) or covariates (e.g., channel pattern). For classes of river systems grouped by channel pattern, climate, and discharge regime, emerging features of facies organization can be identified. Statistically, it is observed that relationships exist (i) between channel pattern and the frequency of the preserved expression of bedforms, and (ii) between controls on river hydrology (climate, discharge regime and seasonal variability) and the record of upper and lower flow-regime conditions. Thus, the results corroborate existing qualitative facies models in some respects. However, observations of the relative dominance of facies in channel deposits demonstrate limited value for interpretations or predictions in subsurface or outcrop studies, as variability within each type of depositional system is significant. Corehole data of fluvial channel deposits may be commonly overinterpreted

Chromosomes from Male Gonads of Acanthochiton Crinitus (Subclass Polyplacophora, Mollusca)

SUMMARYThe haploid number 9 of Acanthochiton crinitus, further confirmed on the diploid number 18, is at present the lowest chromosome number found in the whole subclass Polyplacophora. Spermatogenesis seems to be chiasmatic because of the presence of some cross-shaped bivalents. Mitotic chromosomes are mainly metacentric or submetacentric with a couple of subtelocentric. Spermatogonial chromosomes at metaphase are characterized by a distant parallel pairing of homologues. Differenciated sex-chromosomes seem to be absent

Chromosomes from Male Gonads of Haliotis Tubercolata and Haliotis Lamellosa (Haliotidae, Archeogasteropoda, Mollusca)

SUMMARYThe diploid number 28 was counted for Haliotis tubercolata and the haploid number 14 has been determined for H. lamellosa and H. tubercolata. Spermatocyte bivalents are very similar in both species and are characteristic for the presence of very few chiasmata

On the geological significance of clastic parasequences

Parasequences recognized in clastic sedimentary successions of shallow-marine origin are considered by some geologists to be the fundamental building blocks of depositional sequences, even though problems in their definition and application have been identified by others, who instead advocate their abandonment as formal sequence stratigraphic units. To elucidate the geological significance of clastic parasequences and inform the debate on their use in stratigraphy, a quantitative characterization of the geometry, facies characteristics and timescale of deposition of 1163 parasequences has been undertaken based on a synthesis of data from outcrop and subsurface studies that are available in the scientific literature. Through a database compilation, the attributes of the studied parasequences are analysed with respect to the interpreted geological origin of the units, and with consideration of sources of bias and uncertainty. Particular emphasis is placed on assessing the following: (i) the importance of heuristics, and of data types and coverage in the recognition of parasequences; (ii) differences in parasequence characteristics observed across deltaic and shoreface depositional systems, and between the Quaternary and the ancient rock record; (iii) possible explanations for the range in timescales of deposition of parasequences; and (iv) the role of autogenic dynamics on the development of deltaic parasequences, partly based on a comparison with the recent evolution of modern deltas. The results demonstrate that parasequence definition and physical correlation suffer from subjectivity, and that significant variability exists in the spatio-temporal and architectural attributes of clastic parasequences. This gives rise to uncertainty that affects the use of parasequences as a framework for comparison of the architecture of packages of strata originating via shoreline regression: this uncertainty must be considered when using analogue data for subsurface predictions or when attempting comparative studies of clastic successions

Assessment of backwater controls on the architecture of distributary channel fills in a tide-influenced coastal-plain succession: Campanian Neslen Formation, USA

The backwater zone of a river is its distal reach downstream of the point at which the streambed elevation reaches sea level. Backwater hydraulics is believed to exert an important control on fluvio-deltaic morphodynamics, but the expressions with which this may be recorded in the preserved stratigraphic record are not well understood. The seaward reaches of modern rivers can undergo flow acceleration and become erosional at high discharges due to drawdown of the in-channel water surface near the river mouth, in relation to the fixed water surface at the shoreline. As coastal-plain distributary channels approach the shoreline they tend to be subject to a reduction in lateral mobility, which could be related to diminished sediment flux at low flow. Current understanding of channel morphodynamics associated with backwater effects, as based on observations from numerical models and modern sedimentary systems, is here used to make predictions concerning the architecture of coastal distributary channel fills in the rock record. On the basis of existing knowledge, distributary channel fills are predicted to be typically characterized by low width-to-thickness aspect ratios, by a clustering of scour surfaces toward their base, by an aggradational infill style, by a facies organization that bears evidence of drawdown-influenced scour filling, possibly resulting in the overprint of tidal signals toward their base, and by co-genetic sand-prone overbank units of limited occurrence, thickness and sand content. To test these predictions, fieldwork was carried out to examine sedimentological characters of channel bodies from an interval of the Campanian Neslen Formation (eastern Utah, USA), which comprises a succession of sandstone, carbonaceous mudstone, and coal, deposited in a coastal-plain setting, and in which significant evidence of tidal influence is preserved. Three types of channel bodies are recognized in the studied interval, in terms of lithology and formative-channel morphodynamics: sand-prone laterally accreting channel elements, heterolithic laterally accreting channel elements and sand-prone aggradational ribbon channel elements. This study concentrates on the ribbon channel bodies since they possess a geometry compatible with laterally stable distributaries developed in the zone of drawdown. Sedimentological and architectural characteristics of these bodies are analyzed and compared with the proposed model of distributary channel-fill architecture. Although conclusive evidence of the influence of backwater processes in controlling the facies architecture of distributary channel fills is not reached, the studied bodies display an ensemble of internal architecture, lithological organization, nature of bounding surfaces and relationships with other units that conforms to the proposed model to a certain extent. The analyzed ribbon sandbodies are all characterized by erosional cut-banks, very limited proportions of mudstone deposits, a lack of genetically related barform units, clustering of scour fills at their base, and a lack of relationships with co-genetic river-fed overbank sandstones. This work provides a guide to future research, which is required to better understand the role of backwater processes in controlling the architecture of distributary channel bodies, their down-dip variations, and how these are expressed in the stratigraphic evolution of prograding coastal plains

FRG-SMRG Coastal rivers avulsion dataset

The dataset includes: (1) measures of avulsion frequency based on the avulsion histories of 57 coastal-plain river systems, based on numbers of avulsion events, active or abandoned channel courses, and delta lobes, all considered per unit time; additional sets of avulsion-frequency proxies are based on normalization of these numbers relative to the size of study areas and to the number of distinct river systems that drain into them. (2) variables detailing some of the environmental boundary conditions and depositional contexts of the case studies. (3) metadata

Quantitative analysis of the stratigraphic architecture of incised-valley fills: a global comparison of Quaternary systems

Facies models of the internal fills of incised valleys developed in shelf and coastal settings during cycles of relative sea-level change are largely conceptual, descriptive and qualitative in form; moreover, they are commonly bespoke to individual examples. Here, a database-driven quantitative statistical analysis of 87 late-Quaternary incised-valley fills (IVFs) has been undertaken to assess the general validity and predictive value of classical facies models for IVFs, and to investigate the relative importance of possible controls on their stratigraphic organization. Based on datasets from the published literature stored in a sedimentological database, the geometry and proportion of systems tracts, and of architectural elements of different hierarchies within IVFs are quantified. These variables were analysed to assess how they vary in relation to parameters that represent potential controlling factors: relative sea-level stage, continental-margin type, drainage-basin area, valley geometry, basin physiography and shoreline hydrodynamics. The stratigraphic organization of the studied coastal-plain IVFs is generally consistent with that represented in facies models, the primary control being the rate and magnitude of relative sea-level change. However, results from this study demonstrate significant variability in the stratigraphic architectures of IVFs, which is not accounted for by existing models. Variations in the facies architecture of coastal-plain and cross-shelf valley fills can be attributed to controls other than sea level, and expressed in relationships with continental-margin type, basin physiography, catchment area, river-system size and shoreline hydrodynamics. The following primary findings arise from this research. (i) Compared to their counterparts on passive margins, coastal-plain IVFs hosted on active margins contain, on average, a higher proportion of fluvial deposits and a lower proportion of central-basin estuarine deposits; estuarine deposits tend however to be thicker. This suggests a control on IVF stratigraphic architecture exerted by distinct characteristics of the tectonic setting of the host continental margins, notably basin physiography, rates and mode of sediment supply, and nature of sediment load. (ii) The thickness and proportion of lowstand systems tract are positively correlated with coastal-plain IVF dimensions, likely reflecting the role of drainage-basin area in dictating the scale of the fluvial systems that carved and infilled the valleys. (iii) Positive correlations are observed between the thickness of fluvial deposits, bayhead-delta deposits and central-basin estuarine deposits, versus coastal-plain IVF dimensions and valley catchment area. This suggests a control exerted by the river-system scale on sediment-supply rates and on the accommodation determined by valley size. (iv) Positive correlations between the thickness and proportion of barrier-complex deposits within cross-shelf IVFs versus mean shelf gradient indicate that the geometry of the shelf might control the establishment and preservation of barrier-island environments in incised valleys located on the shelf. (v) Correlations between the width of coastal-plain IVFs and present-day mean tidal range at the shoreline indicate that tidal dynamics may contribute to the widening of the incised valleys. Positive correlation is observed between the proportion of tide-dominated elements in highstand IVF deposits and IVF width, suggesting possible effects of interplays between hydrodynamic conditions and the geometry of incised valleys on their infills. This study highlights the complexity of the internal fills of incised valleys, which must be considered when attempting the application of facies models of IVFs to rock-record interpretations or as predictive tools in subsurface studies