Taxonomie čeledi Enicocephalidae (Heteroptera: Enicocephalomorpha) Orientální oblasti a nové objevy v morfologii nových taxonů

The infraorder Enicocephalomorpha represents the basalmost group of Heteroptera, the sister group of all remaining Heteroptera (= Euheteroptera) (ŠTYS 1989). The infraorder, whose unique phylogenetic position was recognized by most modern authors (ŠTYS & KERZHNER 1975), is currently divided into two families, Aenictopecheidae and Enicocephalidae. Some new taxa are described (Results: Parts A-H) and special attention is paid to newly discovered or overlooked morphological characters of the group, some of them important for all the Heteroptera. In Part A is described first member of worldwide distributed genus Systelloderes from the Oriental region, S. loebli Štys & Baňař, 2007 and the term neopatella is established for for sclerites within the femoro-tibial intersegmental membrane. Xenicocephalus josifovi Štys & Baňař, 2008 is described in Part B, representing first known complete adult of this peculiar genus. Unique type of raptorial foreleg among all Enicocephalomorpha is described and illustrated and its function by predation is assumed. In Part C is described new genus Phaenicocleus Štys & Baňař, 2009 from northern Borneo, based on males of three species. New diagnostic characters in Enicocephalomorpha are used. Phaenicocleus granulosus Baňař & Štys, 2011 is described in Part D, based on the...Infrařád Enicocephalomorpha je bazální skupinou ploštic, sesterskou skupinou všech zbývajících Heteroptera (= Euheteroptera) (ŠTYS 1989). Infrařád, jehož unikátní fylogenetické zařazení bylo rozpoznáno většinou moderních autorů (ŠTYS & KERZHNER 1975), je v současné době členěn na dvě čeledi - Aenictopecheidae a Enicocephalidae. V předložené dizertační práci jsou popsány některé nové taxony infrařádu Enicocephalomorpha (Výsledky: Části A-H), přičemž byla zvláštní pozornost věnována nově objeveným, nebo dosud přehlíženým morfologickým znakům této málo známé skupiny, případně znakům, důležitým pro celý řád Heteroptera. V Části A je popsán první zástupce široce rozšířeného rodu Systelloderes z Orientální oblasti, S. loebli Štys & Baňař, 2007 a byl stanoven termín neopatella pro sklerity mezi přední holení a stehnem. Xenicocephalus josifovi Štys & Baňař, 2008, který je popsán v Části B, reprezentuje první známý kompletní exemplář tohoto zvláštního rodu. Byl u něho popsán nový typ loupeživé přední nohy, unikátní v rámci celého infrařádu Enicocephalomorpha. V práci je diskutována potenciální potravní specializace tohoto druhu. Nový rod Phaenicocleus Štys & Baňař, 2009, stanovený pro tři druhy, všechny známé pouze podle jednoho samce, je popsán ze severního Bornea v Části C. V této práci jsou poprvé...Department of ZoologyKatedra zoologieFaculty of SciencePřírodovědecká fakult

Novi nalazi invazivne nearktičke vrste stjenice Leptoglossus occidentalis (Heteroptera: Coreidae) u Hrvatskoj

Additional records of the invasive Nearctic true bug Leptoglossus occidentalis Heidemann, 1910 (Heteroptera: Coreidae: Coreinae: Anisoscelini) from Croatia are given from the islands of Rab, Brač, and Hvar. A new host plant, Pinus halepensis, is reported.U radu se donose novi nalazi invazivne nearktičke vrste stjenice Leptoglossus occidentalis Heidemann, 1910 (Heteroptera: Coreidae: Coreinae: Anisoscelini) za Hrvatsku, i to s otoka Raba, Brača i Hvara te se govori o novoj biljci domaćinu, alepskom boru Pinus halepensis

The first unique-headed bug (Hemiptera, Enicocephalomorpha) from Cretaceous Iberian amber, and the Gondwanan connections of its palaeoentomological fauna

Enicocephalomorpha, also known as unique-headed bugs, are a seldom-collected infraorder of heteropteran insects whose evolutionary relationships have puzzled entomologists for more than a century. Unique-headed bugs are exceptionally rare in the fossil record, which hinders our understanding of the morphological transformations of the lineage across time and also affects the calibration of molecular clock estimates used to date the origins of the infraorder. Here, we report the discovery of Enicocephalinus ibericus sp. nov. from Iberian amber in the Ariño deposit in Spain, early Albian (Early Cretaceous) in age. The new species represents the second oldest fossil enicocephalomorphan to date, and the second record of this infraorder from European deposits. Remarkably, the closest relative of E. ibericus is the congeneric E. acragrimaldii Azar from Lebanese amber that is c. 20 myr older (Barremian), indicating a long-term persistence of the Enicocephalinus lineage across geological time. A review of the existing literature enabled us to record a total of 20 congeneric insect species that have been found in both Lebanese and Iberian ambers, suggesting the existence of previously underappreciated entomofaunal connections between southern Laurasia (the European archipelago) and northern Gondwana during the Cretaceous. We show that the palaeoentomological record holds remarkable potential for elucidating the faunistic exchanges and palaeobiogeographical patterns in the peri-Tethyan region during the Cretaceous

A new species of Systelloderes (Hemiptera: Heteroptera: Enicocephalidae) from South Africa

Baňař, Petr (2008): A new species of Systelloderes (Hemiptera: Heteroptera: Enicocephalidae) from South Africa. Acta Entomologica Musei Nationalis Pragae 48 (2): 233-240, DOI: 10.5281/zenodo.534086

Taxonomie čeledi Enicocephalidae (Heteroptera: Enicocephalomorpha) Orientální oblasti a nové objevy v morfologii nových taxonů

Infrařád Enicocephalomorpha je bazální skupinou ploštic, sesterskou skupinou všech zbývajících Heteroptera (= Euheteroptera) (ŠTYS 1989). Infrařád, jehož unikátní fylogenetické zařazení bylo rozpoznáno většinou moderních autorů (ŠTYS & KERZHNER 1975), je v současné době členěn na dvě čeledi - Aenictopecheidae a Enicocephalidae. V předložené dizertační práci jsou popsány některé nové taxony infrařádu Enicocephalomorpha (Výsledky: Části A-H), přičemž byla zvláštní pozornost věnována nově objeveným, nebo dosud přehlíženým morfologickým znakům této málo známé skupiny, případně znakům, důležitým pro celý řád Heteroptera. V Části A je popsán první zástupce široce rozšířeného rodu Systelloderes z Orientální oblasti, S. loebli Štys & Baňař, 2007 a byl stanoven termín neopatella pro sklerity mezi přední holení a stehnem. Xenicocephalus josifovi Štys & Baňař, 2008, který je popsán v Části B, reprezentuje první známý kompletní exemplář tohoto zvláštního rodu. Byl u něho popsán nový typ loupeživé přední nohy, unikátní v rámci celého infrařádu Enicocephalomorpha. V práci je diskutována potenciální potravní specializace tohoto druhu. Nový rod Phaenicocleus Štys & Baňař, 2009, stanovený pro tři druhy, všechny známé pouze podle jednoho samce, je popsán ze severního Bornea v Části C. V této práci jsou poprvé...The infraorder Enicocephalomorpha represents the basalmost group of Heteroptera, the sister group of all remaining Heteroptera (= Euheteroptera) (ŠTYS 1989). The infraorder, whose unique phylogenetic position was recognized by most modern authors (ŠTYS & KERZHNER 1975), is currently divided into two families, Aenictopecheidae and Enicocephalidae. Some new taxa are described (Results: Parts A-H) and special attention is paid to newly discovered or overlooked morphological characters of the group, some of them important for all the Heteroptera. In Part A is described first member of worldwide distributed genus Systelloderes from the Oriental region, S. loebli Štys & Baňař, 2007 and the term neopatella is established for for sclerites within the femoro-tibial intersegmental membrane. Xenicocephalus josifovi Štys & Baňař, 2008 is described in Part B, representing first known complete adult of this peculiar genus. Unique type of raptorial foreleg among all Enicocephalomorpha is described and illustrated and its function by predation is assumed. In Part C is described new genus Phaenicocleus Štys & Baňař, 2009 from northern Borneo, based on males of three species. New diagnostic characters in Enicocephalomorpha are used. Phaenicocleus granulosus Baňař & Štys, 2011 is described in Part D, based on the...Katedra zoologieDepartment of ZoologyFaculty of SciencePřírodovědecká fakult

Taxonomy of Oriental Enicocephalidae (Heteroptera: Enicocephalomorpha) and morphological novelties of new taxa.

The infraorder Enicocephalomorpha represents the basalmost group of Heteroptera, the sister group of all remaining Heteroptera (= Euheteroptera) (ŠTYS 1989). The infraorder, whose unique phylogenetic position was recognized by most modern authors (ŠTYS & KERZHNER 1975), is currently divided into two families, Aenictopecheidae and Enicocephalidae. Some new taxa are described (Results: Parts A-H) and special attention is paid to newly discovered or overlooked morphological characters of the group, some of them important for all the Heteroptera. In Part A is described first member of worldwide distributed genus Systelloderes from the Oriental region, S. loebli Štys & Baňař, 2007 and the term neopatella is established for for sclerites within the femoro-tibial intersegmental membrane. Xenicocephalus josifovi Štys & Baňař, 2008 is described in Part B, representing first known complete adult of this peculiar genus. Unique type of raptorial foreleg among all Enicocephalomorpha is described and illustrated and its function by predation is assumed. In Part C is described new genus Phaenicocleus Štys & Baňař, 2009 from northern Borneo, based on males of three species. New diagnostic characters in Enicocephalomorpha are used. Phaenicocleus granulosus Baňař & Štys, 2011 is described in Part D, based on the..

Taxonomy of Oriental Enicocephalidae (Heteroptera: Enicocephalomorpha) and morphological novelties of new taxa.

The infraorder Enicocephalomorpha represents the basalmost group of Heteroptera, the sister group of all remaining Heteroptera (= Euheteroptera) (ŠTYS 1989). The infraorder, whose unique phylogenetic position was recognized by most modern authors (ŠTYS & KERZHNER 1975), is currently divided into two families, Aenictopecheidae and Enicocephalidae. Some new taxa are described (Results: Parts A-H) and special attention is paid to newly discovered or overlooked morphological characters of the group, some of them important for all the Heteroptera. In Part A is described first member of worldwide distributed genus Systelloderes from the Oriental region, S. loebli Štys & Baňař, 2007 and the term neopatella is established for for sclerites within the femoro-tibial intersegmental membrane. Xenicocephalus josifovi Štys & Baňař, 2008 is described in Part B, representing first known complete adult of this peculiar genus. Unique type of raptorial foreleg among all Enicocephalomorpha is described and illustrated and its function by predation is assumed. In Part C is described new genus Phaenicocleus Štys & Baňař, 2009 from northern Borneo, based on males of three species. New diagnostic characters in Enicocephalomorpha are used. Phaenicocleus granulosus Baňař & Štys, 2011 is described in Part D, based on the..

A new Xenicocephalus species from Ecuador (Heteroptera, Enicocephalomorpha, Enicocephalidae)

Xenicocephalus tomhenryi sp. n. (Insecta: Hemiptera: Heteroptera: Enicocephalomorpha: Enicocephalidae) is established for a single macropterous female from Ecuador. The enigmatic genus now includes three species known from only two Neotropical adults and an incomplete female specimen. The new species is described and illustrated, extensive comparative diagnoses for Xenicocephalus species are provided, and nomenclature, distribution, and biology of the genus are reviewed. The architecture of the raptorial forelegs of Xenicocephalus is unique among Enicocephalomorpha, and the genus is classified as subfamily incertae sedis

Enicocephalus flavicollis Westwood 1837

Enicocephalus flavicollis Westwood, 1837 - male, redescription (Figs. 1 –6, 10– 17) Material examined. Lectotype: male, first label: // I. S ti. Vincentii // [handwritten = Insula Sancti Vincentii (Latin)]; second label: // Enicocephalus / flavicollis Westw. // [handwritten]; third label: // Type Hem: 4 / 6 / Enicocephalus / flavicollis / Westwood / Hope Dept. Oxford // [partly printed and partly handwritten]. Specimen bears red label: // Enicocephalus flavicollis / Westwood, 1837 / LECTOTYPE ♂ / P. Štys & P. B aňař det. 2011 // [printed]. Specimen dry mounted, glued to triangle card, in very bad condition: midlobe of pronotum (and some sternal parts) strongly damaged, antennal segments III, IV, left mid- and hindtibiae and tarsi are missing, left hindfemur mounted separately (Fig. 1). Deposited in Hope’s collection, University Museum of Natural History, Oxford. Paralectotype: male, first label: // Windward side / St. Vincent, W.I. / H.H. Smith 190 // [printed; number ‘ 190 ’ handwritten]; second label: // 95-206. //; third label: // H enicocephalus [originally Enicocephalus - overwritten] / flavicollis Westw. // [handwritten]; specimen bears red label: // Enicocephalus flavicollis / Westwood, 1837 / PARALECTOTYPE ♂ / P. Št ys & P. B aňař det. 2011 // [printed]. Dry mounted specimen glued to a triangle, in good condition, right antennal segment 4 and left hind leg missing. Deposited in Department of Entomology, Natural History Museum, London. Measurements (in mm): lectotype first (paralectotype in parentheses, italics). Total body length —ca 3.9 (specimen damaged) (3.87). Head (without neck). Total length— 0.62 (0.63); posterior lobe, length— 0.24 (0.23), width— 0.38 (0.37); distance of eye to apex of antennifer— 0.13 (0.13); diatone (maximum width across eyes)— 0.39 (0.38); dorsal synthlipsis (minimum interocular distance)— 0.18 (0.19); eye, length— 0.17 (0.16); gena, length— 0.03 (0.03), width (minimum)— 0.19 (0.18). Labium. Total length— 0.47 (0.46); segment I, length— 0.06 (0.05); segment II, length— 0.07 (0.07); segment III, length— 0.20 (0.20); segment IV, length— 0.13 (0.13). Antenna. Segment I, length— 0.14 (0.13); segment II, length— 0.29 (0.28), basal width— 0.02 (0.02), distal width— 0.05 (0.05); segment III, length— 0.37 (0.37); segment IV, length— 0.36 (0.36). Pronotum. Total length (median)—ca 0.58–0.62 (specimen damaged) (0.60); collum, length (median)— 0.13 (0.13), width (maximum)— 0.29 (0.29); midlobe, length (median)—ca 0.25–0.29 (specimen damaged) (0.27), width (maximum)— 0.54 (0.55); hindlobe, length (maximum)— 0.40 (0.38), length (mediane)— 0.19 (0.18), width (maximum)— 0.80 (0.78). Foreleg. Femur, length— 0.82 (0.81), width (maximum)— 0.20 (0.19); tibia, length— 0.74 (0.72), width (maximum)— 0.17 (0.17). Coloration (Figs. 1, 3, 5). Bicolorous, piceous or yellow, the colours strongly contrasting except on abdomen. No red or scarlet pigments. Head and thorax. Dorsum. Dark body parts piceous brown (dorsum of head, most of the posterior lobe of pronotum, mesoscutellum); neck of the head somewhat lighter; non-melanized parts unicolorous, contrastingly yellow (collum, midlobe of pronotum; linear, transverse, short, anterior strip of the hindlobe of pronotum). The yellow transverse strip of posterior lobe of pronotum with 1 + 1 submedial extensions shaped as segments of circle. Lateral sides and venter of head and thorax. Anterior lobe of head (from apex to postocular constriction) piceous, posterior lobe yellow; prothorax yellow (the yellow area confluent with the anterior yellow strip on posterior lobe of pronotum), pterothorax (incl. all supracoxal lobes) piceous, pterosterna yellowish. Antennae piceous brown; proximal part of segment IV non-contrastingly lighter. Labium: segments I and II brownish, III and IV yellow; legs (inclusive coxae) uniformly yellow. Forewings piceous brown with blackish hue (Figs. 1, 3), veins concolorous except for the noncontrastingly yellowish marginal vein (costal margin cum ambient vein up to the level of cu-an). The light coloration of the marginal vein invisible in transmitted light owing to semitransparency of the wing; ventral side matching the dorsal one. Hindwings blackish-brown. Abdomen. Dorsum not examined; venter: segments 1–3 yellowish brown, 3–7 piceous, 8 and 9 yellowish brown; posteromedial part of 2 and medial parts of 3–5 black. Coloration of paralectotype London male matching the redescription of Oxford lectotype male (Wyhodzinsky& Schmidt 1991) except for the lectotype abdomen said to be whitish. Microsculpture. Cuticle strongly lustrous and smooth. No setigerous tubercles. Posterior lobe of pronotum irregularly shallowly rugulose. Ventral surface of fore trochanter with about 5 transverse rows of serrate microstructures. Ventral face of forefemur with a continuous percurrent longitudinal strips of minute black granules; these terminating nearly apically, leaving the area contacting basis of foretibia bare but provided with one large posterior dens; no microtrichiae or spicules present. Venter of abdomen with minute transverse wrinkles. Vestiture. Macrotrichia (further on ‘setae’) golden, rather soft, usually short and straight, rarely curved. Head. Dorsal view. Anterior lobe of head covered by short and appressed, very dense short setae, obscuring its surface; longer curved setae occurring apically and in front of eyes only. Posterior lobe with sparse straight to curved, rather short setae. Lateral view. Dorsum with long, very dense, regularly curved, diagonal setae directed anteriad (excl. on preocular lobe), those on posterior lobe obscuring its outline. Venter with long, anteriorly directed, long straight or curved setae forming a conspicuous barb on the posterior lobe. Ventr al view. Very long and irregular macrotrichial cover all over. Eyes with a few straight interommatidial setae in anterior half, with many curved setae in posterior half. Antennae with short, dense, diagonal pubescence; segment 2 with few semi-erect hairs on its inner side, segment 3 on both sides and apically, segment 4 all over. Labium. Segment 3 with moderately long and moderately dense diagonal straight macrotrichia interspersed with long, thin, erect to suberect trichobothrium-like setae along both sides (4 + 4) and clustering in the apical part. Thorax. Dorsal view. Lateral outline: collum with moderately dense, straight short setae, midlobe with curved setae gradually longer caudad, hindlobe with very dense, curved, short setae. Mesoscutellum with long curved dense setae. Lateral view. Dorsum densely covered by extremely short, and very dense cover of straight to curved, diagonal, minute setae, only those on collum with a few erect setae. Prothoracic presupracoxale with a cluster of strikingly long hairs. Forewings with short, curved hairs (mixture of golden and black ones) distributed in a manner modal for Enicocephalus (cf. Wygodzinsky & Schmidt 1991). Intervenial membrane bare. Hindwings bare. Legs. Anterior faces of all coxae densely pilose. Foreleg. Dorsal face of femur with long diagonal setae, with interspersed few trichobothrium-like diagonal setae (recognizable only by their thinness); ventral face with long, thin, straight setae. Tibia covered all over by short, medium long, mostly appressed to diagonal straight setae (similar to those on dorsal face of femur), occurring on the anterior longitudinal tibial depression as well; dorsal and ventral faces additionally with numerous trichobothrium-like setae, all thin and all erect, rarely straight, mostly curved at the tip, nearly as long as local tibial diameter. The latter kind of setae occurring all over tarsus. Mid - and hindtibiae with a long diagonal vestiture all over, setae longer than tibial diameters; the same true for mid - and hindtarsi. The setae of dorsal and ventral tibial faces thin, trichobothrium-like (?). Abdomen. Margin without specialized setae, uniformly covered with uniformly curved setae becoming gradually longer posteriad, longest all along edge of segments 7 and particularly 8; the latter posteriorly with 3 + 3 very long setae, the longest exceeding apex of pygophore. Venter. All segments (exc. pygophore) densely covered with very short and uniform diagonal pubescence with setae slightly dilated (not gradually pointed) at tips, a few longer setae present at the medial keel only. Structure. Very small species, but rather robustly built. Head. Dorsal view (Figs. 1 –2, 6, 11). Anterior lobe short, insertion of antennae far from apex of anteclypeus, midway beween eyes and anteclypeus, gena very short, ratio gena lenght to its width 0.17; ratio of gena length to length of eye 0.19. Postocular constriction deep and long (about as long as the gena) and sharply delimited. Postocular lobe strikingly transverse, laterally asymmetrically rounded, widest behind the middle, ratio of its length to maximum width 0.62, no structural indication of the median except for a slight interocellar concavity. Ocelli situated on low tubercles, interocellar distance shorter than distance eye–ocellus. Head. Lateral view. Dorsal outline of the anteocular part of anterior lobe straight, slanting ventrad, strongly convex over the eye. Posterior lobe strongly convex, bulbous, its outline much exceeding that of the anterior lobe (more than shown by Wygodzinsky & Schmidt (1991, Fig. 80 B)). Ventral outline of preocular part of anterior lobe straight, that of posterior lobe moderately convex. Eyes large, strongly convex and outstanding, inner margins convex in dorsal view, ommatidia separately convex. Dorsal ocular index 2.00. Lateral view: dorsal margin far distant from dorsal margin of head, ventral margin slightly exceeding ventral margin of head. Ventral view: eyes strongly approaching each other. Antennae short, first segment unusually thick, 2 nd subclavate, strongly widening towards apex, segments III and IV stick-shaped, much thinner than segment II (but not flagelliform). Antennal formula (longest segment first) III-II-IV-I. Labium. Segments I and II very short, their dorsal faces flat as well as that of segment III; the latter rather short, conical. Labial formula (longest segment first) III-IV-II-I. Thorax. Pronotum in dorsal view . (Figs. 1 –2, 6, 11). Collum long and laterally rounded, lacking lateroventral tubercles and any structurally indicated median, but with an unusual feature (natural or artifact?): posterior 3 / 4 with one medial and 1 + 1 sublateral subangular shallow impressions, contacting each other and reaching the short and sharp constriction between collum and midlobe. Midlobe. Dorsal view. (Owing to the light colour and strong lustre of the area the description of surface sculpture is open to reinterpretation). Lateral margins strongly rounded, level of maximum width close to the anterior margin, posterior margin nearly straight, not interrupted. Ratio of midlobe maximum width to median length 3.05. Medial impression: antero-medial sector slightly depressed, with two impressed lines meeting in the centre and delimiting an anteromedial acutangular triangle with an apex pointing centrad; apex of the triangle sloping into a deep pit with a more shallow lateral linear extensions and rampart-like posterior margin coinciding with the posterior margin of the lobe itself (the impression resembling Oncylocotis -like inverse T-shaped impression.). The impression containing a linear, impressed median. Sublateral impressions present as simple, shallow, depressions, not interrupting the posterior margin of the midlobe and situated anterolaterad to the median depression. Posterior lobe. Dorsal view. Lateral margins regularly rounded inclusive indistinctly indicated posterolateral angles; medial part of posterior margin deeply angularly excised, lateral parts of this excision convex. No median structure. Ratio of hindlobe maximum width to its maximum length 2.05. Pronotum in lateral view. Collum with no ventrolateral tubercles. Lateral sides of pronotum not clearly delimited from supracoxal lobes, no horizontal lateral ridge or sulcus present. ‘Proepimeral lobe’ short, rounded, ventrally not reaching the level of apex of posterior prosupracoxale. Mesoscutellum with a convex surface, short, strongly rounded lateral margins, and unusually broad and long, apically rounded mucro separated from the disc by a transverse impression. Wings macropterous, not caducous. Forewings much exceeding the abdomen, venation (Fig. 13) as illustrated by Wygodzinsky & Schmidt (1991, Fig. 80 C), only the distal sector of AA + AP continuous, complete (checked on both the specimens). Hindwings longer than abdomen Legs. Foreleg. For microsculpture of trochanter and femur see above. Dorsal face of distal part of trochanter convex, slightly elevated over the neighbouring part of femur; ventral basis of femur slightly angularly produced and exceeding the surface of trochanter ventrad; ratio of femur length to maximum width 4.26. Tibia abruptly widening on dorsoventral plane in about a third of its length; ratio of tibia length to maximum width 4.24. Longitudinal (anterior) tibial depression starting at the points of tibial dilatation and running up to the cleaning comb. (Foretarsus pressed to the distal edge of tibia on both forelegs - consequently, apicitibial and tarsal armatures could not have been studied). Inner (anterior) claw large, posterior one represented by a short rounded hump only. Mid - and hindtibiae with two apical bristle combs each; claws nearly isomorphic (the inner (anterior one) slightly longer. Abdomen. Venter sclerotized, sulci between ventral laterotergites and sterna marked by deep impression and vaguely delimited laterosternal plates. Segments 3–7 with a sharp median keel, segment 8 without the keel, simply trapezoidal, posterolateral angles not protruding, embracing strikingly large, broadly transverse pygophore. Pygophore (Figs. 15–16) (as seen in strictly posterior view and examined under 100 magnification; dorsum covered by abdominal segment 8 and wings) strongly transverse, posterior face oval, delimited by a rampart-like swollen margin of the posterior foramen of the pygophore. Guide slender and small, of a modal shape, with distinct shaft, head and frame, distal margin of guide head truncate. Supradistal plate (Fig. 16) seen as a sclerite dorsad and anterad to guide, parameres situated laterad to the former. Diagnostic characters. Small species (3.9 mm), contrastingly bicolorous (lacking red pigment, yellow and brown - shared with E. boraceianus, E. guarani, E. schuhi (length 6.5–6.7 mm), and E. tupi only), legs uniformly yellow, collum and midlobe of pronotum uniformly yellow, the yellow part of pronotum involving also a short stripe on the disc of the hindlobe (unique!). Discal cell of forewing strikingly long and pointed, reaching nearly the wing margin (unique!). Male forelegs strikingly slender and genitalia small. Outer fore claw extremely reduced (shared with E. semirufus and E. usingeri only). Endemic to St. Vincent Island, sympatric occurrence of any other Enicocephalus species unknown.Published as part of Štys, Pavel & Baňař, Petr, 2012, Taxonomy and nomenclature of Enicocephalus flavicollis Westwood — type species of the type genus of the Enicocephalidae (Hemiptera: Heteroptera: Enicocephalomorpha), pp. 53-68 in Zootaxa 3237 on pages 56-59, DOI: 10.5281/zenodo.21417

A new Cervinotaptera species from northern Madagascar (Hemiptera, Heteroptera, Aradidae)

A new species, Cervinotaptera tomhenryi, sp. n. (Hemiptera: Heteroptera: Aradidae: Mezirinae), from Montagne d’Ambre National Park in northern Madagascar is described and illustrated. The newly described species is compared with the only other known species, Cervinotaptera guilberti Heiss & Marchal, 2012