The Roles of Platelet GPIIb/IIIa and αvβ3 Integrins during HeLa Cells Adhesion, Migration, and Invasion to Monolayer Endothelium under Static and Dynamic Shear Flow

During their passage through the circulatory system, tumor cells undergo extensive interactions with various host cells including endothelial cells and platelets. Mechanisms mediating tumor cell adhesion, migration, and metastasis to vessel wall under flow condition are largely unknown. The aim of this study was to investigate the potential roles of GPIIb/IIIa and αvβ3 integrins underlying the HeLa-endothelium interaction in static and dynamic flow conditions. HeLa cell migration and invasion were studied by using Millicell cell culture insert system. The numbers of transmigrated or invaded HeLa cells significantly increased by thrombin-activated platelets and reduced by eptifibatide, a platelet inhibitor. Meanwhile, RGDWE peptides, a specific inhibitor of αvβ3 integrin, also inhibited HeLa cell transmigration. Interestingly, the presence of endothelial cells had significant effect on HeLa cell migration regardless of static or cocultured flow condition. The adhesion capability of HeLa cells to endothelial monolayer was also significantly affected by GPIIb/IIIa and αvβ3 integrins. The arrested HeLa cells increased nearly 5-fold in the presence of thrombin-activated platelets at shear stress condition (1.84 dyn/cm2 exposure for 1 hour) than the control (static). Our findings showed that GPIIb/IIIa and αvβ3 integrins are important mediators in the pathology of cervical cancer and provide a molecular basis for the future therapy, and the efficient antitumor benefit should target multiple receptors on tumor cells and platelets

Sr-Nd isotopic geochemistry of Holocene sediments from the South Yellow Sea: Implications for provenance and monsoon variability

Elemental geochemical and Sr-Nd isotopic signatures are used to decipher terrigenous sediments provenances and transport mechanisms in the South Yellow Sea during the Holocene. Sr-87/Sr-86 ratios in the Chinese and Korean riverine sediments overlap each other, whereas epsilon Nd values of Korean riverine sediments are generally less radiogenic in comparison to the Changjiang and Huanghe. Moreover, eNd values of these two large rivers appear unaffected by mineral sorting and are relative stable during the Holocene. We propose a three end-members (i.e., the Changjiang, the Huanghe, and Korean rivers) mixing model to explain sediment provenances in the Central Yellow Sea Mud (CYSM). Mixing calculations show that the Huanghe is the major sediment contributor to the CYSM before similar to 8 ka (thousand years before 1950 CE), whereas the Changjiang has become the predominant sediment source after similar to 8 ka. Holocene changes in riverine sediment supplies to the CYSM are closely related to the oceanic circulation, monsoon climate, and drainage changes. After examining several hypotheses to explain the variations in Sr-87/Sr-86 ratios of Core YSC-1 during the past similar to 8 kyr, we tentatively attribute that to changes in the erosion patterns of the Changjiang Basin. This in turn is associated with the asynchronous evolution of monsoon precipitation in the upper (Indian Summer Monsoon) and middle-lower Changjiang (East Asian Summer Monsoon). Therefore, our results highlight significant influences of monsoon climate on erosion patterns within the Changjiang catchment at millennial timescales

Boosting Few-Shot Text Classification via Distribution Estimation

Distribution estimation has been demonstrated as one of the most effective approaches in dealing with few-shot image classification, as the low-level patterns and underlying representations can be easily transferred across different tasks in computer vision domain. However, directly applying this approach to few-shot text classification is challenging, since leveraging the statistics of known classes with sufficient samples to calibrate the distributions of novel classes may cause negative effects due to serious category difference in text domain. To alleviate this issue, we propose two simple yet effective strategies to estimate the distributions of the novel classes by utilizing unlabeled query samples, thus avoiding the potential negative transfer issue. Specifically, we first assume a class or sample follows the Gaussian distribution, and use the original support set and the nearest few query samples to estimate the corresponding mean and covariance. Then, we augment the labeled samples by sampling from the estimated distribution, which can provide sufficient supervision for training the classification model. Extensive experiments on eight few-shot text classification datasets show that the proposed method outperforms state-of-the-art baselines significantly.Comment: Accepted to AAAI 202

Measuring cell deformation by microfluidics

Microfluidics is an increasingly popular method for studying cell deformation, with various applications in fields such as cell biology, biophysics, and medical research. Characterizing cell deformation offers insights into fundamental cell processes, such as migration, division, and signaling. This review summarizes recent advances in microfluidic techniques for measuring cellular deformation, including the different types of microfluidic devices and methods used to induce cell deformation. Recent applications of microfluidics-based approaches for studying cell deformation are highlighted. Compared to traditional methods, microfluidic chips can control the direction and velocity of cell flow by establishing microfluidic channels and microcolumn arrays, enabling the measurement of cell shape changes. Overall, microfluidics-based approaches provide a powerful platform for studying cell deformation. It is expected that future developments will lead to more intelligent and diverse microfluidic chips, further promoting the application of microfluidics-based methods in biomedical research, providing more effective tools for disease diagnosis, drug screening, and treatment

Hydroporus tuvaensis Pederzani 2001

Hydroporus tuvaensis Pederzani, 2001 This species was described by Pederzani (2001: 234) after a small series of specimens from Sush village (ca. 52.05 N 94.17 E), ca. 50 km NW Kizyl city, Tuva district, Russia, and about 200 km north of the border to the eastern part of Mongolia; altitude ca. 800–1000 m (triangle in Figs. 13 and 14). We had the opportunity to study one male paratype (Fig. 3) and two female paratypes (CHH, CHF). This is the largest of the five species of the complex (Pederzani 2001: 238 provided: TL: 3.70–3.88 mm, TL/MW: 1.94–2.03). Additionally, it has the entire upper surface shiny and, thus, cannot be mixed up with H. sejilashan sp. n. As in H. tibetanus and H. sejilashan sp. n. the anterior protarsal claw is simple and more or less shaped as the posterior one. Pederzani (2001: 216) reported seven females from Mongolia with doubtful identities, which are distinctly smaller than the "normal" H. tuvaensis. We have studied these and a few further similar specimens (CHF, CHS, CJH, NMB) from that region—including males—and are not yet sure about their identity.Published as part of Jia, Fenglong, Zhao, Shuang & Fery, Hans, 2012, Hydroporus sejilashan sp. n., a new diving beetle of the acutangulus - complex from Xizang, China (Qinghai-Tibet Plateau), and notes on other taxa of the genus (Coleoptera, Dytiscidae, Hydroporinae), pp. 55-67 in Zootaxa 3223 (1) on page 64, DOI: 10.11646/zootaxa.3223.1.4, http://zenodo.org/record/21083

Hydroporus tibetanus Zaitzev 1953

Hydroporus tibetanus Zaitzev, 1953 Hydroporus tibetanus was described by Zaitzev (1953: 169; see also the English translation Zaitzev 1972: 179) from Amdo (China, Qinghai [Tibet]; ca. 32.27 N 91.68 E; altitude ca. 4700 m; star "a" in Figs. 13–14) as "closely related to" Hydroporus nigrita (Fabricius, 1972) and, thus, was treated as member of the nigrita -group by Nilsson (2001: 163). Zaitzev seemingly overlooked the shape of the metacoxal lines and the setation on the metatrochanters; he wrote "... dorsum and venter more shining; body markedly more weakly shagreened [compared with H. nigrita]..." Shaverdo (2004: 260) studied the two syntypes of H. tibetanus (ZISP), designated a male lectotype (Fig. 4) and found out that—due to the widely smooth elytra—this species belongs to the planus - group and, in particular, to the acutangulus -complex because of the parallel metacoxal lines and the setal fringe on each metatrochanter. We have also studied the types of this taxon and can confirm Shaverdo's observations, but want to add that the base of the median lobe as well as the right paramere of the lectotype are both damaged. Additionally, it must be mentioned that the lectotype lacks the posterior claw of the left protarsus (the anterior claw is present and clearly not strongly curved near base), the anterior claw of the right protarsus, the last three tarsomeres of the left mesotarsus, the last three tarsomeres of the left metatarsus and the complete right metatarsus. We want to point to the fact that the lectotype is reticulated more or less as typical H. acutangulus: entire surface shiny; pronotum smooth on disc, reticulate laterally; elytra smooth in first third, behind with some indistinct short lines between punctures, before apex with distinct small meshes, but also here not matted. In the female paralectotype the elytral reticulation with small meshes is already present in the second third and becomes more impressed towards the apex; the smooth areas on the pronotum are even more extended than in the male. The entire upper surface appears by no means less shiny than that of the male. After Zaitzev's description (1953), no subsequent specimens of H. tibetanus have been collected. The species was only listed in some catalogues (Hua 2002, Nilsson 1995, 2001, 2003, 2011, Nilsson & Fery 2006) or treated in overviews of members of the nigrita - or planus -groups (Fery & Pesic 2006, Guéorguiev 1966, Shaverdo 2004). Recently, however, we have studied several specimens which we could determine as this species. The collecting data are as follows: 3.7. 1999, Sichuan, Ganzi (= Garzê) Tibetan Autonomous Region, Batang County, Shaluli Mountains, 57 km NE Batang, pools, ca. 4500 m, A. Pütz leg. (CAP, CHF) (star "b" in Figs. 13–14). These specimens are similar to the lecto- and paralectotype of H. tibetanus in most respects, in particular, their upper surface is shiny in both sexes. They are, however, somewhat darker than the types. Some females are reticulated on the entire elytra and large parts of the pronotum. Other females have the upper surface reticulated as the female paralectotype of H. tibetanus. The median lobe of the males in ventral view (Fig. 8 b) is similar to that of the lectotype of H. tibetanus (Fig. 8 a). We are not absolutely sure about the identity of Pütz' specimens, but since no other material from the region around Amdo has become available, we assume that this population belongs to H. tibetanus. Measurements: Lectotype (male): TL: 3.2 mm, MW: 1.6 mm, TL/MW: 2.00; paralectotype (female): TL: 3.2 mm, MW: 1.65 mm, TL/MW: 1.94; specimens from Ganzi: TL: 2.95–3.3 mm, MW: 1.5–1.7 mm, TL/MW: 1.9–2.03. We prefer not to present a key to species at this time because any key for this group would be rather primitive and not very useful. Construction of a key for this species group should be postponed until the variability of all group members is known sufficiently. We compare below the new species with the types of H. tibetanus as well as with specimens from the population of H. tibetanus from Ganzi: - Hydroporus sejilashan sp. n. is larger and slightly less elongate than H. tibetanus. - In dorsal view the sides of the pronotum are less curved in H. sejilashan sp. n. In H. tibetanus the base of the pronotum is not broader than the base of the elytra. - The new species has the entire surface reticulated and matt. Hydroporus tibetanus has a shiny surface (also in reticulated areas) and in the males and most females the reticulation of the elytra is restricted to the apex. - Hydroporus sejilashan sp. n. has the dorsal and ventral surface almost totally black, only the tarsi and first antennomeres are dark brownish. The lectotype of H. tibetanus has the tarsi somewhat lighter, the antennomeres are all more or less uniformly brown, and the elytra are dark brown; the female paralectotype and the specimens from Ganzi, however, are generally darker than the lectotype. - The punctation of the surface is distinctly coarser in H. tibetanus, and also denser on head; in the posterolateral depression of the pronotum and before the posterior margin (except centrally) it is very coarse and longitudinally deformed; the punctures of the irregular puncture line behind the anterior pronotal margin are also very coarse and in part longitudinally deformed. In the new species, however, the punctation is generally simple and by far not so coarse in the respective areas. Due to the coarser elytral punctation, the elytral puncture lines appear slightly less distinct in H. tibetanus. - The first three protarsomeres are more dilated in H. sejilashan sp. n.; especially the second protarsomere is almost two times as broad as long and the third one more or less of round shape; the differences between male and female protarsomeres are obvious. In the lectotype of H. tibetanus the second protarsomere is only one and a half times as broad as long and the third one is more elongate; differences between the protarsomeres of males and females are rather slight. The male mesotarsomeres of the new species are also broader than those of the females and broader than those of H. tibetanus. - Antennomeres five to eleven are slightly longer in the new species. - The median lobe of H. sejilashan sp. n. is in ventral view slightly narrower in the apical third and more pointed (compare Figs. 6 a with 8 a–b). In lateral view (Fig. 6 b) a comparison is impossible because the median lobe of the lectotype of H. tibetanus is partly destroyed. A comparison with that of a male from Ganzi (Fig. 8 c) shows that the lobe of the new species is less bent near the base and slightly broader in apical third.Published as part of Jia, Fenglong, Zhao, Shuang & Fery, Hans, 2012, Hydroporus sejilashan sp. n., a new diving beetle of the acutangulus - complex from Xizang, China (Qinghai-Tibet Plateau), and notes on other taxa of the genus (Coleoptera, Dytiscidae, Hydroporinae), pp. 55-67 in Zootaxa 3223 (1) on pages 61-62, DOI: 10.11646/zootaxa.3223.1.4, http://zenodo.org/record/21083

Hydroporus acutangulus Thomson 1856

Hydroporus acutangulus Thomson, 1856 The species was described by Thomson (1856: 202) after specimens from northern Sweden, "Lappland" (very roughly estimated to 67 N 26 E; circle (1) in Fig. 13). The type localities of the five other known synonyms are also given as circles in Fig. 13: (2) Hydroporus punctatulus J. Sahlberg, 1889 (Russia, Kola peninsula, Chavanga; 66.113 N 37.777 E); (3) H. sumakovi (Russia, Novaya Zemlya, Belushya Guba; 71.535 N 52.327 E) (circle with question mark, see notes below); (4) H. pectoralis (Russia, Siberia, Neuleva river, Spirino; 61.167 N 69.814 E); Hydroporus zaitzevi Jacobson, 1908 is a replacement name for H. pectoralis and, thus, has the same type locality as the latter; (5) H. aenescens (Russia, Siberia, Yenisei, Bryokhovsky Island, ca. 70.83 N 83.00E, and Nikandrovsky Island, ca. 70.67 N 83.00E). For more details about the taxa mentioned above see Nilsson (2001) and the comprehensive discussion in Falkenström (1929: 151). We have studied several specimens from Sweden, Finland, northern Norway and Russia (Republic of Karelia) (MNB, CHF). In addition we have studied the male from the Russian Far East, Primorsky krai, Ussuriysky Rayon, Kaimanovka, ca. 43.63 N 132.24 E (CLH; circle 6 in Fig. 13) which has been cited in Pederzani (2001: 236). Hydroporus acutangulus is almost entirely black or has the elytra very dark brownish (Fig. 1); all specimens studied appear at least darker than those H. polaris which are at our disposal. The males have the elytra shiny and not reticulate behind the base; however, in the posterior part of the first third, traces of reticulation can be found: not complete meshes, but short indistinct lines between the punctures. More distally, this reticulation becomes more prominent, and near the apex it is replaced by small complete meshes. In the northern European males studied the pronotum is reticulated with small meshes, except a small area on disc. In the male from the Russian Far East this smooth area is extended more to the sides and reaches until the base. Females often have distinct reticulation on large parts of the upper surface (Nilsson & Holmen 1995: 48), but are, nevertheless, shiny. Further comparative notes can be found below under H. polaris. The species can be easily distinguished from H. sejilashan sp. n. by the arcuate anterior protarsal claws (males) and the shiny surface.Published as part of Jia, Fenglong, Zhao, Shuang & Fery, Hans, 2012, Hydroporus sejilashan sp. n., a new diving beetle of the acutangulus - complex from Xizang, China (Qinghai-Tibet Plateau), and notes on other taxa of the genus (Coleoptera, Dytiscidae, Hydroporinae), pp. 55-67 in Zootaxa 3223 (1) on page 62, DOI: 10.11646/zootaxa.3223.1.4, http://zenodo.org/record/21083

Rhantus fengi sp. n. from Xizang, China, and notes on Laccoporus nigritulus (Gschwendtner) (Coleoptera, Dytiscidae)

Rhantus fengi sp. n. from Mount Sejila, Xizang, China is described and illustrated. Laccoporus nigritulus (Gschwendtner, 1936) is redescribed and illustrated; Laccoporus viator Balfour-Browne, 1939, syn. n. is established as its junior subjective synonym

Hydroporus sejilashan, sp. n.

Hydroporus sejilashan sp. n. Type locality: China, Xizang Autonomous Region, ca. 20 km SEE Linzhi [= Nyingchi], Sejilashan mountains; ca. 29.62 N 94.60 E, altitude ca. 4100–4200 m. Type material: Holotype: 3, " China, Xizang (Tibet), Linzhi, Sejilashan mountains, ca. 29.62 N 94.60 E, 4100- 4200 m, 12.– 18.8.2009, leg. Fenglong Jia" [printed], a further label with same text, but with Chinese letters, " Holotype, Hydroporus sejilashan sp. n., Jia, Zhao & Fery det." [red, printed] (SYSU). Paratypes: 121 exs, same label data, but with the respective red paratype label (SYSU, CGC, CHF). Habitus elongate oval (Fig. 5), appearing slightly parallel-sided; maximum width behind middle of total length, more or less in middle of elytral length. Dorsal and ventral surface almost entirely black, microreticulated and matt. Pronotum at posterior angles slightly broader than elytra at base (but by far not so distinctly broader than in the members of the sibiricus -group), thus, outline in dorsal view here with a slight discontinuity. Head black, with indistinct transverse dark brownish marking near anterior margin of clypeus and on vertex; above antennal cavity indistinct transparent brownish spot. Interocular distance distinctly smaller than half of pronotal width at posterior angles. Between eyes with two nearly triangular clypeal grooves. Reticulation of head distinct, on large parts with even, polygonal, more or less isodiametric meshes; at anterior margin and on vertex transversely elongate; in clypeal grooves meshes smaller and more strongly impressed. Punctation sparse, absent on vertex, diameter of punctures equalling that of meshes on clypeus; in clypeal grooves punctation coarser and denser; next to inner margin of eye with a distinct line of impressed punctures. Setation absent. First and second antennomeres dark brown; third and fourth proximally dark brown, distally black; rest of antennomeres black. First maxillary and labial palpomeres reddish brown, others largely darkened. Third and fifth antennomeres rather short, fourth still shorter, almost as wide as long; fifth to eighth about one and a half times as long as wide, ninth and tenth somewhat longer, eleventh more than two times longer than wide. Pronotum entirely black. Maximum width between posterior angles; in dorsal view sides almost straight in posterior two thirds, before curved to anterior angles. Lateral bead broader than half of diameter of antennomeres, more or less of same width over entire length. Between disc and posterior margin pronotum slightly impressed over about half of its width, thus, disc appearing somewhat vaulted; with shallow impression at posterior angles. Surface rather uniformly reticulated, meshes more or less of same diameter as on clypeus; in posterolateral impressions somewhat smaller. Punctation on disc similar to that on clypeus and frons, behind and towards sides coarser and slightly denser; middle of pronotum with a very large puncture; near anterior margin with several finer punctures, behind them a line of irregularly placed rather coarse punctures perceptible. Most punctures provided with a transparent whitish, rather long seta (ca. 0.05 mm). Elytra entirely black as pronotum; surface appearing slightly more matt than on head and pronotum. Base of elytra slightly narrower than pronotal base. In dorsal view sides of elytra rather weakly curved in anterior two thirds, behind stronger curved to apex. In lateral view elytral margin very slightly ascending towards humeral angle, straight directly before angles, far behind slightly curved; epipleuron visible until humeral angle; elytral bead distinctly thinner than pronotal bead, in lateral view both forming an angle at bases of pronotum and elytra. Reticulation on elytra more or less uniform, meshes larger than those on pronotum. Punctation also rather uniform, punctures sparse, coarser than those on head and disc of pronotum; diameter of punctures same as that of meshes; punctures spaced approximately three to four puncture widths apart. Near sides punctures somewhat smaller and still sparser. Puncture lines almost imperceptible, only visible when adequately illuminated; indicated by slightly enlarged and/or more closely placed punctures; each elytron with two discal lines and one sublateral line; in anterior half elytra alongside first (inner) line additionally slightly impressed. Sutural puncture line absent. Setation distinct, similar to that on pronotum. Ventral surface almost entirely black, including gula. Mouthparts, pro- and mesocoxae, and apex of metacoxal processes dark brownish; apex of prosternal process, hind margins of fourth to sixth abdominal ventrites narrowly transparently brownish. Genae distinctly reticulate, but shiny; gula smooth, with some coarse punctures. Prosternal process with blade narrowly lanceolate, in cross-section tectiform; sides not flattened, coarsely punctured and with long setae; margin beaded; apex not distinctly pointed, narrowly rounded; between procoxae with a weak protuberance, before protuberance with a few transverse carinae; anteriorly process not prolonged as narrow convexity onto prosternum. Prosternum anteromedially flat and very rugosely sculptured. Posterior margins of metacoxal processes straight and forming more or less one line together. Metacoxal lines subparallel, reaching posterior margin of metaventrite. Large parts of ventral surface reticulated, but appearing not so matt as upper surface; meshes mostly elongate, often weak and incomplete, only on last two abdominal ventrites more impressed; centre of metaventrite smooth. Sides of metaventrite, metacoxal plates, and first two abdominal ventrites with coarse, but not very dense punctation; smooth centre of metaventrite with sparse and fine punctures; elytral epipleura and third and fourth abdominal ventrites also sparsely covered with fine punctures; on last two ventrites punctures again coarser. Middle of third to fifth ventrite with one shallow large puncture, with several very long setae originating in it. Almost all other punctures on ventral surface bearing one rather long transparent seta. Intralinear space of metacoxal processes covered with very dense setae; this setal area prolonged anteriorly onto small posteromedial area on metaventrite (this area much more developed anteriad than illustrated for H. acutangulus in fig. 88 of Nilsson & Holmen 1995: 49). Legs with all trochanters and pro- and mesotibiae dark reddish brown, metatibae somewhat darker; femora black, proximal and distal ends indistinctly brownish. Tarsomeres more or less of same colour as respective tibiae, proximally indistinctly lighter. Measurements: Holotype: TL: 3.2 mm, TL-H: 3.0 mm, MW: 1.7 mm, TL/MW: 1.88. Paratypes: TL: 3.1–3.5 mm, TL-H: 3.0– 3.2 mm, MW: 1.65–1.75 mm, TL/MW: 1.88–1.89. Males: Median lobe of aedeagus in ventral (a) and lateral (b) view as in Fig. 6; left paramere as in Fig. 7. For comparison those of H. tibetanus are illustrated in Figs. 8–10. First three tarsomeres of male dilated; second one rather short, length only one half of width; third more or less as long as wide, outline almost circular; fifth indistinctly longer than fourth. First and second pro- and mesotarsomeres with sucker hairs and a few additional elongate oval sucker cups. Anterior protarsal claw indistinctly broader than posterior one, both more or less similarly shaped and evenly curved. (anterior one not thickened at base and here not strongly arcuate, and then straight as in H. acutangulus.) Margin of pro- and mesotrochanters with a short line of approximately six coarse punctures, each with a rather long seta. Hind margin of metatrochanter with a conspicuous fringe of light yellowish setae. Females: Similar to males, but pro- and mesotarsomeres less dilated, without sucker cups, and metatrochanters lacking setal fringe. Reticulation on ventral surface more impressed than in males, almost all meshes complete, smooth area on centre of metaventrite absent or very strongly reduced. Gonocoxosternum and gonocoxae as in Figs. 11–12. Variability: There is little variability in H. sejilashan sp. n. The brownish parts on the ventral surface are sometimes more extended. Third and fourth antennomeres are sometimes almost black, similar to following ones. The puncture lines on elytra are in some specimens more perceptible. The base of the pronotum is in some males only slightly broader than the base of the elytra; this is also the case in about half of the females. The shape of the metacoxal lines also varies a little: in some cases these are slightly diverging shortly before hind margin of the metacoxal plates, in others slightly converging. Notes: Due to the matt surface, fine punctation and weak puncture lines of the elytra, the new species at first glance resembles a dark Hydroporus nigellus Mannerheim, 1853 (belonging to the species-group of the same name) but not any of the members of the acutangulus -complex nor a single one of the planus -group. An inspection of the dorsal and ventral surfaces of the new species (of males as well as females), however, readily shows that it belongs to the acutangulus -complex—rather broad pronotal lateral bead, subparallel metacoxal lines, strong setation on their intralinear space, and a striking setal fringe on the hind margin of the male metatrochanters. Distribution: Eastern Qinghai-Tibet Plateau; so far H. sejilashan sp. n. is only known from the type locality. Etymology: The new species is named after the mountain range Sejila Shan where it has been found. It is a noun in the nominative case in apposition. Ecology: The collecting area in the Sejilashan Mountains is covered by swamps with dense bushes and grass in altitudes of 4100–4260 m. In addition, several shallow pools can be found which are due to Yak activities. The new species was found in swamps with dense vegetation, and shallow pools of stagnant and very clear water with only sparse vegetation; some specimens were collected under grass roots. It was always found together with Boreonectes emmerichi (Falkenström, 1936).Published as part of Jia, Fenglong, Zhao, Shuang & Fery, Hans, 2012, Hydroporus sejilashan sp. n., a new diving beetle of the acutangulus - complex from Xizang, China (Qinghai-Tibet Plateau), and notes on other taxa of the genus (Coleoptera, Dytiscidae, Hydroporinae), pp. 55-67 in Zootaxa 3223 (1) on pages 58-60, DOI: 10.11646/zootaxa.3223.1.4, http://zenodo.org/record/21083