How to Measure Group Selection in Real-world Populations

Multilevel selection and the evolution of cooperation are fundamental to the formation of higher-level organisation and the evolution of biocomplexity, but such notions are controversial and poorly understood in natural populations. The theoretic principles of group selection are well developed in idealised models where a population is neatly divided into multiple semi-isolated sub-populations. But since such models can be explained by individual selection given the localised frequency-dependent effects involved, some argue that the group selection concepts offered are, even in the idealised case, redundant and that in natural conditions where groups are not well-defined that a group selection framework is entirely inapplicable. This does not necessarily mean, however, that a natural population is not subject to some interesting localised frequency-dependent effects – but how could we formally quantify this under realistic conditions? Here we focus on the presence of a Simpson’s Paradox where, although the local proportion of cooperators decreases at all locations, the global proportion of cooperators increases. We illustrate this principle in a simple individual-based model of bacterial biofilm growth and discuss various complicating factors in moving from theory to practice of measuring group selection

The Efficacy of Group Selection is Increased by Coexistence Dynamics within Groups

Selection on the level of loosely associated groups has been suggested as a route towards the evolution of cooperation between individuals and the subsequent formation of higher-level biological entities. Such group selection explanations remain problematic, however, due to the narrow range of parameters under which they can overturn within-group selection that favours selfish behaviour. In principle, individual selection could act on such parameters so as to strengthen the force of between-group selection and hence increase cooperation and individual fitness, as illustrated in our previous work. However, such a process cannot operate in parameter regions where group selection effects are totally absent, since there would be no selective gradient to follow. One key parameter, which when increased often rapidly causes group selection effects to tend to zero, is initial group size, for when groups are formed randomly then even moderately sized groups lack significant variance in their composition. However, the consequent restriction of any group selection effect to small sized groups is derived from models that assume selfish types will competitively exclude their more cooperative counterparts at within-group equilibrium. In such cases, diversity in the migrant pool can tend to zero and accordingly variance in group composition cannot be generated. In contrast, we show that if within-group dynamics lead to a stable coexistence of selfish and cooperative types, then the range of group sizes showing some effect of group selection is much larger

Dynamical decompactification from brane gases in eleven-dimensional supergravity

Brane gas cosmology provides a dynamical decompactification mechanism that could account for the number of spacetime dimensions we observe today. In this work we discuss this scenario taking into account the full bosonic sector of eleven-dimensional supergravity. We find new cosmological solutions that can dynamically explain the existence of three large spatial dimensions characterised by an universal asymptotic scaling behaviour and a large number of initially unwrapped dimensions. This type of solutions enlarge the possible initial conditions of the Universe in the Hagedorn phase and consequently can potentially increase the probability of dynamical decompactification from anisotropically wrapped backgrounds.Comment: 8 figures, JHEP3 styl

Defining the gap between research and practice in public relations programme evaluation - towards a new research agenda

The current situation in public relations programme evaluation is neatly summarized by McCoy who commented that 'probably the most common buzzwords in public relations in the last ten years have been evaluation and accountability' (McCoy 2005, 3). This paper examines the academic and practitioner-based literature and research on programme evaluation and it detects different priorities and approaches that may partly explain why the debate on acceptable and agreed evaluation methods continues. It analyses those differences and proposes a research agenda to bridge the gap and move the debate forward

Advanced Computer Dormant Reliability Study Final Report

Reliability of integrated circuits and discrete components of electronics for computer and dormant module for Minuteman

Clustering properties of ultrahigh energy cosmic rays and the search for their astrophysical sources

The arrival directions of ultrahigh energy cosmic rays (UHECRs) may show anisotropies on all scales, from just above the experimental angular resolution up to medium scales and dipole anisotropies. We find that a global comparison of the two-point auto-correlation function of the data with the one of catalogues of potential sources is a powerful diagnostic tool. In particular, this method is far less sensitive to unknown deflections in magnetic fields than cross-correlation studies while keeping a strong discrimination power among source candidates. We illustrate these advantages by considering ordinary galaxies, gamma ray bursts and active galactic nuclei as possible sources. Already the sparse publicly available data suggest that the sources of UHECRs may be a strongly clustered sub-sample of galaxies or of active galactic nuclei. We present forecasts for various cases of source distributions which can be checked soon by the Pierre Auger Observatory.Comment: 11 pages, 8 figures, 4 tables; minor changes, matches published versio

Vibrating Winding Branes, Wrapping Democracy and Stabilization of Extra Dimensions in Dilaton Gravity

We show that, in the context of dilaton gravity, a recently proposed democratic principle for intersection possibilities of branes winding around extra dimensions yield stabilization, even with the inclusion of momentum modes of the wrapped branes on top of the winding modes. The constraints for stabilization massaged by string theory inputs forces the number of observed dimensions to be three. We also discuss consequences of adding ordinary matter living in the observed dimensions.Comment: Added a section discussing the linear and non-linear stability of the equilibrium point of the scale factors of the extra dimensions. Corrected a typo in the original field equations and other typos. Added and changed references. Final version appeared in JHE

Comparison of herbage yield, nutritive value and ensilability traits of three ryegrass species evaluated for the Irish Recommended List

peer-reviewedThis study examined 169 of the newest varieties of three ryegrass species, perennial (Lolium perenne L.), Italian (Lolium multiflorum Lam.) and hybrid (Lolium boucheanum Kunth), from Recommended List trials in Ireland. The traits examined were yield, dry matter concentration, three nutritive value traits (in vitro dry matter digestibility, water-soluble carbohydrate on a dry matter basis and crude protein concentration) and two ensilability traits (buffering capacity and water soluble carbohydrate concentration on an aqueous phase basis). Varietal monocultures of each species underwent a six cut combined simulated grazing and silage management in each of two years following sowing. Perennial ryegrass yielded less than both other species in one-year-old swards, but less than only Italian ryegrass in two-year-old swards, but generally had the higher in vitro dry matter digestibility and crude protein values. Italian ryegrass displayed the most favourable ensilability characteristics of the three species with perennial ryegrass less favourable and hybrid ryegrass intermediate. Overall, despite the high yields and favourable nutritive value and ensilability traits recorded, the general differences between the three ryegrass species studied were in line with industry expectations. These findings justify assessing the nutritive value and ensilability of ryegrass species, in addition to yield, to allow farmers select species that match farming enterprise requirements.We acknowledge the Department of Agriculture, Food and the Marine for funding provided through the Research Stimulus Fund (07 526

Linear Perturbations in Brane Gas Cosmology

We consider the effect of string inhomogeneities on the time dependent background of Brane Gas Cosmology. We derive the equations governing the linear perturbations of the dilaton-gravity background in the presence of string matter sources. We focus on long wavelength fluctuations and find that there are no instabilities. Thus, the predictions of Brane Gas Cosmology are robust against the introduction of linear perturbations. In particular, we find that the stabilization of the extra dimensions (moduli) remains valid in the presence of dilaton and string perturbations.Comment: 17 pages, 1 figur