Iron-rich solar particle events measured by SOHO/ERNE during two solar cycles

We study the differences in the heavy ion composition of solar energetic particle (SEP) events between solar cycles 23 and 24. We have surveyed the SOHO/ERNE heavy ion data from the beginning of solar cycle 23 until the end of June 2015, that is, well into the declining phase of cycle 24. We used this long observation period to study the properties of heavy ions (from C to Fe) and to compare the two solar cycles in this respect. We surveyed the data for SEP events with enhancements in the Fe/C and Fe/O intensity ratios in the energy range 5-15 MeV per nucleon, and associated the events with solar flare and coronal mass ejections (CME) when possible. We studied the properties of heavy ions in these events and compared the average relative abundances of heavy ions between the two solar cycles. We found that fewer days had C and O intensities higher than ~10$^{-3}$ cm$^{-2}$sr$^{-1}$s$^{-1}$(MeVn$^{-1}$)$^{-1}$ during solar cycle 24 than during cycle 23. For Fe this difference was clear even at lower intensities. We also found that fewer days had Fe/(C+O) > 0.183 during cycle 24. We identified 86 SEP events with at least one Fe-rich day, 65 of which occurred during cycle 23 and only 21 during cycle 24. We found that impulsive events have been almost completely absent during cycle 24. Mean abundances of heavy ions in the events were found to be significantly lower during cycle 24 than in cycle 23. Our results reflect the reduced solar activity in cycle 24 and indicate lower efficiency of particle acceleration processes for both gradual and impulsive SEP events in cycle 24.Comment: 14 pages, 9 figures and 3 tables. To be published in A&

Foundations of multiple black hole evolutions

We present techniques for long-term, stable, and accurate evolutions of multiple-black-hole spacetimes using the `moving puncture' approach with fourth- and eighth-order finite difference stencils. We use these techniques to explore configurations of three black holes in a hierarchical system consisting of a third black hole approaching a quasi-circular black-hole binary, and find that, depending on the size of the binary, the resulting encounter may lead to a prompt merger of all three black holes, production of a highly elliptical binary (with the third black hole remaining unbound), or disruption of the binary (leading to three free black holes). We also analyze the classical Burrau three-body problem using full numerical evolutions. In both cases, we find behaviors distinctly different from Newtonian predictions, which has important implications for N-body black-hole simulations. For our simulations we use analytic approximate data. We find that the eighth-order stencils significantly reduce the numerical errors for our choice of grid sizes, and that the approximate initial data produces the expected waveforms (after a rescaling of the puncture masses) for black-hole binaries with modest initial separations.Comment: Revtex 4, 13 pages, 15 figure

Multidimensionality and intra-individual variation in host manipulation by an acanthocephalan

Trophically-transmitted parasites frequently alter multiple aspects of their host's phenotype. Correlations between modified characteristics may suggest how different traits are mechanistically related, but these potential relationships remain unexplored. We recorded 5 traits from individual isopods infected with an acanthocephalan (Acanthocephalus lucii): hiding, activity, substrate colour preference, body (pereon) coloration, and abdominal (pleon) coloration. Infected isopods hid less and had darker abdominal coloration than uninfected isopods. However, in 3 different experiments measuring hiding behaviour (time-scales of observation: 1h, 8h, 8 weeks), these two modified traits were not correlated, suggesting they may arise via independent mechanisms. For the shorter experiments (1h and 8h), confidence in this null correlation was undermined by low experimental repeatability, i.e. individuals did not behave similarly in repeated trials of the experiment. However, in the 8-week experiment, hiding behaviour was relatively consistent within individuals, so the null correlation at this scale indicates, less equivocally, that hiding and coloration are unrelated. Furthermore, the difference between the hiding behaviour of infected and uninfected isopods varied over 8 weeks, suggesting that the effect of A. lucii infection on host behaviour changes over time. We emphasize the importance of carefully designed protocols for investigating multidimensionality in host manipulatio

Eye fluke-induced cataracts in natural fish populations: is there potential for host manipulation?

Manipulation of host phenotype (e.g. behaviour, appearance) is suggested to be a common strategy to enhance transmission in trophically transmitted parasites. However, in many systems, evidence of manipulation comes exclusively from laboratory studies and its occurrence in natural host populations is poorly understood. Here, we examined the potential for host manipulation by Diplostomum eye flukes indirectly by quantifying the physiological effects of parasites on fish. Earlier laboratory studies have shown that Diplostomum infection predisposes fish to predation by birds (definitive hosts of the parasites) by reducing fish vision through cataract formation. However, occurrence of cataracts and the subsequent potential for host manipulation in natural fish populations has remained poorly explored. We studied the occurrence of eye fluke-induced cataracts from 7 common fish species (Gymnocephalus cernuus, Rutilus rutilus, Leuciscus leuciscus, Alburnus alburnus, Osmerus eperlanus, Coregonus lavaretus and Gasterosteus aculeatus) from the Bothnian Bay in the Baltic Sea. We found that the parasite-induced cataracts were common in fish and they also reached high levels which are likely to predispose fish to predation. However, we observed such cataracts only in species with the highest parasite abundances, which suggests that only certain hosts may be strongly affected by the infectio

Acanthocephalan size and sex affect the modification of intermediate host colouration

For trophically transmitted parasites, transitional larval size is often related to fitness. Larger parasites may have higher establishment success and/or adult fecundity, but prolonged growth in the intermediate host increases the risk of failed transmission via natural host mortality. We investigated the relationship between the larval size of an acanthocephalan (Acanthocephalus lucii) and a trait presumably related to transmission, i.e. altered colouration in the isopod intermediate host. In natural collections, big isopods harboured larger worms and had more modified (darker) abdominal colouration than small hosts. Small isopods infected with a male parasite tended to have darker abdominal pigmentation than those infected with a female, but this difference was absent in larger hosts. Female size increases rapidly with host size, so females may have more to gain than males by remaining in and growing mutually with small hosts. In experimental infections, a large total parasite volume was associated with darker host respiratory operculae, especially when it was distributed among fewer worms. Our results suggest that host pigment alteration increases with parasite size, albeit differently for male and female worms. This may be an adaptive strategy if, as parasites grow, the potential for additional growth decreases and the likelihood of host mortality increase

Lasten ajankäyttö ja vähintään kohtuukuormitteinen fyysinen aktiivisuus varhaiskasvatuksessa

Background: The role of early childhood education and care (ECEC) is crucial in the implementation of the physical activity recommendations during long ECEC days. Young children should engage in 180 minutes of physical activity spread throughout the day, including 60 minutes of moderate-to-vigorous physical activity (MVPA). To attain the goal more information is needed on children’s time use and PA in children’s activities. Aim: The purposes of this quantitative observational study were twofold: (a) to examine children’s time use and (b) to describe which children’s activities best generate physical activity, specifically MVPA, in ECEC. Setting: Altogether, 2,879 Finnish children, aged 1- to 6-years, participated in this study. Methods: The data of 57,881 observations were analysed using cross-tabulation and a z-test. Results: Daily routines (e.g. eating, dressing, napping and transition), task or seatwork and material play accounted for almost 70% of children’s sedentary behaviour and less than 10% of MVPA. The best MVPA generators were physical activities (that generated 57.4% of all MVPA in boys vs. 62.8% in girls, respectively), rule play (11% in boys vs. 6.7% in girls), and role play (10.1% in boys vs. 9.9% in girls), as well as action not allowed (3.3% in boys vs 1.2% in girls). Conclusions: Educators should integrate physical activities, rule play and role play more in ECEC day to support the realization of the recommendations.Peer reviewe

Close encounters of three black holes

We present the first fully relativistic longterm numerical evolutions of three equal-mass black holes in a system consisting of a third black hole in a close orbit about a black-hole binary. We find that these close-three-black-hole systems have very different merger dynamics from black-hole binaries. In particular, we see complex trajectories, a redistribution of energy that can impart substantial kicks to one of the holes, distinctive waveforms, and suppression of the emitted gravitational radiation. We evolve two such configurations and find very different behaviors. In one configuration the binary is quickly disrupted and the individual holes follow complicated trajectories and merge with the third hole in rapid succession, while in the other, the binary completes a half-orbit before the initial merger of one of the members with the third black hole, and the resulting two-black-hole system forms a highly elliptical, well separated binary that shows no significant inspiral for (at least) the first t~1000M of evolution.Comment: 4 pages, 5 figure

Liikunnan lukutaidon tukeminen : Tapaustutkimus luokanopettajaopiskelijoiden käsityksistä

Physical literacy precedes and influences the physically active lifestyle, which is a key element of physical education. Especially, primary school physical education has an important task in the formation of the physically active lifestyle because it reaches all age cohorts. Classroom teachers are mainly in charge of implementing the curriculum in the primary schools. Thus, it is important to know more about their perceptions on physical literacy. Our research question was: How do classroom teacher students perceive the ways of supporting physical literacy? The data of this case study consists of four semi-structured interviews, which were analysed using content analysis. The concept of physical literacy, developed by Whitehead (1993), was used to categorize the data. Furthermore, the answers in the categories were combined in themes that the result of responses to the research question. As a key result of this study, it was found that classroom student teachers perceive the ways of supporting physical literacy as follows: 1) Motivation can be supported by atmosphere, teaching methods, contents and learning environments; (2) Physical competence and self-confidence is supported by experiences of success, developmentally appropriate physical activities and motor skills and 3) Knowledge and understanding can be supported by creating meanings and adjusted objectives, evaluation and individualization. Our results suggest that future classroom teachers have a quite broad understanding of supporting physical literacy, which is in line with their broad educational duties that differ from those of subject teachers. The results of our research are especially useful for classroom teachers. An approach that emphasizes personal potential can be one solution to challenges of immobility. In the future, it is important to study what kind of pedagogical practices based on physical literacy could lead to the development of teaching in physical education in primary schools.Physical literacy precedes and influences the physically active lifestyle, which is a key element of physical education. Especially, primary school physical education has an important task in the formation of the physically active lifestyle because it reaches all age cohorts. Classroom teachers are mainly in charge of implementing the curriculum in the primary schools. Thus, it is important to know more about their perceptions on physical literacy. Our research question was: How do classroom teacher students perceive the ways of supporting physical literacy? The data of this case study consists of four semi-structured interviews, which were analysed using content analysis. The concept of physical literacy, developed by Whitehead (1993), was used to categorize the data. Furthermore, the answers in the categories were combined in themes that the result of responses to the research question. As a key result of this study, it was found that classroom student teachers perceive the ways of supporting physical literacy as follows: 1) Motivation can be supported by atmosphere, teaching methods, contents and learning environments; (2) Physical competence and self-confidence is supported by experiences of success, developmentally appropriate physical activities and motor skills and 3) Knowledge and understanding can be supported by creating meanings and adjusted objectives, evaluation and individualization. Our results suggest that future classroom teachers have a quite broad understanding of supporting physical literacy, which is in line with their broad educational duties that differ from those of subject teachers. The results of our research are especially useful for classroom teachers. An approach that emphasizes personal potential can be one solution to challenges of immobility. In the future, it is important to study what kind of pedagogical practices based on physical literacy could lead to the development of teaching in physical education in primary schools.Peer reviewe

Seasonal changes in host phenotype manipulation by an acanthocephalan: time to be transmitted?

Many complex life cycle parasites exhibit seasonal transmission between hosts. Expression of parasite traits related to transmission, such as the manipulation of host phenotype, may peak in seasons when transmission is optimal. The acanthocephalan Acanthocephalus lucii is primarily transmitted to its fish definitive host in spring. We assessed whether the parasitic alteration of 2 traits (hiding behaviour and coloration) in the isopod intermediate host was more pronounced at this time of year. Refuge use by infected isopods was lower, relative to uninfected isopods, in spring than in summer or fall. Infected isopods had darker abdomens than uninfected isopods, but this difference did not vary between seasons. The level of host alteration was unaffected by exposing isopods to different light and temperature regimes. In a group of infected isopods kept at 4°C, refuge use decreased from November to May, indicating that reduced hiding in spring develops during winter. Keeping isopods at 16°C instead of 4°C resulted in higher mortality but not accelerated changes in host behaviour. Our results suggest that changes in host and/or parasite age, not environmental conditions, underlie the seasonal alteration of host behaviour, but further work is necessary to determine if this is an adaptive parasite strategy to be transmitted in a particular seaso