First report of Thrips carthami Shumsher (Thys.: Thripidae) from Iran

در بررسی فونستیک بال­ریشکداران استان ایلام طی سال­های 1395-1394، نمونه­های متعلق به گونه­ Thrips carthami Shumsher, 1946 از روی برگ­های درختان بادام کوهی واقع در جنگل­های زاگرس (کوه گچان) جمع­آوری و شناسایی شد. این گونه برای اولین بار از ایران گزارش می­شود و به این ترتیب تعداد گونه­های جنس Thrips در ایران به 34 گونه می­رسد. ویژگی­های ریخت‌شناسی و دامنه انتشار جغرافیایی رکورد جدید نیز ارایه می شود

First report of the genus and species Cephalothrips monilicornis (Reuter) (Thysanoptera: Phlaeothripidae) from Doha, Qatar

The genus Cephalothrips Uzel (Thysanoptera: Phlaeothripidae) with eight species in the world, is one the small group in subfamily Phlaeothripinae. Here, Cephalothrips monilicornis (Reuter, 1880) is reported for the first time from Doha, Qatar. Diagnostic morphological characters and geographical distribution of the newly recorded species are briefly discussed. The specimens examined are deposited in the collection of Department of Plant Protection, College of Agriculture, Ilam University, Iran (ILAMU)

Thrips (Insecta: Thysanoptera) of China

A new checklist of Thysanoptera from China (including Taiwan) is provided. In total 566 species in 155 genera are listed, of which there are 313 species in the suborder Terebrantia, comprising 290 species in 74 genera in family Thripidae, 18 species in three genera in Aeolothripidae, two species in one genus in Melanthripidae and three species in one genus in Merothripidae. In the suborder Tubulifera 253 species in 76 genera are listed in the single family Phlaeothripidae. Two species, Aeolothrips collaris and Odontothrips meliloti, are newly recorded for the fauna of China

Identification of Thrips Species on Garlic Fields in Hamedan Province and Determination of Dominant Species

Introduction: Garlic (Allium sativum) (family Amaryllidaceae) are one of important crops in Hamedan province. There are several insects and mites that by feeding on this plant cause to damage garlic yield. Among the insect pests, the most dangerous pests of garlic are thrips species (Insecta: Thysanoptera). In this group, onion thrips, Thrips tabaci Lindeman is widely distributed and is a dominant species. Its sap sucking causes tiny and silvery spots on the leaves that may spread all over the leaf surface with an unfavorable effect on yield production. In addition, imagoes and larvae living and feeding on this plant cause more serious damages. Due to serious damage of thrips in garlic fields of Hamedan province, it is necessary to identify thripsspecies for pest control programs. Materials and Methods: Thrips specimens were collected on leaves of garlic in Hamedan province (Maryanaj and Heydareh) situated in the west of Iran, during 2012–2013. Herein, specimens were collected by shaking plants to white dish and specimens were kept in 70 % ethanol and transferred to the laboratory. All collected material was macerated in 5% KOH and subjected to dehydration in an ethanol series before being mounted onto glass slides. Subsequently, thrips specimens mounted onto slides in Hoyer’s medium form of the protocol given in Mirab-balou and Chen (2010). All descriptions, measurements and photos were made with a Leica DM IRB microscope, with a Leica Image 1000 system. Thrips specimens were identified by author in species level. Specimens are deposited in the collection of Department of Plant Protection, College of Agriculture, Ilam University, Iran (ILAMU). In addition, dominant thrips species were also determined. For this purpose, regular samplings were done in fields of two regions of Hamedan province every two weeks, and totally, 2040 specimens (adult) were collected. Results and Discussion: The world fauna of Thysanoptera, are presently known to be consisted of about 6000 species placed in two suborders and nine families. A large number of thrips species are considered pests, because they feed on economical crops. In this study, a total of eight species in seven genera and three families were collected and identified, including Aeolothrips intermedius Bagnall and Rhipidothrips gratiosus Uzel from family Aeolothripidae, Aptinothrips rufus (Haliday), Frankliniella intonsa (Trybom), Scolothrips longicornis Priesner, Thrips alliorum (Priesner) and Thrips tabaci Lindeman from family Thripidae, and Haplothrips reuteri (Karny) from family Phlaeothripidae. All of the species existed in two years in both two regions. Thrips tabaci was the dominant species (64.89%) in garlic fields. Among the predatory thrips, Aeolothrips intermedius and Scolothrips longicornis were present in greater numbers. However, their number was not enough to reduce the number of phytophagous thrips. The predatory species feeds mainly on the larvae and imagoes of onion thrips but they feed on mites as well. An identification key for thrips species associated with garlic is also given. Conclusion: In this study, eight species of thrips were collected on garlic fields of Hamedan province which Thrips tabaci was dominant species (64.89%); and three of which were identified as predatory species. Up to the present, several thrips were collected and recorded from Hamedan province by the author, but there is no study on thrips associated to garlic; therefore, this study was firstly carried out in Hamedan province. There are several insect pests on garlic, and T. tabaci was also reported as important pest on garlic fileds in this province. Onion thrips, T. tabaci is one of the important pests in the world, and it has more than 300 host plants. At present, it is widely distributed in Iran and is a key insect pest in most onion and cotton cultivation areas as well as ornamental plants. In addition, thrips species in the genera Thrips and Frankliniella spread plant diseases through the transmission of viruses, such as Tospoviruses, Tomato Spotted Wilt Virus, and the Impatiens Necrotic Spot Viruses; So, it is necessary to pay more attention to the management of T. tabaci . In garlic crops of Hamedan province, T. tabaci was established on the plant from earlier time to near harvest; and because of high damaging, the farmers used several insecticides within one year

Asprothrips punctulosus Tong, Wang & Mirab-Balou, 2016, sp. n.

<i>Asprothrips punctulosus</i> sp. n. <p>(Figs 5–6, 14–20, 23)</p> <p> <i>Female</i>. Macroptera (Fig 5). Body generally brown. Antennal segments I–III and VI –VIII brown, IV clear yellow, V yellow with distal half brown (Fig 19). Fore wing greyish brown (Fig 18). All femora and tibiae brown, tarsi yellow. Abdomen tergites I–VII brown laterally, VIII–X brown, tergites II–VII light brown between setal pair S2 with dark brown spot posteromedially (Figs 5, 15).</p> <p>Head (Fig 14) 1.8 times as wide as long; two pairs of minute ocellar setae present, pair II situated at middle between anterior ocellus and compound eye, pair III arising near anterior margin of posterior ocelli within ocellar triangle; 3 pairs of postocular setae present, first pair below the hind ocelli; vertex between eyes including ocellar triangle sculptured with irregularly reticulation, occipital region of vertex sculptured with transverse dotted lines. Mouth cone short and rounded. Antennal segment II globular with ridges on striae, III with a noticeable pedicel, IV–V with short pedicel, IV subequal to VI in length; V with small simple outer sense cone, VI with 3 sense cones, inner one longest arising submedially, reaching apex of VIII; microtrichia rows present on III–VI, segments III and VI with 3 microtrichia rows, IV and V with 4 rows.</p> <p>Pronotum (Fig 14) transverse approximately twice as wide as long, sculptured with transverse anastomosing lines bearing internal granules on anterior 1/3 and transverse anastomosing striae bearing numerous internal wrinkles and granules on posterior 2/3, with less than 30 minute discal setae, 4 pairs of posteromarginal setae. Mesonotum (Fig 16) smooth on anterior third with two campaniform sensilla, posterior 2/3 with transverse anastomosing smooth striae, median setal pair situated far from posterior margin. Metanotum (Fig 16) reticulate medially with irregular granules between reticles, median setae far back from anterior margin, campaniform sensilla absent. Leg surface reticulate, all tarsi 2-segmented. Fore wing costa with about 22 setae, first vein with 6– 7 proximal and 2 distal setae, second vein with 7 setae; main posterior fringe hairs straight except for some fine hairs weakly wavy; microtrichia of fore wing only present on two veins, along margins and apical sixth, not uniformly covering surface of fore wing.</p> <p>Abdominal tergites I–VII laterally with strong transverse sculpture lines bearing short microtrichia on anterior 1/3 and tuberculate microtrichia on posterior 2/3 but smooth between setal pair S2 (Fig 15); paired campaniform sensilla situated between setae S1 and S2, much closer to S2 on tergites II–VII; tergites VIII–IX entirely covered with strong transverse sculpture bearing tuberculate microtrichia except for groove medially; VIII with posteromarginal comb, but minute medially; posterior margin of IX overhanging X, medially with a pair of thick and strong setae (Fig 17); tergite X with a complete longitudinal dorsal split. Abdominal sternites II–VII sculptured with transverse anastomosing striae bearing short microtrichia laterally, sternite II with 2 pairs of setae on posterior margin, III–VII each with 3 pairs of setae.</p> <p> <b>Measurements</b> (holotype female in microns). Total distended body length 1180. Head length (width) 90 (175); eye length (width) 70 (42). Pronotum length (width) 106 (218). Length of antenna 203; length (width) of antennal segments I 18 (27), II 28 (29), III 34 (20), IV 36 (20), V 30 (18), VI 34 (15), VII 10 (6), VIII 13 (4). Fore wing length740.</p> <p> <i>Male.</i> Macroptera (Fig 6). Most character and color pattern states similar to female. Antennal segments brown except for I yellowish brown and IV clear yellow (Fig 20). Abdominal sternites without pore plates. The paramere shape of genitalia as Figure 23.</p> <p> <b>Measurements</b> (paratype male in microns). Total distended body length 1120. Head length (width) 95 (165); eye length (width) 68 (43). Pronotum length (width) 90 (200). Length of antenna 186; length (width) of antennal s egments I 20 (26), II 26 (26), III 32 (18), IV 32 (18), V 26 (17), VI 30 (15), VII 9 (6), VIII 11 (4). Fore wing length 630.</p> <p> <b>Specimens examined.</b> Holotype female (in SCAU): <b>CHINA</b>, <b>Jiangxi</b>: Chongyi County, Yangling National Forest Park (25°39'N, 114°18'E), collected from an unidentified dicotyledonous shrub, 22.viii.2015 (Xiaoli Tong). 1 female and 11 males, collected with holotype, same locality and collector.</p> <p> <b>Distribution.</b> China (Jiangxi).</p> <p> <b>Etymology.</b> The specific epithet, <i>punctulosus</i>, meaning “many spots”, is from the Latin adjective and refers to the abdominal tergites II–VIII each with a brown spot posteromedially.</p> <p> <b>Comments.</b> This species shares morphological affinities with <i>bucerus</i> as both species have similar sculpture on the body surface and a dark brown spot on abdominal tergites II–VII posteromedially, and the males lack sternal pore plates. However, in addition to the different shape of the male parameres (Figs 22–23), <i>punctulosus</i> differs from <i>bucerus</i> by the clear yellow antennal segment IV which is subequal in length to VI, and segment V is bicolored; the posterior margin of IX bears a pair of thick and strong setae medially, and tergite X has a complete longitudinal dorsal split. In contrast, in <i>bucerus</i> the antennal segments are uniformly dark brown except segment I yellowish brown, and IV is much longer than VI; the posterior margin of tergite IX bears a pair of short horn-like setae medially, and tergite X has an incomplete longitudinal dorsal split. The sexes of <i>punctulosus</i> are similar to each other in most characters and color pattern (Figs 5–6), but the male of <i>bucerus</i> is quite different from the female in color pattern (Figs 3–4).</p>Published as part of <i>Tong, Xiaoli, Wang, Zhaohong & Mirab-Balou, Majid, 2016, Two new species and one new record of the genus Asprothrips (Thysanoptera: Thripidae) from China, pp. 181-188 in Zootaxa 4061 (2)</i> on pages 185-187, DOI: 10.11646/zootaxa.4061.2.8, <a href="http://zenodo.org/record/256313">http://zenodo.org/record/256313</a&gt

Mortality Rate of Frankliniella occidentalis Under Recommended Concentration of Some Insecticides and the Amount of Its Attraction to Colored Sticky Traps in Apple Orchard

Introduction: Western flower thrips, Frankliniella occidentalis (Pergande) is a serious pest of fruit crops in flowering stage worldwide. Many researches have been studied different methods of western flower thrips control. Two control methods including pesticide application and the use of colored sticky traps are commonly used by farmers due to the ease of use and low running costs. Chemical control is known as the main tool in controlling of this pest. In recent decades, mixing several kinds of pesticides by farmers has been become common strategy which seems to be due to their synergistic effects and decreasing of pest resistance one. The current study was done to detect the best color sticky trap in monitoring and to determine the most effective pesticide in controlling western flower thrips. Materials and Methods: In the current study, the pest attraction rate by 3 sticky color traps (Blue, Yellow, White) and efficacy of five pesticides were investigated against western flower thrips in two separately randomized complete block design in the apple orchard around Shiraz city. The treatments included: deltametrin + imidacloprid, acetamiprid, antifeedant, azadirachtin and oxydemeton methyl. Mortality percent of insects in different treatments was calculated using the Henderson-Tilton formula. Before conducting the experiment, trees was not treated by any pesticides for one year. Each trap was hung in the middle of the trees´ canopy at about 1.5 meter height from the ground. Sticky cards (10 x 25 cm) were visited weekly to record the number of the captured western flower thrips. Each trap was replaced with new one weekly. Number of western flower thrips was analyzed through one-way analysis of variance (ANOVA) using the SPSS version 9. The significant differences among means were compared using the Duncan's multiple range test at 95% confidence interval whenever treatment effects were significant. Results and Discussion: Analysis of variance showed that there were significant differences among treatments after 14 days post-treatment (F5, 10 = 6.51, P = 0.006). Mean comparisons showed that treatments deltametrin + imidacloprid, acetamiprid and antifeedant had significant difference with control. Pesticide efficacies based on Henderson-Tilton formula at 14 days post-treatment showed that acetamiprid provided the best control (83.3%) of F. occidentalis, followed by deltametrin + imidacloprid (77.7%) and antifeedant (70.1%). Effectiveness of acetamiprid, which is a systemic pesticide and belongs to neonicotinoids, against a wide range of insect pests has been reported by many researchers. The treatment oxydemeton methyl caused low mortality (less than 30%) in F. occidentalis. The reason of that may be related to resistance of this insect to oxydemeton methyl. This pesticide is older than other tested compounds and has been used by farmers against fruit pests from long time ago. Further studies are needed to confirm these findings like bioassay of this pesticide. In addition, some researchers have shown that inappropriate application of chemical compounds resulted in high level and widespread resistance of thrips to chemicals. Results of color traps showed that blue traps attracted significantly more thrips than other colors in the all sampling dates. On average, the blue, yellow and white traps captured 34.31 ± 13.85, 12.56 ± 4.78 and 4.87 ± 1.83 thrips respectively. In general, there are many studies have been conducted to determine the effect of color traps on thrips. Some studies have reported that thrips are attracted to blue, while the others have reported that yellow is more attractive for them. There are evidences that F. schultzei is also caught by red color. With regard to color preference of western flower thrips, there are many disagreements among researchers. Different researchers have stated different reasons for these contradictions that the most important are the availability of food sources, thrips density, environmental conditions and the duration time of the experiment carrying out. Conclusion: The results of the present study indicated that a remarkable ability of blue traps in catching of the western flower thrip. In addition, results showed that acetamiprid, deltametrin + imidacloprid and antifeedant could considerably reduce the population density of F. occidentalis. Although the pesticides acetamiprid, deltametrin and imidacloprid have been recommended in IPM programs, in organic farming, it is approved that application of pesticides are much more limited. It seems that antifeedant is environment-friendly and can be recommended in organic farming. However, more research is needed to explore the impact of this compound on other agricultural systems. In conclusion, blue sticky color and deltametrin + imidacloprid, acetamiprid and antifeedant could be useful in IPM planes against western flower thrips

Two new synonyms among Chinese Thripinae (Thysanoptera: Thripidae)

Mirab-Balou, Majid, Tong, Xiao-Li (2015): Two new synonyms among Chinese Thripinae (Thysanoptera: Thripidae). Zootaxa 3941 (1): 149-150, DOI: http://dx.doi.org/10.11646/zootaxa.3941.1.1

Asprothrips bimaculatus Michel & Ryckewaert

Asprothrips bimaculatus Michel & Ryckewaert (Figs 1 –2, 21) Asprothrips bimaculatus: Michel & Ryckewaert 2014: 496. Female. Adequately described and illustrated by Michel & Ryckewaert (2014). Male. Similar to female, but body paler; abdominal tergite VI without a pair of brown patches; sternites II–VII each with one small, rounded pore plate; genitalic paramere shape see Figure 21. Specimens examined. CHINA, Guangxi, Nanning City, Nalou Town (22 ° 36 'N, 108 ° 39 'E), 1 male, 1 female collected from Pennisetum purpureum (Poaceae), 3.x. 2012 (Shulan Yang). Taiwan, Pingdong, Kenting, Ngoluanpi Park (21 ° 54 ' 09″N, 120 ° 51 ' 15 ″E), 1 female from grass (Poaceae), 5.viii. 2012 (Xiaoli Tong). Distribution: Martinique in the French West Indies, Malaysia (ThripsWiki 2015), China. Comments. Unlike the type specimens of A. bimaculatus that have a pair of brown patches on abdominal tergite VI, the brown patches in the Chinese specimens are faded and only faintly visible. However, the male that is recorded here for the first time is devoid of such brown patches. With the description below of two new species from China, Asprothrips now includes seven described species that are all from Asia but with seminigricornis widespread in greenhouses of several countries (Mound 1999, Sartiami & Mound 2013), and bimaculatus that shows a remarkable disjunct distribution between Martinique in the Caribbean and Asia (Malaysia) (ThripsWiki 2015 —identified by Ng Foo Yong). The new record of this species from China, including the male, supports the probability that it is originally from Asia, in common with the other members of the genus, and has been introduced to the western hemisphere.Published as part of Tong, Xiaoli, Wang, Zhaohong & Mirab-Balou, Majid, 2016, Two new species and one new record of the genus Asprothrips (Thysanoptera: Thripidae) from China, pp. 181-188 in Zootaxa 4061 (2) on page 182, DOI: 10.11646/zootaxa.4061.2.8, http://zenodo.org/record/25631

A new species of the genus Karnyothrips (Thysanoptera,\ua0Phlaeothripidae) from China

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Asprothrips bucerus Tong, Wang & Mirab-Balou, 2016, sp. n.

Asprothrips bucerus sp. n. (Figs 3 –4, 7–13, 22) Female. Macroptera (Fig 3). Body generally brown, abdomen slightly paler than head and thorax. Antennae dark brown except for segment I yellowish brown, segments VI–VIII lighter than II–V (Fig 8). Fore wing greyish brown (Fig 13). All femora and tibiae brown but tarsi yellow. Abdominal tergites I–VII brown laterally and VIII–X brown, tergite I yellowish brown with a dark brown spot anteromedially (Fig 3), tergites II–VII yellowish brown medially with a dark brown spot posteromedially (Figs 11–12). Head (Fig 7) 1.8 –2.0 times as wide as long; two pairs of minute ocellar setae present, pair II situated at middle between anterior ocellus and compound eye, pair III arising near anterior margin of posterior ocelli within ocellar triangle; postocular setal row of four setae extending near cheeks; vertex between eyes including ocellar triangle sculptured with irregularly reticulation, occipital region of vertex sculptured with transverse dotted lines. Mouth cone short and rounded. Antennal segment II largest, globular with ridges on striae, III small and pedicellate (Fig 8), IV longest, longer than VI; V with short pedicel and a small simple outer sense cone, VI with 3 sense cones, inner one longest arising medially, reaching apex of segment VIII; microtrichia rows present on segments III–V, segment III with 3 microtrichia rows, IV with 5 dorsal rows, V with 3 rows, VI with only a few sparse microtrichia. Pronotum (Fig 7) transverse approximately twice as wide as long, sculptured with transverse dotted lines on anterior half, posterior fourth with some anastomosing striae bearing numerous internal granules, with about 40 minute discal setae and 4 pairs of posteromarginal setae. Mesonotum (Fig 10) with transverse anastomosing striae without internal wrinkles or granules, two campaniform sensilla on anterior fourth, median setal pair situated far from posterior margin. Metanotum (Fig 10) reticulate medially with wrinkles between reticles, median setae far back from anterior margin, campaniform sensilla absent. Leg surface reticulate, all tarsi 2 -segmented. Fore wing costa with 22–24 setae, first vein with 8 proximal and 2 distal setae, second vein with 9–11 setae; main posterior fringe hairs straight except for some fine hairs weakly wavy; microtrichia of fore wing only present on two veins, along margins and apical sixth, not uniformly covering surface of fore wing. Abdominal tergites I–VII laterally with strong transverse sculpture lines bearing short microtrichia on anterior 1 / 3 and tuberculate microtrichia on posterior 2 / 3, smooth between setal pair S 2 (Fig 11); paired campaniform sensilla between setae S 1 and S 2, much closer to S 2 on tergites II–VII; VIII–IX entirely covered with strong transverse sculpture bearing tuberculate microtrichia except for groove medially; VIII with posteromarginal comb, but minute medially; posterior margin of IX overhanging X, medially with a pair of short horn-like setae directed medially (Fig 12); tergite X with incomplete longitudinal dorsal split, divided only in distal half. Abdominal sternites II–VII sculptured with transverse anastomosing smooth striae; II with 2 pairs of setae on posterior margin, III–VII each with 3 pairs. Measurements (holotype female in microns). Total distended body length 1100. Head length (width) 85 (170); eye length (width) 62 (38). Pronotum length (width) 100 (200). Length of antenna 190; length (width) of antennal segments I 24 (28), II 24 (31), III 34 (22), IV 36 (22), V 25 (19), VI 25 (13), VII 10 (6) and VIII 12 (4). Fore wing length 630. Male. Macroptera (Fig 4). Similar in structure but different in color to female. Body generally brown except abdomen much paler than head and thorax. Antennal segments I–II yellow, III yellowish brown, IV–VIII brown (Fig 9). Fore wing greyish brown. Fore and mid legs brown, hind leg yellow. Abdominal tergites I–III yellowish brown, II–VII brown laterally but yellow between setal pair S 2 except for brown spot posteromedially, VIII–X brown. Abdominal sternites without pore plates. Paramere shape of genitalia as Figure 22. Measurements (paratype male in microns). Total distended body length 895. Head length (width) 65 (145); eye length (width) 55 (35). Pronotum length (width) 88 (167). Length of antenna 175; length (width) of antennal s egments I 22 (24), II 22 (26), III 30 (19), IV 32 (20), V 26 (17), VI 22 (14), VII 9 (6), VIII 12 (4). Fore wing length 550. Specimens examined. Holotype female (in SCAU): CHINA, Guizhou: Maolan National Nature Reserve, Weng’ang (25 ° 14 ' 12 ″N, 107 ° 52 ' 30 ″E), collected from Viburnum henryi (Caprifoliaceae), 19.vii. 2015 (Zhaohong Wang). Paratypes (in SCAU): 5 males, 2 females from Viburnum henryi (Caprifoliaceae), collected with holotype, 1 male, 1 female from Beilschmiedia kweichowensis (Lauraceae), same locality and collector. Guangdong, Xinyi City, Mt. Tianmashan (22 ° 27 'N, 110 ° 41 'E), 4 females on 11.vi. 2011 and 6 females on 18.ix. 2011 (Tao Song), both from Sarcandra glabra (Chloranthaceae). Gaozhou City, Yuntan Town, Mt. Sanguanshan, 2 females from Syzygium jambos (Myrtaceae), on 5.ix. 2015 (Zhaohong Wang). Non-paratype: CHINA, Guangdong, Boluo County, Mt. Luofushan (23 ° 16 'N, 114 °03'E), 1 female, from Cinnamomum camphora (Lauraceae), 14.x. 1976 (Weiqiu Zhang). Distribution. China (Guizhou, Guangdong) Etymology. The specific epithet, bucerus, is from the Latin adjective, meaning “horned”, in reference to the posterior margin of abdominal tergite IX in female having a pair of short horn-like setae medially. Comments. According to the collection records above, this species is associated with a wide range of unrelated plants. It is similar to fuscipennis in having the body, antennae and fore wing brown, but is distinguished by the following combination of characteristics: (1) segment I yellowish brown and segment IV longest; (2) tergite IX entirely covered with strong transverse sculpture bearing tuberculate microtrichia and posterior margin with a pair of short horn-like setae medially; (3) tergite X with incomplete longitudinal dorsal split, divided only in distal half. In contrast, the antennae of fuscipennis are uniformly greyish brown with segment VI longest; tergite IX reticulate anteriorly and striate posteriorly, posterior margin with pair of long fine setae medially; tergite X with complete longitudinal dorsal split. Zhang & Tong (1988) identified one poorly preserved specimen of this new species as fuscipennis. At that time, this was the only specimen of Asprothrips known from China, therefore fuscipennis has not yet been recorded from this country.Published as part of Tong, Xiaoli, Wang, Zhaohong & Mirab-Balou, Majid, 2016, Two new species and one new record of the genus Asprothrips (Thysanoptera: Thripidae) from China, pp. 181-188 in Zootaxa 4061 (2) on pages 182-185, DOI: 10.11646/zootaxa.4061.2.8, http://zenodo.org/record/25631