17 research outputs found
Goblin spider genus Neoxyphinus.
75 p. : ill., maps ; 26 cm.The Neotropical spider genus Neoxyphinus Birabén, 1953, is revised, comprising 10 species found from the West Indies to northern Argentina. The genus is characterized by a unique combination of characters, which include the presence of a set of low tubercles or large spikes on the posterior surface of carapace; male endites with an apical, retrolateral excavation bearing a subapical toothlike apophysis; palpal bulb strongly inflated; embolus compact, S-shaped, with large, round ejaculatory opening; epigynal atrium large, with angular lateral corners. The large, round ejaculatory duct may be a synapomorphy of the genus. The genus Hawkeswoodoonops Makhan and Ezzatpanah, 2011, is synonymized with Neoxyphinus and the two species included in that genus are considered as nomina dubia. The monotypic genus Decuana Dumitrescu and Georgescu, 1987, is also synonymized with Neoxyphinus and its type species, D. hispida Dumitrescu and Georgescu, 1987, is transferred to the genus and redescribed. Confirming a hypothesis raised in the literature, Dysderina termitophila Bristowe, 1938, is transferred to Neoxyphinus and recognized as the senior synonym of the type species N. ogloblini Birabén, 1953. Two other species are transferred from Dysderina to Neoxyphinus: D. keyserlingi Simon, 1907, here recognized as the senior synonym of D. rugosa Bristowe, 1938, and D. furtiva Chickering, 1968, of which the male is described for the first time. In addition to the redescription of N. termitophilus, n. comb., N. hispidus, n. comb., N. xyphinoides (Chamberlin and Ivie, 1942), N. keyserlingi, n. comb., and N. furtivus, n. comb., five new species are described: N. petrogoblin Abrahim and Ott, from the Amazon basin in Brazil, Colombia, Ecuador, and Peru; N. gregoblin Abrahim and Santos, from Venezuela; N. axe Abrahim and Brescovit, from Bahia, Brazil; N. boibumba Abrahim and Rheims, from Pará, Brazil; and N. barreirosi Abrahim and Bonaldo, from Brazil, Colombia, Guyana, and Venezuela. With the exception of N. boibumba, known only from males, all new species are described from both sexes
A review of the Neotropical goblin spider genus Neoxyphinus Birabén 1953 (Araneae, Oonopidae)
The Neotropical genus Neoxyphinus Birabén 1953 is revised, comprising nine species found from Venezuela to northern Argentina. The monotypic genus Decuana Dumitrescu and Georgescu 1987 is synonymized with Neoxyphinus and
its type species, D. hyspida Dumitrescu and Georgescu 1987, is transferred to this genera. Confirming suspicions rose in literature, Dysderina termitophila Bristowe 1938 is transferred to Neoxyphinus and recognized as senior synonym of N. ogloblini Birabén 1953. Six new species are described, all of which known from both sexes: N. petrogoblin, from Amazonas, Brazil and Huanuco, Peru; N. gregoblin, from Andres Bello, Venezuela; N. axe, from Bahia, Brazil; N. barreirosi, from north Brazil; N. brega, from Brazil, Colombia, Guyana and Venezuela and N. caterete, from São Paulo, Brazil
A new species of Alpaida (Araneae, Araneidae) from Caxiuanã National Forest, Oriental Amazonia, Brazil Uma nova espécie de Alpaida (Araneae, Araneidae) da Floresta Nacional de Caxiuanã, Amazônia Oriental, Brasil
Alpaida guto n. sp. is described based on males and females from State of Pará, Brazil. This species appears to be the most abundant Alpaida O. P.-Cambridge, 1889 species in Caxiuanã National Forest, eastern Amazonia. The males seem to be close to A. antonio Levi, 1988 but can be easily distinguished by the terminal apophysis with rounded distal lobe and long, slender basal prong; females differ from those of A. yotoco Levi, 1988 by the median lobe with a basal constriction and by the bilobed median plate.<br>Alpaida guto sp. n. é descrita com base em machos e fêmeas do Estado do Pará, Brasil. Esta parece ser a espécie de Alpaida O. P.-Cambridge, 1889 mais abundante na Floresta Nacional de Caxiuanã, Amazônia Oriental. Os machos assemelham-se aos de A. antonio Levi, 1988 mas podem ser facilmente distinguidos pela apófise terminal com lobo distal arredondado e prolongamento basal longo e delgado; as fêmeas diferem daquelas de A. yotoco Levi, 1988 pelo lobo mediano com uma constrição basal e pela placa mediana bilobada
Neoxyphinus hispidus Abrahim & Brescovit & Rheims & Santos & Ott & Bonaldo 2012, new combination
Neoxyphinus hispidus (Dumitrescu and Georgescu), new combination Figures 198–209; map 2 Decuana hispida Dumitrescu and Georgescu, 1987: 92, pl. 2, 3 (two males, one female syntypes from Rancho Grande [probably Parque Nacional Henri Pittier, Aragua], Venezuela, XI.11.1982, T. Orguidan and V. Decu col., in “Grigore Antipa” National Museum of Natural History, Bucharest, not available). DIAGNOSIS: Males resemble those of N. boibumba in having abdominal anterior denticles and lacking carapace spikes, but differ by the longer pedicel, inserted obliquely in relation to the postepigastric scutum (fig. 203). Females differ from those of N. furtivus and N. keyserlingi by the smooth carapace, epigynal atrium strongly rebordered anteriorly and median element with long accessory prongs (figs. 216, 217). Females may be further distinguished by the presence of two prolateral spines on femur I (fig. 212), contrasting with those of N. furtivus, with one and with those of all remaining species, devoid of spines. MALE (PBI_ OON 40608, MNRJ): Total length 2.61. Carapace orange-brown, posterolateral surface without spikes, with a pair of raised blunt tubercles near the posterior margin (fig. 203), surface of elevated portion of pars cephalica smooth, sides finely reticulate, lateral margin with small, inconspicuous tubercles. Clypeus margin unmodified, straight in frontal view. Eyes: ALE separated by less than their radius. Sternum as long as wide, pale orange, median concavity absent, microsculpture absent, furrows shallow, setae evenly scattered, originating from small tubercles. Chelicerae, endites, and labium pale orange. Book lung covers large, ovoid. Pedicel tube medium sized. Dorsal scutum orange-brown, surface of middle and sides reticulate, anterior half with projecting denticles. Postepigastric scutum orange-brown. Legs yellow. Spine formula: tibia I v2-4-2, metatarsus I v2-2-2; tibia II v2-4-2, metatarsus II v2-2-2. Genitalia: sperm pore situated between anterior and posterior spiracles. Palp proximal segments yellow, cymbium and bulb white. Embolus with well-developed apical projection, without prolateral prong or prolateral prong, with a prolateral basal notch (figs. 209–211). FEMALE (PBI_ OON 40626): Total length 2.84. Dorsal scutum anterior half without projecting denticles. Carapace posterior surface with additional retrolateral protruded tubercle. Leg spination femora: I v0-0-2; tibiae: I, II v4-4-2; metatarsi: I, II v2-2-2. Epigynal median element large, curved, with conspicuous accessory prongs (figs. 216, 217). DISTRIBUTION: Known from the type locality and from Lara Department, Venezuela (map 2). MATERIAL EXAMINED: VENEZUELA: Lara: Parque Nacional Yacambu, Dec. 2002, A. Pérez and A. Giupponi, 1♂ (PBI _ OON 40608, MNRJ). Aragua: Rancho Grande–Maracy Road (First stream SE of Rancho Grande), May 01, 1992, L. Herman, 1♀ (PBI _ OON 40626, AMNH). Figs. 198–204. Neoxyphinus hispidus (Dumitresco and Georgesco), male. 198. Habitus, dorsal view. 199. Same, lateral view. 200. Carapace, anterior view. 201. Same, posterior view. 202. Abdomen, dorsal view. 203. Same, lateral view. 204. Same, ventral view. Figs. 205–209. Neoxyphinus hispidus (Dumitresco and Georgesco), male. 205. Abdomen, anterior view. 206. Mouthparts, posterior view. 207. Left palp, prolateral view. 208. Same, ventral view. 209. Same, retrolateral view. Figs. 210–217. Neoxyphinus hispidus (Dumitresco and Georgesco), female. 210. Carapace, dorsal view. 211. Same, posterior view. 212. Leg I, prolateral view. 213. Abdomen, dorsal view. 214. Same, ventral view. 215. Same, lateral view. 216. Internal female genitalia and tracheal system, dorsal view. 217. Epigynum, ventral view. Abbreviations: ME, median element; Re, receptaculum; Tc, tracheoles; TT, tracheal trunk.Published as part of Abrahim, Naiara, Brescovit, Antonio D., Rheims, Cristina A., Santos, Adalberto J., Ott, Ricardo & Bonaldo, Alexandre B., 2012, A Revision of the Neotropical Goblin Spider Genus Neoxyphinus Birabén, 1953 (Araneae, Oonopidae), pp. 1-76 in American Museum Novitates 2012 (3743) on pages 48-51, DOI: 10.1206/3743.2, http://zenodo.org/record/536034
Neoxyphinus furtivus Abrahim & Brescovit & Rheims & Santos & Ott & Bonaldo 2012, new combination
Neoxyphinus furtivus (Chickering), new combination Figures 5, 17, 23, 29, 47, 275–312; map 7 Dysderina furtiva Chickering, 1968: 13, figs. 26, 27 (female holotype from Jamaica, Saint Catherine, 3 miles north from Spanishtown, 16.Oct.1957, in MCZ, PBI_ OON 40607, examined). DIAGNOSIS: As in N. barreirosi and N. keyserlingi, the specimens of N. furtivus are devoid of both carapace spikes and abdominal denticles. Both males and females of this species can be distinguished by the reticulate surface of carapace and dorsal scutum (figs. 279, 280, 296, 297, 305, 308). Males differ from all other species by the presence of a small embolic prolateroapical lamella (figs. 291, 294). Females may be further distinguished by the presence of one prolateral spine on femur I, contrasting with two in N. hispidus and none in the remaining species. Map 7. Records of Neoxyphinus furtivus. MALE (PBI_ OON 14382, MPEG 10506): Total length 1.98. Carapace orange-brown, posterolateral surface without spikes, surface of elevated portion of pars cephalica and sides strongly reticulate, lateral margin without tubercles. Clypeus margin slightly rebordered, straight in frontal view. Eyes: ALE separated by less than their radius. Sternum as long as wide, orange-brown, median concavity absent, with a median field of small pits, setae evenly scattered, originating from surface. Book lung covers large, ovoid. Pedicel tube medium sized. Dorsal scutum orange-brown, surface of middle and sides strongly reticulate, anterior half without projecting denticles. Postepigastric scutum orange-brown. Legs pale orange. Spine formula: tibia I v2-4-0, metatarsus I v2-1-2. Genitalia: sperm pore situated between anterior and posterior spiracles. Palp proximal segments, cymbium, and bulb yellow. Embolus rounded, without both prolateral prong and prolateral lamella, but with an additional prolatero-apical lamella (figs. 287–295). FEMALE (PBI_ OON 14383, MPEG 13670): Total length 1.93. Carapace surface of elevated portion of pars cephalica and sides finely reticulate. Clypeus margin unmodified. Book lung covers ovoid. Dorsal scutum middle and sides surface finely reticulate. Spine formula: femur I p0-1-0, tibia I v5-3-2, metatarsus I v2-2-2; tibia II v4-4-0, metatarsus II v3-2-0; tibia III v1-1-1, metatarsus III v1-1-1. Epigynal atrium wide, with median element conical, placed anteriorly in our preparations (fig. 312). DISTRIBUTION: Known from Jamaica to northern Brazil (map 7). Figs. 275–286. Neoxyphinus furtivus (Chickering), male. 275, 276. Habitus, dorsal view. 277. Same, lateral view. 278. Carapace, anterior view. 279. Same, lateral view. 280. Same, posterior view. 281. Same, dorsal view. 282. Sternum and mouthparts, ventral view. 283. Abdomen, anterior view. 284. Same, dorsal view. 285. Same, lateral view. 286. Same, ventral view. Figs. 287–295. Neoxyphinus furtivus (Chickering), male. 287. Left palp, prolateral view. 288. Same, ventral view. 289. Same, retrolateral view. 290, 293. Left palp, embolus, prolateral view. 291, 294. Same, ventral view. 292, 295. Same, retrolateral view. Figs. 296–301. Neoxyphinus furtivus (Chickering). 296, 297. Male habitus and carapace, dorsal view. 298. Female abdomen, frontal view. 299. Male abdomen, lateral view. 300. Male endite, posterior view. 301. Male sternum and mouthparts, ventral view. Figs. 302–312. Neoxyphinus furtivus (Chickering), female. 302, 303. Habitus, dorsal view. 304. Same, lateral view. 305. Carapace, dorsal view. 306. Sternum and mouthparts, ventral view. 307. Carapace, anterior view. 308. Same, posterior view. 309. Abdomen, dorsal view. 310. Same, ventral view. 311. Same, lateral view. 312. Epigynum, dorsal view. OTHER MATERIAL EXAMINED: JAMAICA: Clarendon: 2.5 miles N of May Pen Rd. to Chapelton, Nov. 10, 1963, A. Chickering (PBI _ OON 27139, MCZ 66654), 1♀. Saint Ann: 3 mi S Moneague, Nov. 07, 1957, A. Chickering (PBI _ OON 27145, MCZ 71719), 2♀. Saint Andrew: Cooper’s Hill, Feb. 10, 1955, P. Bellinger (PBI _ OON 27142, MCZ 71725), 1♀; Ferry, E Red Hills Road, Sept. 27, 1957, A. Chickering (PBI _ OON 27147, MCZ 71720), 1♀; Hope Gardens, 18°01′59″N 76°45′W, Nov. 27–Dec. 06, 1963, A. Chickering (PBI _ OON 27141, MCZ 71718), 2♀; Jack’s Hill Road, 18°03′00″N 76°44′59″W, Dec. 06, 1957, A. Chickering (PBI _ OON 27146, MCZ 71723), 1♀; Stony Hill, 18°05′01″N 76° 48′00″W, May 28, 1956, C. Hoff (PBI _ OON 27163, MCZ 71726), 1♀, Oct. 18, 1957, A. Chickering (PBI _ OON 27138, MCZ 71724), 1♀. Saint Catherine: 3 mi N Spanishtown, Oct. 16, 1957 (PBI _ OON 27144, MCZ 71721), 2♀; Ewarton, 18°10′44″N 77°05′17″W, Nov. 29, 1957, A. Chickering (PBI _ OON 27148, MCZ 71722), 1♀. TRINIDAD AND TOBAGO: Arima: Arima Valley: No specific locality, 800–1200 ft, Feb. 10–22, 1964, Wygodzinsky and Rosen (PBI _ OON 38046, AMNH), 2♀; Simla, Apr. 05–25, 1964, A. Chickering (PBI _ OON 27143, MCZ 71728), 13♀, Apr. 20, 1964, same (PBI _ OON 38047, AMNH), 1♀. Saint George: St. Augustine: University campus, 10°38′20″N 61°23′45″W, Apr. 05–07, 1964, A. Chickering (PBI _ OON 27140, MCZ 71727), 24♀. BRAZIL: Roraima: Amajari: Reserva Biológica Ilha de Maracá, 03°22′01″N 61°25′00″W, Jan. 31–Feb. 14, 1992, A.B. Bonaldo (PBI _ OON 40125, MCTP 1836), 1♀. Amazonas: Manaus: 03°06′07″S 60°01′30″W, Jan. 28, 1992, A.B. Bonaldo (PBI _ OON 40124, MCTP 1479), 2♀. Pará: Belém: Bosque Rodrigues Alves–Jardim Botânico da Amazônia, 01°25′49.0″S 48°27′22.3″W, all collected by J.A.P. Barreiros, Apr. 01, 2001 (PBI _ OON 14412, MPEG 10501), 1♀, Apr. 08, 2001 (PBI _ OON 14402, MPEG 10503), 1♀, May 12, 2001 (PBI _ OON 14417, MPEG 10490), 1♀, Apr. 16, 2001 (PBI _ OON 14418, MPEG 10500), 1♀, Apr. 22, 2001 (PBI _ OON 14387, MPEG 10498; PBI _ OON 14401, MPEG 10497; PBI _ OON 14416, MPEG 10499), 3♀, Sept. 27, 2001 (PBI _ OON 14419, MPEG 10495), 1♀, Oct. 18, 2001 (PBI _ OON 14404, MPEG 10491; PBI _ OON 14428, MPEG 10496), 2♀, Oct. 25, 2001 (PBI _ OON 14391, MPEG 10483; PBI _ OON 14392, MPEG 10480; PBI _ OON 14400, MPEG 10504; PBI _ OON 14434, MPEG 10482), 7♀, Nov. 08, 2001 (PBI _ OON 14382, MPEG 10506; PBI _ OON 14383, MPEG 13670; PBI _ OON 14393, MPEG 10484; PBI _ OON 14410, MPEG 10488; PBI _ OON 14429, MPEG 10505; PBI _ OON 14435, MPEG 10481; PBI _ OON 14547, MPEG 10486), 1♂, 10♀, 2001 (PBI _ OON 14385, MPEG 10507; PBI _ OON 14389, MPEG 10492; PBI _ OON 14397, MPEG 10493; PBI _ OON 14408, MPEG 10487; PBI _ OON 14413, MPEG 10502; PBI _ OON 14414, MPEG 10485; PBI _ OON 14415, MPEG 10494; PBI _ OON 14427, MPEG 10489), 8♀; Reserva Mocambo, 01°26′28.7″S 48°24′46.2″W, Dec. 06–12, 2007, B.V.B. Rodrigues and J.M.B. Pereira-Filho (PBI _ OON 14633, MPEG 14013; PBI _ OON 14635, MPEG 14015; PBI _ OON 14636, MPEG 14016; PBI _ OON 14637, MPEG 14017; PBI _ OON 14638, MPEG 14018; PBI _ OON 14640, MPEG 14020; PBI _ OON 14641, MPEG 14021; PBI _ OON 14642, MPEG 14022; PBI _ OON 14643, MPEG 14023; PBI _ OON 14644, MPEG 14024), 10♀, Dec. 13–15, 2007, same (PBI _ OON 14634, MPEG 14014), 1♀, May 07, 2008, B.V.B. Rodrigues, J.M.B. Pereira-Filho, N. Abrahim and N.C. Bastos (PBI _ OON 14639, MPEG 14019), 1♀. Goianésia: Fazenda Rio Capim, 03°18′50″S 48°28′54″W, July 30, 2002, Equipe IPAN (PBI _ OON 14388, MPEG 10742; PBI _ OON 14398, MPEG 10741; PBI _ OON 14421, MPEG 10732), 2♂, 1♀, Aug. 30, 2002, same (PBI _ OON 14409, MPEG 10739), 1♀, June 15–29, 2003, same (PBI _ OON 14394, MPEG 10735; PBI _ OON 14439, MPEG 10727), 2♂, June 17, 2003, same (PBI _ OON 14407, MPEG 10730; PBI _ OON 14426, MPEG 10743; PBI _ OON 14440, MPEG 10715; PBI _ OON 14384, MPEG 10744; PBI _ OON 14390, MPEG 10740; PBI _ OON 14438, MPEG 10716), 2♀, 5♂. Melgaço: Estação Científica Ferreira Penna, FLONA Caxiuana, 01°44′15.5″S 51°26′42.0″W, Aug. 17, 2003, J.A.P. Barreiros (PBI _ OON 14755, MPEG 10372), 1♀, Oct. 25, 2003, same (PBI _ OON 14405, MPEG 10345), 1♀, Oct. 28, 2003, same (PBI _ OON 14754, MPEG 10374), 1♂. Novo Repartimento: Fazenda Arataú, 04°19′51″S 49°47′47″W, June 19–Aug. 12, 2003, Equipe IPAN (PBI _ OON 14423, MPEG 10724) 1♀, June 25–26, 2003, same (PBI _ OON 14395, MPEG 10719; PBI _ OON 14399, MPEG 10734; PBI _ OON 14403, MPEG 10710; PBI _ OON 14406, MPEG 10726; PBI _ OON 14411, MPEG 10713; PBI _ OON 14424, MPEG 10728; PBI _ OON 14430, MPEG 10720; PBI _ OON 14431, MPEG 10721; PBI _ OON 14436, MPEG 10722; PBI _ OON 14546, MPEG 10729), 2♂, 9♀, Aug. 11, 2006, same (PBI _ OON 14425, MPEG 10745), 1♀. Tailândia: Fazenda Santa Marta, May 13–14, 2003, Equipe IPAN (PBI _ OON 14396, MPEG 10733; PBI _ OON 14422, MPEG 10738; PBI _ OON 14432, MPEG 10731), 3♀; July 10–30, 2003, same (PBI _ OON 14433, MPEG 10714), 1♀, July 19–30, 2003, same (PBI _ OON 14420, MPEG 10711; PBI _ OON 14386, MPEG 10723), 1♂, 1♀. No precise location, Mata, Varzea, Oct. 2005, L. Macambira (PBI _ OON 14548, MPEG 10708), 3♀.Published as part of Abrahim, Naiara, Brescovit, Antonio D., Rheims, Cristina A., Santos, Adalberto J., Ott, Ricardo & Bonaldo, Alexandre B., 2012, A Revision of the Neotropical Goblin Spider Genus Neoxyphinus Birabén, 1953 (Araneae, Oonopidae), pp. 1-76 in American Museum Novitates 2012 (3743) on pages 61-67, DOI: 10.1206/3743.2, http://zenodo.org/record/536034
Tarsal organ morphology and the phylogeny of goblin spiders (Araneae, Oonopidae), with notes on basal genera
Based on a survey of a wide variety of oonopid genera and outgroups, we hypothesize new
synapomorphies uniting the Oonopidae (minus the South African genus Calculus Purcell, which is transferred to the Orsolobidae). The groundplan of the tarsal organ in Oonopidae is hypothesized to be an exposed organ with a distinctive, longitudinal ridge originating from the proximal end of the organ, and a serially dimorphic pattern of 4-4-3-3 raised receptors on legs I–IV, respectively. Such organs typify the diverse, basal, and ancient genus Orchestina Simon. Several other genera whose members resemble Orchestina in retaining two plesiomorphic features (an H-shaped,transverse eye arrangement and a heavily sclerotized, thick-walled sperm duct within the male
palp) are united by having tarsal organs that are partly (in the case of Cortestina Knoflach) or fully capsulate (in the case of Sulsula Simon, Xiombarg Brignoli, and Unicorn Platnick and Brescovit).
The remaining oonopids are united by the loss of the heavily sclerotized palpal sperm duct, presumably reflecting a significant transformation in palpal mechanics. Within that large assemblage,a 4-4-3-3 tarsal organ receptor pattern and an H-shaped eye arrangement seem to be retained only in the New Zealand genus Kapitia Forster; the remaining genera are apparently united by a reduction in the tarsal organ pattern to 3-3-2-2 raised receptors on legs I–IV and by the acquisition of a clumped eye arrangement. Three subfamilies of oonopids are recognized: Orchestininae
Chamberlin and Ivie (containing only Orchestina; Ferchestina Saaristo and Marusik is placed as a junior synonym of Orchestina), Sulsulinae, new subfamily (containing Sulsula, Xiombarg, Unicorn,and Cortestina), and Oonopinae Simon (containing all the remaining genera, including those previously placed in the Gamasomorphinae). The type species of Sulsula and Kapitia, S. pauper (O. P.-Cambridge) and K. obscura Forster, are redescribed, and the female of S. pauper is described for the first time. A new sulsuline genus, Dalmasula, is established for Sulsula parvimana Simon and four new species from Namibia and South Africa
A revision of the neotropical goblin spider genus Neoxyphinus Birabén, 1953 (Araneae, Oonopidae) /
no. 374
FIGURES 233–247 in Tarsal Organ Morphology and the Phylogeny of Goblin Spiders (Araneae, Oonopidae), With Notes on Basal Genera
FIGURES 233–247. Dalmasula lorelei, new species, female (233–241), D. parvimana (Simon), male (242–247). 233. Posterior median spinneret, apical view. 234. Posterior lateral spinneret, apical view. 235. Claws of leg I, anterior view. 236. Trichobothrial base from tibia III, dorsal view. 237. Tarsal organ from leg I, dorsal view. 238. Same, leg II. 239. Same, leg III. 240. Same, leg IV. 241. Same, palp. 242. Habitus, dorsal view. 243, 246. Left palp, prolateral view. 244, 247. Same, retrolateral view. 245. Habitus, ventral view.Published as part of Platnick, Norman I., Abrahim, Naiara, Álvarez-Padilla, Fernando, Andriamalala, Daniela, Baehr, Barbara C., Baert, Léon, Bonaldo, Alexandre B., Brescovit, Antonio D., Chousou-Polydouri, Natalia, Dupérré, Nadine, Eichenberger, Beata, Fannes, Wouter, Gaublomme, Eva, Gillespie, Rosemary G., Grismado, Cristian J., Griswold, Charles E., Harvey, Mark S., Henrard, Arnaud, Hormiga, Gustavo, Izquierdo, Matías A., Jocqué, Rudy, Kranz-Baltensperger, Yvonne, Kropf, Christian, Ott, Ricardo, Ramírez, Martín J., Raven, Robert J., Rheims, Cristina A., Ruiz, Gustavo R.S., Santos, Adalberto J., Saucedo, Alma, Sierwald, Petra, Szüts, Tamás, Ubick, Darrell & Wang, Xin-Ping, 2012, Tarsal Organ Morphology and the Phylogeny of Goblin Spiders (Araneae, Oonopidae), With Notes on Basal Genera, pp. 1-52 in American Museum Novitates 2012 (3736) on page 24, DOI: 10.1206/3736.2, http://zenodo.org/record/536018