Revision of Newsteadia (Homoptera: Coccoidea) of the Nearctic and Neotropic Regions, with Descriptions of New Species

Six new species of Newsteadia are described from the Nearctic and Neotropic Regions. A key is presented for the identification of 10 species currently included in the genus from these regions. The number of species in the genus worldwide is 37. A modified concept of the genus is presented and some new characters are given as diagnostic of the genus. The zoogeography of the genus is discussed

Scale insect (Hemiptera, Coccomorpha) survey of soil samples from southern Asia with description of two new species of Rhizoecidae

Seventeen species of Ortheziidae, Rhizoecidae and Pseudococcidae (Hemiptera: Coccomorpha) were extracted from East and South Asian soil sample collection of the Hungarian Natural History Museum. Among 1080 soil samples almost 100 included scale insect specimens. Two new species of the family Rhizoecidae, Rhizoecus muranyii Kaydan sp. n. and Ripersiella danyii Kaydan et Konczné Benedicty sp. n. are described and illustrated, and identification keys of East and South Asian Rhizoecus and Ripersiella species are provided

Coccus pseudomagnoliarum (Kuwana) (Hemipterea: Coccidae) teknőspajzstetű megjelenése városi környezetbe telepített nyugati ostorfán (Celtis occidentalis L)

A cikk szerzôi a Coccus pseudomagnoliarum (Kuwana, 1914) (Hemiptera: Coccidae) teknôspajzstetû elsô magyarországi megjelenésérôl számolnak be. A fajt eddig csak egy tápnövényen, a nyugati ostorfán (Celtis occidentalis) azonosították – jellemzôen csak városi környezetben, országos viszonylatban két helyszínrôl – Budapestrôl és Kecskemétrôl. Feltételezhetô, hogy a faj régebben jelen van városi környezetben. A faj javasolt magyar neve: szürke ostorfa-teknôspajzstetû

Pseudomontanococcus baloghi Kozar and Konczne Benedicty, sp. n.

Pseudomontanococcus baloghi Kozár and Konczné Benedicty sp. n. (Fig 5) Type material. Holotype, adult female: Papua New Guinea, Mt. Wilhelm, near to Brass Tarn, from moss and litter among Va cc in i um and Coprosma plants, 5.viii. 1969, by Berlese funnel by Dr. J. Balogh, NG-Mt-B. 42 (PPI). Paratype: same data as holotype, 1 slide with 2 nd -instar female nymph (PPI). Adult female (Fig. 5) Described from 1 specimen. Mounted specimen. Body elongate oval, 1.373 mm long, 0.855 wide. Antenna 7 segmented, segment lengths (Μm) I 58, II 55, III 30, IV 43, V 20, VI 20, VII 40; each antennal segment with few setae; segment II with a sensory pore; segments V & VI both with a falcate sensory seta 32–44 Μm long; apical segment with three sensory falcate setae, longest 78 Μm long; apical seta 75 Μm long. Frontal tubercle or lobe absent. Eye visible on margin. Venter. Labium 79 Μm long; with a total of 6 pairs of setae, all spinose; basal segment weakly developed, with 1 pair of setae. Legs long and well-developed (lengths in Μm), anterior legs: coxae 84, trochanter 65, femur 137, tibia 108, tarsus 103, claw 32; middle legs: coxae 84, trochanter 62, femur 132, tibia 108, tarsus 110, claw 30; hind legs: coxae 89, trochanter 62, femur 139, tibia 109, tarsus 101, claw 32, tarsal digitules knobbed, each 51–55; claw digitules each 34–35 and slightly knobbed. All coxae with spinulae on anterior surface; metacoxae with many small, translucent pores on posterior surface. Trochanter each with two pores on each side and with two long flagellate setae. Other leg segments with spinose and flagellate setae; tarsi each with a sensory pore. Claws without a denticle. Anterior and posterior spiracles each with a large group of five-locular pores in atrium; width of anterior peritreme 26 µm. Multilocular pores each 5 µm in diameter, with 5–9 loculi (usually 5), scattered medially on posterior abdominal segments and head, and also in a wide submedian band on all segments. Abdomen with a few flagellate setae medially, plus a marginal band of large spines and several spinose setae in a submarginal band. Cruciform pores absent. Microtubular ducts few, in a fairly narrow submarginal band. Macrotubular ducts few, present in a submarginal band and medially on thorax and head; each duct with inner ductule shorter than outer ductule; inner ductule with a flower-like terminal gland. Spinulae present only on abdomen. Vulva unclear. Each anal lobe with a large spinose suranal seta 53– 77 Μm long, a short spinose seta near each lateral margin 38 Μm long, and a long flagellate submarginal seta, 102 µm long. Segment VII with weak submedian sclerotization. Dorsum. Dorsal setae conical and spinose, of two sizes: largest setae along margin, robust, with 2 on each abdominal segment, each 41–52 µm long, intermixed with smaller setae, each about 21 µm long; also with long and short setae in a longitudinal median band, with 4 to about 14 setae on each segment, each seta 19–43 µm long; some small spinose setae also present submedially on head and thorax. Shorter, less strongly spinose setae, each 12–16 µm long, sparse on most segments. Macrotubular ducts each 18–25 Μm long, 2–3 Μm wide, abundant throughout. Microtubular ducts each 14 µm long and less than 1 Μm wide, with a bilocular inner end, scattered among dorsal setae, and sometimes with 1 or 2 present at base of a few large spinose setae. Anal ring unclear, 48 Μm wide, 52 Μm long, with a sparse row of pores along outer margin; with 8 flagellate setae, each about 125 µm long. Anal lobes large, sclerotized, 156 µm long and 127 µm wide, with a spinose seta on both inner and outer margins, inner seta 36 Μm long, outer seta 41 µm long, and a longer apical seta, 50 µm long. Anal lobes with dorsal surface covered with sclerotized protuberances, each protuberance with a microtubular duct; inner margin of lobes with sclerotized teeth. Spinulae present only on last abdominal tergites. Cauda present medially at base of anal lobes, 17–25 µm wide, and about twice as long as wide. With two areas of dense sclerotisation submedially on segment VII, each with strong protuberances and an associated microduct. Comment. For comparison with P. m a r t i n i, see under that species. Etymology. The new species is named after the late Professor Dr. János Balogh from Hungary, in acknowledgement of his collecting efforts that have yielded several important new Eriococcidae, including the present species. Second-instar female (Fig. 6) Described from 1 specimen. Mounted female. Body elongate oval, 0.958 mm long and 0.67 wide. Antenna 6 segmented, segment lengths (Μm) I 35, II 30, III 35, IV 15, V 20, VI 28; each antennal segment with few setae; segment II with a sensory pore; segments IV and V each probably with a falcate sensory seta but that on segment IV not clearly seen; falcate seta on segment V 45 Μm long; apical segment with three sensory falcate setae; apical seta 60 Μm long. Frontal tubercle or lobe absent. Eye visible on margin. Venter. Labium 53 Μm long; with total of 6 pairs of setae, all spinose; basal segment weakly developed, with one pair of setae. Stylet loop reaching metacoxae. Legs long and strong (lengths in Μm): anterior legs: coxae 46, trochanter 43, femur 88, tibia 66, tarsus 65, claw 20; middle legs: coxae 52, trochanter 40, femur 88, tibia 67, tarsus 70, claw 21; hind legs: coxae 54, trochanter 40, femur 83, tibia 70, tarsus 67, claw 22; tarsal digitules knobbed, each 36–39; claw digitules each 26–30 and slightly knobbed. Middle and hind coxae with spinulae on anterior surfaces; translucent pores absent on all coxae. Trochanter with two pores on each side and two long flagellate setae. Claws without a denticle. Legs with few strong spine-like and hair-like setae; tarsi each with a sensory pore. Anterior and posterior spiracles each with a large group of five-locular pores in atrium; width of anterior peritreme 15 µm. Multilocular pores each with 5–9 loculi (mostly 5 -locular), 4 µm in diameter, scattered medially on posterior abdominal segments and head, and also forming a longitudinal submedian band on all segments. Abdominal margin with a row of robust spines; lanceolate setae present on midventer, plus several small spinose setae in a submarginal band. Cruciform pores absent. Microtubular ducts few, in a submarginal band associated with spinose setae. Macrotubular ducts absent. Each anal lobe with a large spinose suranal seta 53 Μm long, a shorter ventral spinose seta medially 24 Μm long, and a long seta in a submarginal position (broken). Segment VII with an area of weak sclerotization submedially. Dermal spinules present on abdominal segments. Dorsum. Dorsal setae conical and spinose, of two sizes: largest setae robust, in a line along margin, with 1 seta per segment on abdomen but more on anterior segments, each 31–37 µm long; also with 2–8 medially on each segment in a longitudinal band, each 14–17 µm long. Shorter setae each 7–9 µm long, present in a sparse row on most segments. Macrotubular ducts absent. Microtubular ducts each 11 µm long, and less than 1 Μm wide, with a bilocular inner end; scattered among dorsal setae, and sometimes with 1 or 2 at base of some larger spinose seta. Anal ring unclear, 28 Μm wide, 32 Μm long, with a sparse row of pores along outer margin; with six flagellate setae, each 98 µm long. Anal lobes large, sclerotized, 77 µm long, 77 µm wide, each with a spinose seta on both inner and outer margins, inner seta 22 µm long, outer seta 19 Μm long, plus a spine in apical position (broken). Anal lobes with dorsal surface covered with sclerotised protuberances, each with a microtubular duct; inner margin of lobes with serrate sclerotized teeth. Cauda present medially at base of anal lobes, 12–22 µm wide, about twice a long as wide. Area of strong sclerotisation present submedially on segment VII. Dermal spinules absent on dorsum. Comment. The 2 nd -instar female nymphs of P. baloghi differ from those of P. m a r t i n i in having (characters states on P. martini in brackets): (i) both setae on trochanter flagellate (one spinose and one flagellate), (ii) metacoxae without translucent pores (present), and (iii) protuberances on dorsal surface of anal lobe and VIIth abdominal segment with associated microtubular ducts (absent).Published as part of Kozár, Ferenc, Benedicty, Zsuzsanna Konczné & Hodgson, Chris, 2008, A new felt scale genus (Hemiptera: Sternorrhyncha; Eriococcidae) from Papua New Guinea, with descriptions of two new species, pp. 47-62 in Zootaxa 1934 on pages 57-61, DOI: 10.5281/zenodo.18492

Pseudomontanococcus martini Kozar and Konczne Benedicty, sp. nov.

<i>Pseudomontanococcus martini</i> Kozár and Konczné Benedicty sp. nov. <p>(Fig 1)</p> <p> <b>Type material</b>. <b>Holotype, Papua New Guinea</b>: adult female: Mt. Wilhelm (ca 4270 m), from moss and litter under <i>Astelia papuana</i> (Liliaceae), 13.ix.1968, NG-M-B 118, collected by Berlese funnel by Dr. J. Balogh, deposited in PPI.</p> <p> <b>Paratypes: Papua New Guinea</b>: 1 adult female with same data as for holotype (PPI); one adult female plus 1 adult male on same slide, data as for holotype but from litter under <i>Papuapteris papuana</i> (Dryopteridae), by Berlese funnel, NG-M-B 117 (BMNH); one adult female + two 2nd -instar male nymphs on two slides, as for holotype but collected on 19.ix.1968, from moss by Berlese funnel, NG-M-B 125 (PPI); 1 2nd - instar male nymph and 1 2nd -instar female nymph on two slides, data as for holotype, by Berlese funnel from moss and litter under <i>Coprosma divergens</i> (Rubiaceae) bushes, NG-M-B 119, 120 (PPI); 1 2nd -instar female nymph, as for holotype but collected under <i>Styphelia suaveolens</i> (Epacridaceae) bushes from moss and litter by Berlese funnel, 13.ix.1968, NG-M-B 116 (PPI); 5 2nd -instar female nymphs + 2nd -instar male nymphs on three slides, as for holotype but collected from moss at 4400-4500 m, 6.viii.1969 (PPI).</p> <p> <b>Adult female</b> (Fig. 1)</p> <p>Described from four specimens. Data for non-holotype specimens given in brackets.</p> <p> <b>Mounted specimens</b>. Body elongate oval, 1.347 (1.295–1.502) mm long, 0.673 (0. 673–0.906 wide). Antennae 6 segmented, segment lengths (Μm) I 43 (42–50), II 36 (35–38), III 38 (36–44), IV 14 (6–14), V 18 (11–20), VI 28 (22–28); antennal segments with few setae; segment II with a sensory pore; segment III not tapered, partially divided due to pseudosegmentation in middle; two preapical segments each 36–40 (30–43) Μm long with a falcate sensory seta; apical segment with apical seta 62 (63–72) Μm long, plus 3 sensory falcate setae, each 58 (54–62) Μm long. Frontal tubercle or lobe absent. Eye present on margin.</p> <p> <b>Venter</b>. Labium 84 (76–84) Μm long; with a total of 6 pairs of setae, all spinose; basal segment weakly developed with 1 pair of setae. Legs long and well-developed (lengths in Μm), anterior legs: coxa 58 (58–65), trochanter 48 (41–48), femur 101 (101–120), tibia 72 (72–84), tarsus 74 (72–78), claw 30 (28–30); middle legs: coxa 58 (58–73), trochanter 48 (44–52), femur 106 (99–117), tibia 70 (70–79), tarsus 74 (72–81), claw 30 (28–31); posterior legs: coxa 72 (70–75), trochanter 48 (46–49), femur 101 (101–120), tibia 79 (79–86), tarsus 79 (68–84), claw 32 (31–32); tarsal digitules knobbed, each 38 (38–45), claw digitules each 28 (26–30) and slightly knobbed. All coxae with spinulae on anterior surface; posterior coxae with numerous small, translucent pores along outer margin on posterior surface. Claws without a denticle. Legs each with a few spinose setae, and with a sensory pore on each tarsus. Spiracles each with a large group of pores in atrium; diameter of anterior peritreme 17 (17–22) µm. Multilocular pores each 5 (4–5) µm in diameter, with 5–9 loculi, mostly 7, forming a wide submedian band on all segments and extending onto anal lobes. Abdomen with a few large lanceolate setae, plus several small spinose setae. Cruciform pores absent. Microtubular ducts present, sparse in a submarginal band. Macrotubular ducts present on all segments; each duct with inner ductule shorter than half-total length; inner ductule with a flower-like terminal gland. Vulva unclear. Each anal lobe with a large spinose suranal seta, 53 (53–72) Μm long, a short spinose ventral seta 26 (26–33) Μm long, plus a long flagellate seta in submarginal position, latter 118 (115–122) Μm long. An area of weak sclerotisation present submedially on segment VII.</p> <p> <b>Dorsum</b>. Dorsal setae conical and spinose; of two sizes: largest setae on margin, with 1 on each side of each abdominal segment, each 27–34 Μm long; plus 4–12 spines medially on each segment forming a longitudinal median band, each 20–25 Μm long. Shorter setae, each 9–17 µm long, present in a sparse row on all segments. Macrotubular ducts numerous, each 30–32 Μm long and 3 Μm wide. Microtubular ducts each 8–9 µm long, and less than 1 Μm wide, with a bilocular inner end; scattered among dorsal setae, and with 1–3 present at base of all large spines. Anal ring unclear but 42–45 Μm wide, 40–46 Μm long, with a sparse row of pores along outer margin; with six long setae, each 116–130 Μm long. Anal lobes sclerotised, 161 (144–168) µm long and 137 (135–137) µm wide, with a spinose seta on both inner and outer margins, inner seta 19 (19–22) µm long, outer seta 22 (22–29) Μm long, plus an apical seta 38 (broken?) µm long; entire dorsal surface of each anal lobe covered with sclerotised protuberances, each with an associated microtubular duct; inner margin of each lobe with sclerotised teeth. Cauda present, 16–18 µm wide, about twice as long as wide. An area of dense sclerotisation present submedially on segment VII.</p> <p> <b>Comment.</b> <i>P. m a r t i n i</i> differs from <i>P. baloghi</i> in having (character-states on latter species in brackets): (i) only one large spine on margin of each abdominal segment (two), (ii) trochanter with aone flagellate seta and one spinose seta (both flagellate), and (iii) most dorsal spinose setae having 2 or 3 associated microtubular ducts (occasionally 1 on <i>P. baloghi</i>).</p> <p> <b>Etymology</b>. The new species is named after Dr. Jon Martin (UK) in acknowledgement of his scale insect collecting efforts, which have yielded several important new taxa in Eriococcidae.</p> <p> <b>Adult male</b> (Fig. 2)</p> <p>Described from 1 specimen.</p> <p> <b>Mounted specimen.</b> Apterous; quite small, total-body length about 1.03 mm; antennae short, each about 1/3rd total-body length; body setose, with a mixture of fleshy setae (fs) and hair-like setae (hs), these often difficult to separate (fs tending to be curved, rather parallel-sided, and not broadening at base; in addition, socket tending to be wider; hs tending to be sharply bent or almost straight, narrowing to a fine point (often flagellate) and broadening at base; their basal socket also tending to be narrower and more obvious); hs very variable in size; fs and hs also present on antennae and legs; with only 1 pair of loculate pores, each with several loculi, present dorsally on head anterior to each dorsal simple eye; no other pores present. Constriction between head and thorax distinct, but none present between thorax and abdomen. Thorax and abdomen membranous apart from penial sheath; glandular pouches and glandular pouch setae absent but with some large hs present on margin of segment VIII. Legs well developed and hirsute. Penial sheath with a distinct constriction about half-way along length.</p> <p> <b>Head.</b> Roundly oval in shape; length about 160 μm, width across genae about 220 μm. Median crest absent both dorsally and ventrally; lateral arms also absent; preoccipital ridge (por) poorly defined, indicated by a series of diverging, shallow dermal ridges; with (on each side) 3 fs + 10 hs dorsal head setae (dhs) + 1 quite large loculate pore (about 6 μm widest, with about 5 or 6 odd-shaped loculi) just anterior to each dorsal simple eye. Hair-like setae (hs) variable in size, longest about 55 μm long; most fleshy setae (fs) about 30 μm long. Dorsal and ventral midcranial ridges absent. Genae (g) not reticulated; each with 5 hs and 4 fs genal setae dorsolaterally. Eyes: with two pairs of round, simple eyes; dorsal eyes (dse) placed slightly more anterolaterally than ventral simple eyes (vse); both dse and vse 25–27 μm wide. Ocelli (o) pronounced, situated rather dorsally, quite large and oval, each about 17 μm long and 8 μm wide; not quite touching postocular ridge (pocr) posteriorly. Ocular sclerite (ocs) lightly sclerotised; faintly reticulated; more dorsal and lateral reticulations each generally with a central cone-like structure; those around vse more elongate and microridges appearing more as dots. Preocular ridge (procr) very short or poorly developed. Postocular ridge (pocr) strongly developed, extending antero-laterally from posterior to each dorsal simple eye past each ocellus and then postero-laterally to near mouth. Interocular ridges absent. Dorsal ocular setae absent. Ventral midcranial ridge setae (vmcrs) plus ventral head setae (vhs) extending all around vse, with, on each side about 15 fs + 12 hs, some of latter quite long. Preoral ridge absent. Cranial apophysis probably about 45 μm long, with a single blunt apex.</p> <p> <b>Antennae</b>: 9 segmented; length 355 μm. Scape (scp) 50 μm long, 65 μm wide, with 1 or 2 fs + 5 hs. Pedicel (pdc): length 50 μm, width 65 μm; without concentric ridges on distal end; with about 15 setae, mainly fs, + a campaniform pore. Segment III very narrow basally near Johnston’s Organ and then broadening rapidly; width about 60 μm widest; segments IV–VIII similar to pedicel but not as narrow proximally, each about 60 μm widest: fs each 30–35 μm long, hs about 35 μm long; lengths of segments (μm): III 58, IV–VIII 40 –43, IX 45. Approximate number of setae per segment: III–VIII about 15, mostly fs; segment VIII with a large antennal bristle (ab) near apex; IX about 20 fs, 5 hs + 3 ab; capitate setae absent.</p> <p> <b>Thorax</b>. Apterous, and therefore thorax and abdomen membranous and fused but segmentation clear. All thoracic segments with bands of setae dorsally, mainly hs, those near margin longest, each up to 85 μm long but most hs about 30–35 μm long; fs about 30–35 μm long. <b>Prothorax</b> dorsally with fairly well-developed pronotal ridge (prnr) but not fused dorsally; pronotal sclerite (prn) represented by a dorsolateral widening of prnr; lateral pronotal setae absent, all setae located medially and probably equivalent to medial pronotal setae (mpns). Ventrally with 2 pairs of long hs + 2 pairs fs, probably equivalent to prosternal setae (stn1s). Prosternum absent. <b>Meso- and metathorax</b> without sclerites; meso- and metasternal apophyses distinct. Mesothorax ventrally with a large group of setae around spiracle and across segment, mostly fs; metathorax with two smaller groups medially (probably equating to anterior and posterior metasternal setae); marginal areas of all thoracic segments mainly without setae. Spiracles: width of mesothoracic (sp2) peritreme 28–30 μm; metathoracic (sp3): peritreme 30 μm.</p> <p> <b>Legs.</b> Subequal in length, broad and strong. Lengths (μm): coxae (cx) I 112; II 120; III 125; setae of coxa III: about 13 fs + 6 hs; long seta on each coxa not differentiated but about 35 μm long. Trochanter (tr) + femur (fm): I 203; II 200; III 210; trochanter III with 2 fs + 6 hs; each trochanter with 3 oval sensoria in a groove on each side; long trochanter seta not differentiated; femur III with about 23 setae, setal type hard to differentiate. Tibia (ti): I 135; II 145; III 140; tibia III with many setae, mainly hs, a few becoming spur-like on distal third of leg; all legs with 2 distal spinose spurs (tibs) on each tibia; length of longest 35–37 μm. Tarsi (ta) 2 segmented (length of both segments combined): I 100; II 90; III 93 μm long (ratio of length of tibia III to length of tarsus III 1:0.66); tarsus III with many setae, mainly spur-like; tarsal spurs setose, about 55 μm long; tarsal campaniform pore present; tarsal digitules (tdt) capitate, significantly longer than claw. Claws (c) long and thin, subequal in length to width of tarsus, slightly curved, without a denticle; length: III: 40 μm; claw digitules (cdt) each capitate, long and thin.</p> <p> <b>Abdomen</b> Membranous throughout, shape similar to thoracic segments but narrower; dorsal abdominal setae (ads) and dorsal pleural setae (dps) similar to those on thorax but former slightly less frequent but with more fs; ventral abdominal setae (avs) and ventral pleural setae (vpls) few. Caudal extensions on segment VII absent. Segment VIII with a short and rounded caudal extension (ce), without either glandular pouches or glandular pouch setae, but with a large group of fs + about 4 long hs in pleural position, longest about 75 μm long.</p> <p> <b>Genital segment</b>: segment IX + penial sheath (ps) longer than broad, length from anterior border to apex: 150 μm, width across anterior margin 125 μm; divided into anterior broad basal section (probably equating to segment IX) and narrower, pointed posterior part, penial sheath proper. Anterior section dorsally with anal (an) opening, about 17 μm wide, and with 0 or 1 fs; dermal surface ventrally with spinulae. Penial sheath (ps) with (on each side of anus) about 8 fs (each about 50 μm long), plus two strong hs, outer about 40 μm long, inner about 60 μm long; ventrally with a line of 3–5 hs along anterior margin, each about 35–40 μm long. Aedeagus (aed) parallel-sided, blade-like, about 83 μm long and about 12 μm wide; basal rod (bra) short but distinct. Apex of penial sheath with a few small sensoria along margins.</p> <p> <b>Comment.</b> The adult male of <i>P. m a r t i n i</i> is a fairly typical eriococcid apterous male, as far as our knowledge goes. It has simple eyes and ocelli, antennae more than 3 segmented, clubbed claw and tarsal digitules, claws rather broad basally and it has tibial spurs. In comparison, the apterous male of <i>Pseudochermes fraxini</i> <b>FIGURE 2.</b> <i>Pseudomontanococcus martini</i> Kozár and Konczné Benedicty <b>sp. n.</b>, adult male. Where A = loculate pore; B = cranial apophysis; D = microridges on dorsal surface of ocular sclerite; D = hair-like seta; E = microridges on ventral surface of ocular sclerite; F = fleshy seta; G = distal end of metathoracic leg; ab = antennal bristle; ads = dorsal abdominal setae; aed = aedeagus; an = anus; avs = ventral abdominal setae; bra = basal rod; c = claw; ce = caudal extension on segment abdominal VIII; cdt = claw digitule; cx = coxa; dhs = dorsal head setae; dps = dorsal pleural setae; dse = dorsal simple eye; fem = femur; fs = fleshy seta; g = gena; hs = hair-like seta; mpns = median pronotal setae; o = ocellus; pdc = pedicel; prn = pronotal sclerite; prnr = pronotal ridge; pocr = postocular ridge; por = postoccipital ridge; procr = preocular ridge; ps = penial sheath; scp = scape; sp2 = anterior spiracle; sp3 = posterior spiracle; stn1s = prosternal setae; ta = tarsus; tdt = tarsal digitules; ti = tibia; vhs = ventral head setae; vmcrs = ventral midcranial ridge setae; vps = ventral pleural setae; vse = ventral simple eye,.</p> <p> (Kalt.) appears even more female like, with no cervical constriction or pronotal ridge, no ridges or sclerites on the head and the abdomen tapers to the penial sheath (Afifi, 1969). In having the latter characters (a cervical contriction, pronotal ridge and distinct ocular ridges, and ocular sclerite), the adult male of <i>P. martini</i> bears some general resemblance to the brachypterous males of <i>Gossyparia spuria</i> (Mod.) and <i>G. salicicola</i> Borchsenius (Afifi, 1969), although the latter do have a sclerotised mesothorax, short wings, and glandular pouches.</p> <p> <b>Second-instar female</b> (Fig. 3).</p> <p>Described from two specimens.</p> <p> <b>Mounted specimens</b>. Body elongate oval, 0.742–0.782 mm long, 0.433–0.484 wide. Antennae 6 segmented, segment lengths (Μm): I 37 – 41, II 24–28, III 19– 20, IV 10–12, V 17, VI 22–25; each segment with few setae; segment II with one sensory pore; segments IV & V each with a falcate sensory seta 26–32 Μm long; apical segment with three sensory falcate setae, each 35–39 Μm long; apical seta 48–50 Μm long. Frontal tubercle or lobe absent. Eye visible on margin.</p> <p> <b>Venter</b>. Labium 48–50 Μm long; with a total of 6 pairs of setae, all spinose, basal segment with one pair of setae. Stylet loop reaching metacoxae. Legs long and well-developed (lengths in Μm), anterior legs: coxae 43, trochanter 39–41, femur 68–75, tibia 55–56, tarsus 58–59, claw 20–21; middle legs: coxae 43–46, trochanter 35–40, femur 71–75, tibia 51–55, tarsus 54–57, claw 20–22; hind legs: coxae 45–50, trochanter 33–34, femur 72–76, tibia 55–57, tarsus 55–60, claw 22–25; tarsal digitules knobbed, each 30–39, claw digitules each 20– 22 and slightly knobbed. All coxae with spinulae on anterior surface; posterior coxae also with some translucent pores on posterior surface. Tarsi each with a sensory pore. Claw without a denticle. Legs with a few spinose setae. Anterior and posterior spiracles similar, each with a group of seven-locular pores in atrium; diameter of anterior peritreme 12 Μm. Multilocular pores mostly with 7 loculi (range 5–9), each pore 4 µm in diameter; scattered in a submedian band extending on to ventral anal lobe, and present submedially on head and near mouthparts. Abdomen with a few spinose, hair-like setae, plus several spine-like setae. Cruciform pores absent. Microtubular ducts few, present in a submarginal band. Macrotubular ducts absent. Each anal lobe with a large spinose suranal seta 38–48 Μm long, a short spinose ventral seta medially 17 Μm long, and a long flagellate seta on outer submargin, 107–113 Μm long.</p> <p> <b>Dorsum</b>. Dorsal setae conical and spinose, of two sizes: largest setae on margin, robust, with 1 on margin of each segment, each 22–23 Μm long, plus 2–6 medially on each segment in a median longitudinal line, each 18–20 Μm long. Short spinose setae, each 6–7 µm long, scattered in two longitudinal submedian rows, each row with 1 or 2 per segment. Macrotubular ducts absent. Microtubular ducts each 8–9 µm long, and less than 1 Μm wide, with a bilocular inner end; scattered among dorsal setae, and with 1 to 3 usually present at base of each larger spine. Anal ring unclear, 34–38 Μm wide, 28–30 Μm long, with a row of pores along outer margin; with 6 long flagellate setae, each about 72 Μm long. Anal lobes sclerotized, each 87 (80–87) µm long; and 89 (77–89) µm wide, with a spinose seta on both inner and outer margins, inner seta 14–16 µm long, outer seta 24–26 Μm long, plus a longer apical seta, 24–26 Μm long. Anal lobes covered by sclerotized protuberances; inner margin with sclerotised teeth. Cauda present medially at base of anal lobes, 13 µm wide, about twice as long as wide.</p> <p> <b>Comment.</b> The second-instar female differs from the second-instar male in the absence of macrotubular ducts, the presence of translucent pores on the posterior coxae, and the shorter sensory setae on the apical segment of the antennae.</p> <p> <b>Second-instar male</b> (Fig. 4).</p> <p>Described from three specimens.</p> <p> <b>Mounted specimens</b>. Body elongate oval, 0.742–0.834 mm long, 0.433–0.505 wide. Antenna 6 segmented, segment lengths (Μm): I 35 – 38, II 27–30, III 18– 21, IV 10–14, V 10– 15, VI 22–24; each with few setae; segment II with a sensory pore; segments IV & V each with a falcate sensory seta 24–38 Μm long; apical segment with three sensory falcate setae, each 54–59 Μm long; apical seta 59–65 Μm long. Frontal tubercle or lobe absent. Eye visible on margin.</p> <p> <b>Venter</b>. Labium 57–63 Μm long; with a total of 6 pairs of setae, all spinose; basal segment with one pair of setae. Stylet loop reaching metacoxae. Legs long and well-developed (lengths in Μm): front legs: coxae 39– 44, trochanter 38–40, femur 75–80, tibia 56–59, tarsus 59–60, claw 22; middle legs: coxae 40–44, trochanter 40–44, femur 73–75, tibia 55–57, tarsus 57–60, claw 20; hind legs coxae 45–48, trochanter 38–42, femur 77– 80, tibia 57–60, tarsus 60–67, claw 23; tarsal digitules knobbed, each 28–34, claw dig

Pseudomontanococcus Kozar and Hodgson, gen. nov.

<i>Pseudomontanococcus</i> Kozár and Hodgson gen. nov. <p> <b> Type species: <i>Pseudomontanococcus martini</i></b> Kozár and Konczné Benedicty <b>sp. nov.</b></p> <p> <b>Diagnosis.</b> Adult female: antennae 6 segmented. Frontal lobe or tubercle absent. Eye present on margin. <b>Venter</b>. Labium 3 segmented, although divisions of segments unclear (more so on immature stages), basal segment with 1 pair of spinose setae; other labial setae also spinose. Stylet loop reaching metacoxae. Legs long and well-developed; tarsal and claw digitules slightly knobbed. All coxae with spinulae on anterior surface; metacoxae with translucent pores on posterior surface. Trochanter with two pores on each side. Claws without a denticle. Legs each with a few spinose setae, and with one sensory pore on each tarsus. Spiracles with a large group of seven-locular pores in atrium. Multilocular pores each with 5–9 loculi, scattered throughout venter but abundant in a wide longitudinal submedian band. Abdomen with a few flagellate setae, plus several small spinose setae. Cruciform pores absent. Microtubular and macrotubular ducts present throughout. A weak area of sclerotisation present submedially on segment VII. Dermal spinules present throughout. Anal lobes each with a strong spinose suranal seta, 1 short spine-like seta, plus a long flagellate seta in submarginal position. <b>Dorsum</b>. Dorsal setae strong, conical and spine-like; of two sizes: larger along margin, and in a longitudinal median band; short spinose setae sparse on all segments. Macrotubular ducts numerous; microtubular ducts scattered among dorsal setae and often with 1–3 present at base of larger spines. Dermal spinules present on both dorsum and venter. Anal ring with a sparse row of pores along outer margin, and with eight long setae. Anal lobes strong and sclerotized, each lobe longer than wide, with a spine-like seta on both inner and outer margins, plus an apical seta; entire dorsal surface of anal lobes covered with sclerotised protuberances. Cauda present, about twice as long as wide. An area of sclerotisation present submedially on segment VII on both dorsum and venter.</p> <p> <b>Comment</b>. The adult females of <i>Pseudomontanococcus</i> share with <i>Montanococcus</i> Henderson (2007) the ventral longitudinal band of multilocular pores. They differ in having (character-states on <i>Montanococcus</i> in brackets): (i) anal lobe setae large and spinose (anal lobe setae relatively small); (ii) basal segment of labium well-developed (basal segment apparently absent); (iii) large conical spinose setae present in a median longitudinal band on dorsum (conical setae very short, in transverse rows); (iv) a dorsal sclerotized plate on penultimate abdominal segment (absent); (v) 1–3 microtubular ducts associated with base of all or most large spinose setae (less clearly associated with setal bases); (vi) a well-developed, sclerotised cauda or dorsal median plate present (absent); (vii) anal lobes rather large but of normal tapering shape (anal lobes massive, with additional lobules); (viii) anal ring setae long and hair-like (anal ring setae sword-shaped on two species but flagellate on 3rd), and (ix) anal ring with only a few large pores, each variable in shape (anal ring with many pores of rather uniform shape and size).</p>Published as part of <i>Kozár, Ferenc, Benedicty, Zsuzsanna Konczné & Hodgson, Chris, 2008, A new felt scale genus (Hemiptera: Sternorrhyncha; Eriococcidae) from Papua New Guinea, with descriptions of two new species, pp. 47-62 in Zootaxa 1934</i> on page 48, DOI: <a href="http://zenodo.org/record/184923">10.5281/zenodo.184923</a&gt

Eriococcus szentivanyi Kozar and Williams, sp. nov.

Eriococcus szentivanyi Kozár and Williams sp. nov. (Fig. 2) Type material. Holotype adult female: Malaysia, Sabah, Mt Kinabalu, 3230 m, extracted from “common trap”, in Leptospermum forest, 29.x. 1982, coll. M. Horak, MEF 8286, deposited in BMNH. Description. Body of slide-mounted adult female, elongate-oval, about 1.43 mm long and 0.78 mm wide. Antennae 6 segmented, segment III longest but with a partial division; segment lengths (µm) I 34, II 36, III 96, IV 22, V 22, VI 31; each segment with few setae, segment II with sensory pore, segment III with almost parallel sides; apical segment with 4 sensory falcate setae, each 24–27 µm long; segments IV and V each with single falcate seta, each 24–29 µm long. Frontal lobes present next to basal antennal segments. Eyes present, near margin. Venter. Labium 3 segmented, about 77 µm long; basal segment with 1 pair of robust setae; setae on distal segment long and capitate. Stylet loop very short, about as long as labium. Legs well developed, long, lengths in (µm): forecoxae each 72, trochanter 48, femur 142, tibia 96, tarsus 110, claw 27. Midcoxae each 77, trochanter 43, femur 135, tibia, 96, tarsus 115, claw 26. Hindcoxae each 96, trochanter 48, femur 147, tibia 101, tarsus 106, claw 30. Tarsal digitules knobbed, 49–55 µm long; claw digitules 32–36 µm long, slightly knobbed. Coxae without spinulae. Hindcoxae and femora each with large irregularly-shaped translucent pores. Each trochanter with 2 sensory pores on each side. Claws each without a denticle. Leg segments with few setae, all flagellate; tarsi each with a sensory pore at proximal end. Spiracles each about 21 µm in diameter, each with a large group of 5–10 locular pores near opening. Multilocular disc pores of 2 sizes; smaller about 4 µm in diameter, larger 5 µm in diameter with 7–10 loculi, present in small numbers over most of surface. Abdominal setae flagellate, each short, longest about as long as width of segment. Minute spine-like setae fairly numerous in a submarginal band and on frons. Microtubular ducts absent. Macrotubular ducts of 2 sizes, each about 4–6 µm wide, 19 µm long, ducts found more medially slightly narrower; each duct with inner ductule as long as outer ductule and with inner gland flower-shaped. Oviduct or internal genital organ not easily discernible. Venter of each anal lobe with 1 subapical seta, one suranal setae near inner margin and one more seta near outer margin. Dorsum. Marginal setae robust, spine-like, weakly truncate, each 41–43 µm long, present singly on margin of each abdominal segment; also with about 16 robust, bluntly-tipped spine-like setae on head margin, each subequal to those posteriorly; spine-like marginal setae absent along thoracic margins. Other dorsal setae minute, blunt and spine-like, each 3–7 µm long, with a rather broad basal socket; mainly present across middle of segments. Macrotubular ducts present, similar to those on venter, fairly evenly distributed throughout dorsum. Microtubular ducts present, each about 5 µm long, each with an oval orifice surrounded by small flat sclerotized area with two wing-like extensions; sparsely scattered among dorsal setae. Loculate pores absent. Anal ring sclerotized, 48 µm long, 45 µm wide, with a sparse row of pores plus 6 flagellate anal ring setae, each about 94 µm long. Anal lobes well developed, about as wide as long, heavily sclerotized and each dentate along inner margin; each with 2 robust truncate setae about 20–24 µm long, and with a very short stout seta on outer margin near base, about 4 µm long. Apical setae robust and flagellate, each 110 µm long. Cauda present, sclerotized, about 48 µm wide. Comments. Eriococcus szentivanyi differs from other species presently included in the genus Eriococcus Targioni Tozzetti sensu lato by the capitate setae on the labium, instead of the usual flagellate setae. Apparently, similar setae are found on the labium in the family Monophlebidae (Koteja, 1974) and in the mealybug genus Dicranococcus Williams. E. szentivanyi is similar to Eriococcus araucariae Maskell in possessing a marginal series of stout spine-like setae but these are noticeably longer than the other dorsal setae. Furthermore, (i) the marginal spinose setae are absent from the thorax of E. szentivanyi whereas they are present on E. araucariae; (ii) there is only one pair of setae present on the basal segment of labium of E. szentivanyi but there are two pairs on E. araucariae; (iii) E. szentivanyi possesses a cauda (apparently absent on E. araucariae) and (iv) E. szentivanyi lacks trilocular pores (present on E. araucariae). Etymology. The new species is named after József Szent Ivány, who contributed much to the knowledge of the Coccoidea and other insect groups of Papua New Guinea.Published as part of Kozár, Ferenc, Williams, Douglas J. & Benedicty, Zsuzsanna Konczné, 2009, A new genus and four new species of the scale insect family Eriococcidae (Hemiptera: Coccoidea) from the Austro-Oriental Region., pp. 1-15 in Zootaxa 1979 on pages 5-7, DOI: 10.5281/zenodo.18524

Sangicoccus

Key to adult female species of Sangicoccus 1. Setae in large marginal groups on dorsal margin bluntly pointed............................................... obtusispinus (Reyne) - Setae in large marginal groups on dorsal margin truncate-conical............................................................................... 2 2. Modified microtubular ducts forming a wide longitudinal band medially on dorsum.............................. reynei sp. n. - Modified microtubular ducts present throughout, not restricted to medially on dorsum............................................ 3 3. Groups of 7 locular pores present on ventral margins ......................................................................... morrisoni sp. n. - Groups of 7 locular pores absent from ventral margins ............................................................ truncatispinus (Reyne)Published as part of Kozár, Ferenc, Williams, Douglas J. & Benedicty, Zsuzsanna Konczné, 2009, A new genus and four new species of the scale insect family Eriococcidae (Hemiptera: Coccoidea) from the Austro-Oriental Region., pp. 1-15 in Zootaxa 1979 on page 14, DOI: 10.5281/zenodo.18524