Mammoth fauna remains from late Pleistocene deposits of the Dmitry Laptev Strait south coast (Northern Yakutia, Russia)

The Yedoma Ice Complex in northern Yakutia provides perfect preservation conditions for frozen remains of vertebrate animals. Even complete mummified specimens of the late Pleistocene Beringian Mammoth fauna such as woolly mammoth, woolly rhinoceros, horse, and bison are occasionally found in permafrost deposits across eastern Siberia, i.e., in West Beringia, although bones are much more commonly found. The present study characterizes mammal bones from late Pleistocene and Holocene permafrost deposits exposed on the Oyogos Yar coast, part of the southern shore of the Dmitry Laptev Strait that connects the Laptev and East Siberian seas. The study applies a method to characterize fossil bone samples by the location of their discovery and by the accuracy of their relation to a depositional horizon. We analyzed a total of 38 finite radiocarbon ages of bone material from mammoth, horse, and musk ox, spanning from about 48.8 to 4.5 ka BP and including both our own data and data from the literature, in addition to previous publications that reported numerous bones with infinite ages from the Oyogos Yar coast. The distribution of bones and tooth along the coastal permafrost exposure is not uniform; it depends upon whether the material was found in situ, on thermo-terraces, or on the shore. The overall bone collection consists of 13 species of which Mammuthus primigenius (woolly mammoth, 41%), Bison priscus (bison, 19%), Equus ex gr., caballus (horse, 19%), and Rangifer tarandus (reindeer, 16%) predominate. The fossil bone species distribution is similar to those of other prominent Yedoma outcrops in the region, i.e., on Bykovsky Peninsula and on Bol’shoy Lyakhovsky Island. Correlation analysis shows that the Oyogos Yar bone sampling sites of different geomorphological settings are similar to each other but not to all sampling sites within the other two locations on Bykovsky Peninsula and on Bol’shoy Lykahovsky Island. High similarities in terms of correlation coefficients between specific sampling sites are often not represented in the cluster analysis

Paleozoological characteristics of the Late Neo-Pleistocene - Holocene sediments of Bykovsky Peninsula, Northern Yakutia (Палеозоологическая характеристика поздненеоплейстоценовых – голоценовых отложений Быковского Полуострова (Северная Якутия)

The present study characterizes palaeozoological findings from the late Pleistocene and Holocene permafrost deposits of the Bykovsky Peninsula. The study proposes a method to differentiate fossil bone samples by the location of their finding and by the accuracy of their relation to a certain depositional horizon. A total of 90 radiocarbon dates of bone material from large mammals has been analyzed. The distribution of bone material along the sedimentary permafrost profile is not uniform. During the last 60.000 years, two periods of environmental conditions favourable for representatives of mammoth fauna are distinguished. “Cold” tundra-steppe environments with either dry and warm or cold summer conditions provided the most favourable settings for the existence of mammoths and hoofed mammals over the territory of the modern Bykovsky Peninsula

Organic matter composition and greenhouse gas production of thawing subsea permafrost in the Laptev Sea

Subsea permafrost represents a large carbon pool that might be or become a significant greenhouse gas source. Scarcity of observational data causes large uncertainties. We here use five 21-56 m long subsea permafrost cores from the Laptev Sea to constrain organic carbon (OC) storage and sources, degradation state and potential greenhouse gas production upon thaw. Grain sizes, optically-stimulated luminescence and biomarkers suggest deposition of aeolian silt and fluvial sand over 160 000 years, with dominant fluvial/alluvial deposition of forest- and tundra-derived organic matter. We estimate an annual thaw rate of 1.3 ± 0.6 kg OC m−2 in subsea permafrost in the area, nine-fold exceeding organic carbon thaw rates for terrestrial permafrost. During 20-month incubations, CH4 and CO2 production averaged 1.7 nmol and 2.4 µmol g−1 OC d−1, providing a baseline to assess the contribution of subsea permafrost to the high CH4 fluxes and strong ocean acidification observed in the region

Ice Complex chronologies and environments in western Beringia

Polygon tundra with tundra-steppe vegetation cover and growing syngenetic ice-wedge nets evolved during stadial and interstadial periods of the late Quaternary in non-glaciated Beringia. The depositional relict of such environments is called Ice Complex (IC; ледовый комплекс [ledovyi kompleks] in Russian) permafrost. The IC archives preserve information of past periglacial and climate landscape conditions of mid- and late Pleistocene Beringian environments. In certain locations of the East Siberian Arctic, IC remnants of different age and extent are known. While using IC deposits as archives of palaeo-landscape and palaeo-environmental dynamics, summer and winter conditions over large time-scales are detectable. Commonly applied summer proxy include palaeontological proxy such as pollen, plant macrofossils, insect fossils and, most prominent, mammal fossils of the Mammoth fauna, while geochemical and stable isotope properties of ground ice allow for insights into freezing and winter conditions. IC chronologies are challenging because the deposition and post-sedimentary preservation of ice-rich permafrost are triggered by palaeo-relief settings and related processes as well as by the intensity of thermokarst. This complicates geochronological interpretations, as representatives of consecutive late Quaternary periods may be found at laterally different positions and altitudes in coastal and riverine exposures. Shifts between permafrost aggradation and degradation over time frequently cause gaps in sequences. Furthermore, numerical dating of IC mainly includes different approaches such as radiocarbon (14C) dating of organic material, infrared and optically-stimulated luminescence (IRSL, OSL) dating on feldspar and quartz grains, radioisotope disequilibria of thorium-230 to uranium-234 (230Th/U) dating of peat, and chlorine-36 to chloride ratios (36Cl/Cl) of ground ice. The application of various geochronologic methods to cover the age intervals of certain IC deposits implies that different permafrost components (organic, mineralic, ice) as well as different geochemical and physical properties have to be employed. At the southern coast of Bol'shoy Lyakhovsky Island at least four distinct IC strata were previously described and dated, which cover among the longest time interval of late Quaternary terrestrial permafrost deposition in East Siberia; starting about 200 kyr ago. With this contribution we seek to present and discuss our current understanding of IC chronologies preserved on the New Siberian Archipelago including MIS2 Yedoma (Sartan) IC, MIS3 Yedoma (Molotkov) IC, MIS5 Buchchagy IC, and MIS7a Yukagir IC. Geocryological and palaeo-environmental proxy data highlight past periglacial landscape and deposition processes to deduce past climate conditions and Beringian palaeo-ecological settings and dynamics

25 years of joint Yedoma Ice Complex studies in Arctic Russia, especially in Sakha/Yakutia

Since 1994, permafrost deposits of the Siberian Yedoma region have been in the focus of the joint Russian-German scientific cooperation in terrestrial Polar research (Figure 1). These studies focused on cryostratigraphic, geochemical, geochronological, and paleontological characteristics at more than 25 individual study sites of the late Pleistocene Yedoma Ice Complex in Siberia and provided a detailed insight into the paleoenvironments and paleoclimate for the westernmost part of Beringia. The multidisciplinary investigations resulted in new ideas and discussions in the ongoing scientific debate on the origin of Yedoma Ice Complex and the main periglacial processes involved in its formation (1,2,3). The Yedoma Ice Complex is an ice-rich type of permafrost deposit widely distributed across Beringia. The Ice Complex aggradation is mainly controlled by the growth of syngenetic ice wedge polygons contributing up to 60 vol% of the entire formation. The clastic sedimentation of ice-oversaturated Yedoma deposits with considerable organic matter content is further controlled by local conditions such as source rocks and periglacial weathering processes, paleotopography, and temporary surface stabilization with autochthonous peat growth and soil formation. Key processes include alluvial, fluvial, and niveo-aeolian transport (4) as well as accumulation in ponding waters and continued in-situ frost weathering over millennial time-scales. Important post-depositional processes affecting Yedoma deposits are solifluction, cryoturbation, and pedogenesis. Major joint Russian-German field studies were conducted on Taymyr Peninsula (5,6,7,8,9,10,11), along the western and central Laptev Sea coasts (12,13,14,15,16,17,18), in the Lena Delta (19,20,21,22), on islands of the New Siberian Archipelago (23,24,25,26,27,28), and the adjacent mainland (29). Further study sites were conducted in the Kolyma Lowland (30), the Yana Highlands (31,32), in the foothills of the Verkhoyan Mountains (33,34,35,36), and in Central Yakutia (37). Comprehensive sampling and further analytical work included not only the Yedoma Ice Complex itself but also included its stratigraphic context of older underlying sequences and younger overlying deposits. The latter often are subaerial or subaquatic deposits associated with late-Glacial to Holocene thermokarst dynamics that led to Yedoma degradation during the deglacial and Holocene warming of these regions (38,39,40). Figure 1: Joint Russian-German fieldwork sites in NE Siberia labeled with the year of expedition. Besides geomorphological and cryolithological studies, extensive paleo-ecological investigations were carried out on zoological (41,42,43,44,45) and botanic fossils (46,47,48,49,50,51) to derive quantitative and qualitative reconstructions late Pleistocene Beringian environments and climate conditions. New methods in geochronology were also tested (52,53,54,55). In addition to the sedimentary components of the frozen deposits, segregated ground ice and in particular the large syngenetic ice wedges of Yedoma Ice Complex were also studied as geochemical and stable isotope archives of paleoclimate (56,57,58, 59,60,61,62). In addition, a range of remote sensing methods in combination with GIS analyses (63,64,65) and geophysical surveys (66) were used for large-scale analyses of landscape changes associated with Yedoma Ice Complex degradation (67,68,69). In the last few years, an additional important focus has been on using modern biogeochemical methods of organic matter analysis to characterize the frozen organic matter in Yedoma Ice Complex deposits and for permafrost carbon pool calculations (70, 71,72,73,74,75,76,77) as well as microbiological studies (78) and genetic studies on fossil DNA (79,80). The rich body of scientific data and literature produced in Russian-German co-authorship within the more than 25 years of joint research on Yedoma Ice Complex represents an important cornerstone for understanding the Late Quaternary evolution of Siberian Yedoma regions, its role in the Earth System, and its feedbacks with climate and ecosystems. References 1. Schirrmeister, L., Dietze, E., Matthes, H., Grosse, G., Strauss, J., Laboor, S., Ulrich, M., Kienast, F., and Wetterich, S. (2020) The genesis of Yedoma Ice Complex permafrost – grain-size endmember modeling analysis from Siberia and Alaska, E&G Quaternary Sci. J., 69, 33–53, doi: 10.5194/egqsj-69-33-2020. 2. Schirrmeister, L., Froese, D., Tumskoy, V., Grosse,G., Wetterich, S. (2013.) Yedoma: Late Pleistocene ice-rich syngenetic permafrost of Beringia. In: Elias S.A. (ed.) The Encyclopedia of Quaternary Science 2nd edition, vol. 3, pp. 542-552. Amsterdam: Elsevier. 3. Schirrmeister, L., Kunitsky, V.V., Grosse, G., Wetterich, S., Meyer, H., Schwamborn, G., Babiy, O., Derevyagin, A.Y., and Siegert, C.: Sedimentary characteristics and origin of the Late Pleistocene Ice Complex on North-East Siberian Arctic coastal lowlands and islands - a review. Quaternary International 241, 3-25, doi: 10.1016/j.quaint.2010.04.004, 2011. 4. Kunitsky, V., Schirrmeister, L., Grosse, G., Kienast, F. (2002). Snow patches in nival landscapes and their role for the Ice Complex formation in the Laptev Sea coastal lowlands, Polarforschung, 70, 53-67, doi:10.2312/polarforschung.70.53. 5. Andreev, A. , Siegert, C. , Klimanov, V. A. , Derevyagin, A. Y. , Shilova, G. N. and Melles, M. (2002) Late Pleistocene and Holocene vegetation and climate changes in the Taymyr lowland, Northern Siberia Quaternary research, 57, pp. 138-150 . 6. Andreev, A. , Tarasov, P. E. , Siegert, C. , Ebel, T. , Klimanov, V. A. , Melles, M. , Bobrov, A. A. , Derevyagin, A. Y. , Lubinski, D. J. and Hubberten, H. W. (2003) Vegetation and climate changes on the northern Taymyr, Russia during the Upper Pleistocene and Holocene reconstructed from pollen records , Boreas, 32 (3), pp. 484-505 . 7. Chizhov, A. B. , Derevyagin, A. Y. , Simonov, E. F. , Hubberten, H. W. and Siegert, C. (1997) Isotopic composition of ground ice at the Labaz Lake region (Taymyr). Kriosfera Zemlii (Earth Cryoshere), 1, No 3, pp. 79-84 . (in Russian), 8. Derevyagin, A.Yu., Chizhov, A.B., Brezgunov, V.S., Siegert, C., Hubberten, H.-W., 1999.Isotopic composition of ice wedges of Cape Sabler (Lake Taymyr). Kriosfera Zemlii (Earth Cryosphere) 3/3, 41-49 (in Russian). 9. Kienast, F., Siegert, C., Dereviagin, A., Mai, H.D. Climatic implications of Late Quaternary plant macrofossil assemblages from the Taymyr Peninsula, Siberia, Global and Planetary Change, Volume 31, Issues 1–4, 265-281, 2001, https://doi.org/10.1016/S0921-8181(01)00124-2. 10. Kienel, U. , Siegert, C. and Hahne, J. (1999) Late Quarternary paeloenvironmental reconstruction from a permafrost sequence (Northsiberian Lowland, SE Taymyr Peninsula) - a multidisciplinary case study, Boreas, 28 (1), pp. 181-193 . 11. Siegert C., Derevyagin A.Y., Shilova G.N., Hermichen WD., Hiller A. (1999) Paleoclimatic Indicators from Permafrost Sequences in the Eastern Taymyr Lowland. In: Kassens H. et al. (eds) Land-Ocean Systems in the Siberian Arctic. Springer, Berlin, Heidelberg. 12. Bobrov, A.A., Müller, S., Chizhikova, N.A., Schirrmeister, L., Andreev, A.A.(2009).Testate Amoebae in Late Quaternary Sediments of the Cape Mamontov Klyk (Yakutia), Biology Bulletin, 36(4), 363-372. 13. Schirrmeister, L., Grosse, G., Kunitsky, V., Magens, D., Meyer, H., Dereviagin, A., Kuznetsova, T., Andreev, A., Babiy, O., Kienast, F., Grigoriev, M., Overduin, P.P., and Preusser, F.: Periglacial landscape evolution and environmental changes of Arctic lowland areas for the last 60,000 years (Western Laptev Sea coast, Cape Mamontov Klyk), Polar Research, 27(2), 249-272, doi: 10.1111/j.1751-8369.2008.00067.x, 2008. 14. Winterfeld, M., Schirrmeister, L., Grigoriev, M., Kunitsky, V.V., Andreev, A., and Overduin, P.P.: Permafrost and Landscape Dynamics during the Late Pleistocene, Western Laptev Sea Shelf, Siberia, Boreas 40(4), 697–713, doi: 10.1111/j.1502-3885.2011.00203.x, 2011. 15. Siegert, C., Schirrmeister, L., and Babiy, O.: The sedimentological, mineralogical and geochemical composition of late Pleistocene deposits from the ice complex on the Bykovsky peninsula, northern Siberia, Polarforschung, 70, 2000, 3-11, doi: 10.2312/polarforschung.70.3, 2002. 16. Schirrmeister, L., Siegert, C., Kuznetsova, T., Kuzmina, S., Andreev, A.A., Kienast, F., Meyer, H., and Bobrov, A.A.: Paleoenvironmental and paleoclimatic records from permafrost deposits in the Arctic region of Northern Siberia, Quaternary International, 89, 97-118, doi: 10.1016/S1040-6182(01)00083-0, 2002. 17. Schirrmeister, L., Siegert, C., Kunitzky, V.V., Grootes, P.M., and Erlenkeuser, H.: Late Quaternary ice-rich permafrost sequences as a paleoenvironmental archive for the Laptev Sea Region in northern Siberia, International Journal of Earth Sciences, 91, 154-167, doi: 10.1007/s005310100205, 2002. 18. Schirrmeister, L., Schwamborn, G., Overduin, P.P., Strauss, J., Fuchs, M.C., Grigoriev, M., Yakshina, I., Rethemeyer, J., Dietze, E., and Wetterich, S.: Yedoma Ice Complex of the Buor Khaya Peninsula (southern Laptev Sea), Biogeosciences 14, 1261-1283, doi: 10.5194/bg-14-1261-2017, 2017. 19. Schirrmeister, L., Kunitsky, V.V., Grosse, G., Schwamborn, G., Andreev, A.A., Meyer, H., Kuznetsova, T., Bobrov, A., and Oezen, D.: Late Quaternary history of the accumulation plain north of the Chekanovsky Ridge (Lena Delta, Russia) - a multidisciplinary approach, Polar Geography, 27(4), 277-319, doi: 10.1080/789610225, 2003. 20. Schirrmeister, L., Grosse, G. Schnelle, M., Fuchs, M., Krbetschek, M., Ulrich, M., Kunitsky, V., Grigoriev, M., Andreev, A. Kienast, F., Meyer, H., Klimova, I., Babiy, O., Bobrov, A., Wetterich, S., and Schwamborn, G.: Late Quaternary paleoenvironmental records from the western Lena Delta, Arctic Siberia, Palaeogeography, Palaeoclimatology, Palaeoecology 299, 175–196, doi: 10.1016/j.quascirev.2009.11.017, 2011. 21. Schwamborn, G., Rachold, V., and Grigoriev, M.N.: Late Quaternary sedimentation history of the Lena Delta, Quaternary International 89, 119–134, doi: 10.1016/S1040-6182(01)00084-2, 2002. 22. Wetterich, S., Kuzmina, S., Andreev, A.A., Kienast, F., Meyer, H., Schirrmeister, L., Kuznetsova, T., and Sierralta, M.: Palaeoenvironmental dynamics inferred from late Quaternary permafrost deposits on Kurungnakh Island, Lena Delta, Northeast Siberia, Russia, Quaternary Science Reviews, 27, 1523-1540, doi: 10.1016/j.quascirev.2008.04.007, 2008. 23. Andreev, A.A., Grosse, G., Schirrmeister, L., Kuzmina, S.A., Novenko, E.Yu., Bobrov, A.A., Tarasov, P. E., Kuznetsova, T.V., Krbetschek, M., Meyer, H., and Kunitsky, V.V.: Late Saalian and Eemian palaeoenvironmental history of the Bol'shoy Lyakhovsky Island (Laptev Sea region, Arctic Siberia), Boreas 33(4), 319-348, doi:10.1080/03009480410001974, 2004. 24. Andreev, A., Grosse, G., Schirrmeister, L., Kuznetsova, T.V., Kuzmina, S.A., Bobrov, A.A., Tarasov, P.E., Novenko, E.Yu., Meyer, H., Derevyagin, A.Yu., Kienast, F., Bryantseva, A., and Kunitsky, V.V.: Weichselian and Holocene palaeoenvironmental history of the Bol’shoy Lyakhovsky Island, New Siberian Archipelago, Arctic Siberia, Boreas 38(1), 72–110, doi: 10.1111/j.1502-3885.2008.00039.x, 2009. 25. Wetterich, S., Rudaya, N., Meyer, H., Opel, T., and Schirrmeister, L.: Last Glacial Maximum records in permafrost of the East Siberian Arctic, Quaternary Science Reviews 30, 3139-3151, doi: 10.1016/j.quascirev.2011.07.020, 2011. 26. Wetterich, S., Rudaya, N., Andreev, A.A., Opel, T., Schirrmeister, L., Meyer, H., and Tumskoy, V.: Ice Complex formation in arctic East Siberia during the MIS3 Interstadial, Quaternary Science Reviews 84: 39-55, doi:. 10.1016/j.quascirev.2013.11.009, 2014. 27. Wetterich, S.; Tumskoy:V.E., Rudaya, N., Kuznetsov, V., Maksimov, F., Opel T. , Meyer H., Andreev, A.A., Schirrmeister, L (2016) Ice Complex permafrost of MIS5 age in the Dmitry Laptev Strait coastal region (East Siberian Arctic). Quaternary Science Reviews, 147:298-31, doi.org/10.1016/j.quascirev.2015.11.016. 28. Wetterich, S., Rudaya, N., Kuznetsov V., Maksimov, F., T. Opel, Meyer, H., Guenther, F., Bobrov, A., Raschke, E., Zimmermann, H., Strauss, J., Fuchs, M.C., Schirrmeister, L. (2019) Recurrent Ice Complex formation in arctic eastern Siberia since about 200 ka. Quaternary Research 92 (2); 530-548, doi.org/10.1017/qua.2019.6. 29. Wetterich, S., Schirrmeister, L., Andreev A. A., Pudenz, M., Plessen, B, Meyer, H., Kunitsky, V. V. (2009). Eemian and Late Glacial/Holocene palaeoenvironmental records from permafrost sequences at the Dmitry Laptev Strait (NE Siberia, Russia), Palaeogeography, Palaeoclimatology, Palaeoecology 279: 73-95 doi:10.1016/j.palaeo.2009.05.002. 30. Strauss, J., Schirrmeister, L., Wetterich, S., Borchers, A, and Davydov S.P.: Grain-size properties and organic-carbon stock of Yedoma Ice Complex permafrost from the Kolyma lowland, northeastern Siberia. GBC. 26: GB3003, doi: 10.1029/2011GB004104, 2012. 31. Ashastina, K., Schirrmeister, L., Fuchs M., and Kienast F.: Palaeoclimate characteristics in interior Siberia of MIS 6–2: first insights from the Batagay permafrost mega-thaw slump in the Yana Highlands, Clim. Past, 13, 795–818, doi: 10.5194/cp-13-795-2017, 2017. 32. Kunitsky, V.V., Syromyatnikov, I.I., Schirrmeister, L., Skachkov, Yu.B., Grosse, G., Wetterich, S., and Grigoriev, M.N.: Ice-rich permafrost and thermal denudation in the Kirgillyakh area, Kriosfera Zemli. 17(1), 56-68, 2013 (in Russian). 33. Popp, S., Diekmann,B., Meyer, H., Siegert, C.,Syromyatnikov, I., Hubberten, H.-W. Palaeoclimate Signals as Inferred from Stable-isotope Composition of Ground Ice in the Verkhoyansk Foreland, Central Yakutia. Permafrost and Periglac. Process. 17: 119–132 (2006) DOI: 10.1002/ppp.556 34. Popp, S., Belolyubsky, I., Lehmkuhl, F., Prokopiev, A., Siegert, C., Spektor, V., Stauch, G., Diekmann,B. Sediment provenance of late Quaternary morainic, fluvialand loess-like deposits in the southwestern VerkhoyanskMountains (eastern Siberia) and implications for regionalpalaeoenvironmental reconstructions. Geol. J.42: 477–497 (2007), DOI: 10.1002/gj.1088 35. Siegert, C. , Sergeyenko, A. I. and Schirrmeister, L. (2017) Late Quaternary Deposits of the Northern Verkhoyansk Mountains: Geochronology and Questions of their Genesis (in Russian), Bulletin of the Commission for Study of the Quaternary = БЮЛЛЕТЕНЬ КОМИССИИ ПО ИЗУЧЕНИЮ ЧЕТВЕРТИЧНОГО ПЕРИОДА, 75 , pp. 100-112 . 36. Siegert, C. , Stauch, G. , Lehmkuhl, F. , Sergeyenko, A. I. , Diekmann, B. , Popp, S. and Belolyubsky, I. N. (2007) Development of glaciation in the Verkhoyansk Range and its foreland during the Pleistocene: Results of new investigations., Regionalnaya Geologiya i Metallogeniya (Regional Geology and Metallogeny), No. 30-31(in Russian)., 222 . 37. Ulrich, M., Morgenstern, A., Günther, F., Reiss, D. Bauch, K. E., Hauber, E., Rössler, S. and Schirrmeister, L. (2010) Thermokarst in Siberian ice-rich permafrost: Comparison to asymmetric scalloped depressions on Mars, Journal of Geophysical Research, 115, E10009 . doi:10.1029/2010JE003640 , 38. Morgenstern, A. , Grosse, G. , Günther, F. , Fedorova, I. and Schirrmeister, L. (2011): Spatial analyses of thermokarst lakes and basins in Yedoma landscapes of the Lena Delta. The Cryosphere, 5(4), 849–867, doi:10.5194/tc-5-849-2011. 39. Morgenstern, A. , Ulrich, M. , Günther, F. , Roessler, S. , Fedorova, I. V. , Rudaya, N. A. , Wetterich, S. , Boike, J. and Schirrmeister, L. (2013). Evolution of thermokarst in East Siberian ice-rich permafrost: A case study, Geomorphology, 201 , 363-379. doi:10.1016/j.geomorph.2013.07.011 40. Biskaborn, B. , Herzschuh, U. , Bolshiyanov, D. Y. , Schwamborn, G. and Diekmann, B. (2013) Thermokarst Processes and Depositional Events in a Tundra Lake, Northeastern Siberia, Permafrost and Periglac. Process.24: 160–174 doi:https://doi.org/10.1002/ppp.1769, 41. Kuznetsova, T. V. , Sulerzhitsky, L. D. , Andreev, A. , Siegert, C. , Schirrmeister, L. and Hubberten, H. W. (2003) Influence of Late Quaternary paleoenvironmental conditions on the distribution of mammals fauna in the Laptev Sea region , Occasional Papers in Earth Sciences, 5 , pp. 58-60 . 42. Kuznetsova T.V., Tumskoy V.E., Schirrmeister L., Wetterich S., (2019.) Paleozoological characteristics of the Late Neo-Pleistocene - Holocene sediments of Bykovsky Peninsula, Northern Yakutia (Палеозоологическая характеристика поздненеоплейстоценовых – голоценовых отложений Быковского Полуострова (Северная Якутия). Zoological Journal 98(11), 1268-1290. Special issue in honor of Andrey Sher. (in Russian) doi: 10.1134/S0044513419110102. 43. Bobrov, A. A. , Andreev, A. , Schirrmeister, L. and Siegert, C. (2004) Testate amoebae (Protozoa: Testacea) as bioindicators in the Late Quaternary deposits of the Bykovsky Peninsula, Laptev Sea, Russia , Palaeogeography palaeoclimatology palaeoecology, 209 , pp. 165-181 . doi:https://doi.org/10.1016/J.PALAEO.2004.02.012 44. Wetterich, S., Schirrmeister, L., Pietrzeniuk, E. (2005). Freshwater ostracodes in Quaternary permafrost deposits from the Siberian Arctic, Journal of Paleolimnology, 34, 363-376. doi:10.1007/s10933-005-5801-y 45. Müller, S., Bobrov, A. A., Schirrmeister, L., Andreev, A. A., Tarasov, P. E. (2009). Testate amoebae record from the Laptev Sea coast and its implication for the reconstruction of Late Pleistocene and Holocene environments in the Arctic Siberia, Palaeogeography, Palaeoclimatology, Palaeoecology 271(3-4), 301-315. doi:10.1016/j.palaeo.2008.11.003 46. Andreev, A.A., Schirrmeister, L., Siegert, C., Bobrov, A.A., Demske, D., Seiffert, M., Hubberten, H.-W. (2002). Paleoenvironmental changes in Northeastern Siberia during the Late Quaternary - evidence from pollen records of the Bykovsky Peninsula, Polarforschung, 70, 13-25, doi:10.2312/polarforschung.70.13. 47. Andreev, A.A.; Schirrmeister, L.; Tarasov , P.E.; Ganopolski , A.; Brovkin V.; Siegert, C.; Hubberten, H.-W. (2011). Vegetation and climate history in the Laptev Sea region (arctic Siberia) during Late Quaternary inferred from pollen records. Journal of Quaternary science reviews. 30, 2182-2199 doi:10.1016/j.quascirev.2010.12.026. 48. Kienast, F. , Schirrmeister, L. , Siegert, C. and Tarasov, P. (2005) Palaeobotanical evidence for warm summers in the East Siberian Arctic during the last cold stage, Quaternary Research, 63 (3), pp. 283-300. doi:https://doi.org/10.1016/j.yqres.2005.01.003 , 49. Kienast, F., Tarasov, P., Schirrmeister, L., Grosse, G., Andreev, A.A. (2008). Continental climate in the East Siberian Arctic during the last interglacial: implications from palaeobotanical records, Global and Planetary Change, 60(3/4), 535-562. doi:10.1016/j.gloplacha.2007.07.004 50. Kienast, F., Wetterich, S., Kuzmina, S., Schirrmeister, L., Andrev, A., Tarasov, P., Nazarova, L., Kossler, A., Frolova, A., Kunitsky, V. V.(2011) Paleontological records indicate the occurrence of open woodlands in a dry inland climate at the present-day Arctic coast in western Beringia during the last interglacial. Quaternary Science Reviews 30: 2134-2159, doi:10.1016/j.quascirev.2010.11.024. 51. Palagushkina, O.V., Wetterich, S., Schirrmeister, L., Nazarova, L.B. (2017) Modern and fossil diatom assemblages from Bol'shoy Lyakhovsky Island (New Siberian Archipelago, Arctic Siberia). Contemporary Problems of Ecology, 10, (4), 380–394. doi: 10.1134/S1995425517040060. 52. Gilichinsky, D. A. , Nolte, E., Basilyan, A.E., Beer, J., Blinov, A., Lazarev, V., Kholodov, A., Meyer, H., Nikolsky, P.A., Schirrmeister, L., Tumskoy, V. (2007). Dating of syngenetic ice wedges in permafrost with 36Cl and 10Be, Quaternary science reviews. 26, 1547-1556. doi:10.1016/j.quascirev.2007.04.004 53. Blinov A.V., Beer J., Tikhomirov D.A., Schirrmeister L., Meyer H., Abramov A.A., Basylyan A.E., Nikolskiy P.A., Tumskoy V.E., Kholodov A.L., Gilichinsky D.A. (2009) Permafrost dating with the cosmogenic radionuclides ( Report 1) (= Датирование многолетнемерзлых пород с помощью космогенных радионуклидов (сообщение 1). Kriosfera Zemli 13,( 2), 3-15 (in Russian). 54. Blinov, A., Alfimov, V., Beer, J., Gilichinsky, D., Schirrmeister, L., Kholodov, A., Nikolskiy, P., Opel, T., Tikhomirov, D., Wetterich, S.(2009).36Cl/Cl ratio in ground ice of East Siberia and its application for chronometry, Geochemistry, Geophysics, Geosystems (G3). 10(1), doi: 10.1029/2009GC002548. 55. Schirrmeister, L., Oezen, D., Geyh, M.A. (2002). 230Th/U dating of frozen peat, Bol'shoy Lyakhovsky Island (North Siberia), Quaternary research, 57, 253-258. doi:10.1006/qres.2001.2306. 56. Meyer, H. , Derevyagin, A. Y. , Siegert, C. and Hubberten, H. W. (2002) Paleoclimate studies on Bykovsky Peninsula, North Siberia - hydrogen and oxygen isotopes in ground ice , Polarforschung 70:, pp. 37-51 . 57. Derevyagin, A. Y., Chizhov, A. , Meyer, H. , Opel, T. , Schirrmeister, L. and Wetterich, S. (2013). Isotopic composition of texture ices, Laptev Sea coast , Kriosfera Zemlii (Earth Cryosphere), XVII (3), pp. 27-34 (in Russian). 58. Meyer, H. , Derevyagin, A. Y. , Siegert, C. , Schirrmeister, L. and Hubberten, H. W. (2002) Paleoclimate reconstruction on Big Lyakhovsky Island, North Siberia - Hydrogen and oxygen isotopes in ice wedges , Permafrost and periglacial processes, 13 , pp. 91-105 . 59. Opel, T., Dereviagin, A., Meyer, H., Schirrmeister, L., Wetterich, S. (2010).Paleoclimatic information from stable water isotopes of Holocene ice wedges at the Dmitrii Laptev Strait (Northeast Siberia), Permafrost and Periglacial Processes. 22 (1), 84-100, doi:10.1002/ppp.667. 60. Opel, T., Wetterich, S., Meyer, H., Dereviagin, A.Yu., Fuchs, M.C., and Schirrmeister, L.: Ground-ice stable isotopes and cryostratigraph

Western Beringia and beyond - three decades of German-Russian paleoenvironmental research on Siberian permafrost

With first joint fieldwork on Taymyr Peninsula during mid-1990s, a successful cooperation of German, Russian, and further international partners on permafrost and Quaternary palaeoenvironments in Siberia was started and resulted in extensive joint research for 3 decades. Studies of permafrost deposits and ground ice provided insights on past environmental and climatic changes, covering several hundreds of thousands of years into the past. They provide multi-proxy evidence for multiple glacial/interglacial cycles and different periods of past climate change or stability in Arctic land environments. Study objects were natural permafrost exposures along coastal sections, thaw slumps, and river banks, studied mostly during summers, complemented by permafrost cores from land, lake and sea ground drilled mostly in spring. Exposure geometry and stratigraphic horizon thickness have been surveyed using laser tachymetry, other measuring equipment, and drones. Based on multi-proxy analyses, mid- and late Quaternary periods were studied, resulting in >300 scientific papers. The approach includes geomorphic studies, various geochronological analyses, analysis of frozen sediments (for ice, carbon, nitrogen, and carbonate contents, grain-size parameters, magnetic susceptibility, heavy mineral compositions), ground ice (stable water isotopes, major ions) and of numerous fossil bioindicators, to reconstruct the Quaternary paleoenvironmental change. Oldest permafrost horizons were dated from the Batagay mega-thaw-slump (Yana Uplands) to about 650 ky with luminescence dating. Here and elsewhere, records of Eemian and Holocene interglacial periods, and environmental conditions associated with it were targeted. Many sites with late Pleistocene Yedoma Ice Complex have been explored. Lateglacial and Holocene warming induced enormous periglacial landscape changes by widespread permafrost degradation and substantial paleoecological changes. For vast Siberian areas where glacial records are not available, we aim on the establishment of permafrost as paleoclimatic archive, emphasizing peculiarities of permafrost age control and record resolution and stressing the great potential for understanding climate variability on glacial-interglacial timescales in Western Beringia