Brooding behaviour in Ophioderma wahlbergii, a shallow-water brittle star from South Africa

Includes bibliographical references.The brooding behaviour and brooded young are described for Ophioderma wahlbergii Müller & Troschel 1842, a large, common brittle star from the coastal waters of South Africa. Twenty specimens were collected each month from June 2013 – May 2014 (n = 240). The species was found to be gonochoric

Contributions to the taxonomy of South African hermit crabs (Crustacea: Decapoda: Paguroidea) – integrating microCT scanning and barcoding

Hermit crabs form an important component of the marine benthos and globally more than 1,200 species have been described. In the unique bioregion of South Africa, hermit crabs are poorly known, and the last taxonomic revision of the group was that of K. H. Barnard in 1950, who recorded only 32 species. This study combines morphological taxonomy, threedimensional (3D) micro-computed tomography (µCT) visualisations, and molecular barcoding to add to, revise, and provide an updated listing of, the regional fauna. The first section of the thesis comprises four chapters, each giving a detailed account of a species either new to science, or to the region. The pagurid hermit crab Goreopagurus poorei, a new species and genus record to the country, is reported and described from deep sea habitats along the Agulhas Shelf, extending the distribution by >10,000 km across the Indian Ocean. Furthermore, three species are described as new to science, one each from the three most common families. The first of these, a deep-water species from a genus of the family Parapaguridae that was previously unknown to South Africa, Paragiopagurus atkinsonae n. sp., is fully described and illustrated, and compared with two other parapagurids that each play a dominant role in the regional benthic offshore invertebrate community. The other two species new to science, Diogenes n. sp. from the family Diogenidae, and Pagurus n. sp. from the family Paguridae, inhabit coastal reefs in subtidal waters off southern KwaZulu-Natal. For the first time in crustacean taxonomy, species descriptions, particularly the one of Pagurus n. sp., are informed by, or based on, µCT imagery of calcified body parts. Following on this technique, Chapter 6 is a short presentation of the 3D raw dataset of seven µCT scans of types and rare museum specimens used in this thesis, which is made publicly available for download. The taxonomic use of the scanning method, with disseminating volumetric data of hermit crabs, is discussed briefly. The final section investigates the fauna as a whole. In Chapter 7, 194 cytochrome c oxidase subunit I gene segments (COI ‘barcodes’) of 43 nominal species plus 12 additional putative species (n = 55 species-like units) were used to validate morphological identifications. Testing this dataset revealed high barcoding efficacy, with nearly 99% identification success rates, and with the best Kimura 2-parameter distance to safely delimit species of hermit crabs of about 3.5%. Chapter 8 updates the regional fauna and provides taxonomic accounts for 62 nominal species which have either been added subsequent to the previous monographic review, or which have undergone taxonomic revision since that time. Of these, 12 are added for the first time here, increasing the number of known South African hermit crab species to 72, an expansion of 56% since Barnard, and about 20% since a recent species list published by W. Emmerson in 2016. Because colour images are provided for 51 out of 72 species, Chapter 8 can also be used as a preliminary guide. However, this study has shown that the hermit crabs of South Africa are by far more diverse than originally thought, and the summary, which includes only the 72 nominal taxa and none of the additional 10 putative species included in the barcoding dataset, is speculated to be only 60–70% complete. Future taxonomic work, especially in the genera Diogenes and Paguristes, will most likely result in many more species descriptions. Therefore, this current study is to be seen as important step towards a fully illustrated taxonomic catalogue on the South African hermit crabs to be produced in the near future

Additions to the marine decapod (Crustacea: Decapoda) fauna of South Africa

This report adds five previously unreported species to the decapod crustacean fauna of South Africa, as well as removing one species previously listed in error. It also documents locality (and/or reference specimen) data for 12 other species, most of which had been depicted in regional field guides, but without reporting when and where they had been collected. Almost all the species added were already known from adjacent African countries and their ranges are here extended into South Africa. Although some of these records are based on photographs, rather than collected specimens, it is argued that such records should be accepted as adequate evidence for inclusion of at least visually-distinctive crustacean species into the regional fauna

Ocean Species Discoveries 1–12 — A primer for accelerating marine invertebrate taxonomy

Discoveries of new species often depend on one or a few specimens, leading to delays as researchers wait for additional context, sometimes for decades. There is currently little professional incentive for a single expert to publish a stand-alone species description. Additionally, while many journals accept taxonomic descriptions, even specialist journals expect insights beyond the descriptive work itself. The combination of these factors exacerbates the issue that only a small fraction of marine species are known and new discoveries are described at a slow pace, while they face increasing threats from accelerating global change. To tackle this challenge, this first compilation of Ocean Species Discoveries (OSD) presents a new collaborative framework to accelerate the description and naming of marine invertebrate taxa that can be extended across all phyla. Through a mode of publication that can be speedy, taxonomy-focused and generate higher citation rates, OSD aims to create an attractive home for single species descriptions. This Senckenberg Ocean Species Alliance (SOSA) approach emphasises thorough, but compact species descriptions and diagnoses, with supporting illustrations and with molecular data when available. Even basic species descriptions carry key data for distributions and ecological interactions (e.g., host-parasite relationships) besides universally valid species names; these are essential for downstream uses, such as conservation assessments and communicating biodiversity to the broader public

Differentiation of three common deep-water hermit crabs (Crustacea, Decapoda, Anomura, Parapaguridae) from the South African demersal abundance surveys, including the description of a new species of Paragiopagurus Lemaitre, 1996

Deep-water hermit crabs of the family Parapaguridae can be abundant (up to 20 kg or 1000 hermit crab individuals per haul) in the trawl bycatch collected during South African demersal abundance research surveys. Until recently, only two parapagurid species had been recognized in the bycatch; Parapagurus bouvieri Stebbing, 1910, and Sympagurus dimorphus (Studer, 1883). Detailed examination of numerous samples of parapagurid specimens from research surveys revealed the existence of a third, undescribed species previously confounded with S. dimorphus, but in fact belonging to a different genus. This new species, Paragiopagurus atkinsonae sp. n. is the 25th in the genus Paragiopagurus Lemaitre, 1996, and has been found only in a small region on the West Coast shelf of South Africa, at depths of 199–277 m. The species is herein fully described and illustrated, including colour images, µCT scans of selected body parts, and CO1 barcode data. The new species is morphologically most similar to P. ventilatus Lemaitre, 2004, a species associated with hydrothermal vents, but differs in armature of the fourth antennal segment (armed with a spine on the dorsolateral distal angle vs. unarmed in P. ventilatus); setation of the antennal flagella (nearly naked vs. with dense setae in P. ventilatus); plumose setation on the third maxillipeds and basal segments of chelipeds (absent vs. present in P. ventilatus); number of rows of scales on the propodal rasp of pereopod 4 (two or three rows vs. one row in P. ventilatus); and degree of telson asymmetry (weakly asymmetrical vs. strongly asymmetrical in P. ventilatus). Paragiopagurus atkinsonae sp. n. is superficially similar to S. dimorphus, with males of the two species showing the same extreme degree of sexual dimorphism on the right cheliped, general light orange colouration, and frequent use of colonial zoanthid carcinoecia for pleonal protection. To aid in future identifications and to facilitate data gathering during surveys, a comparison of P. atkinsonae sp. n. with S. dimorphus is provided, along with descriptions of colouration and photographs of live specimens of all three parapagurid species. Information on taxonomy of the species is summarized, as well as knowledge of their distribution in the demersal research survey regions of South Africa

Odontodactylus japonicus

<i>Odontodactylus japonicus</i> (de Haan, 1844) new record <p>Fig. 9</p> <p> <i>Gonodactylus japonicus</i> de Haan, 1844: pl. 51 fig. 7 [type locality Japan].</p> <p> <i>Gonodactylus edwardsii</i> Berthold, 1845: 48.</p> <p> <i>Gonodactylus japonicus</i> – de Haan 1849: 255 [text]. — Miers 1880: 116.</p> <p> <i>Odontodactylus japonicus</i> – Alexander 1916: 10. — Holthuis 1941: 276. — Stephenson & McNeill 1955: 248–249. — Stephenson 1960: 61. — Manning 1965: 260; 1967b: 7–10, fig. 2; 1995: 20, 82. — Graham <i>et al.</i> 1993: 73. — Yamaguchi & Baba 1993: 176–178, fig. 9. — Ahyong & Norrington 1997: 103. — Moosa 2000: (list). — Ahyong 2001: 81–83, fig. 39. — Liu 2008: 1267 (list).</p> Diagnosis <p>Ocular scales oblique to bodyline, appressed medially. A2 scale with anterior margin smooth, without setae in adults. Rostral plate triangular, but appearing trapezoid dorsally; lateral margins sinuous; apex deflexed. Raptorial claw dactylus with 5–8 teeth on inner margin; proximal margin strongly inflated; without basal notch. AS 1–5 posterolateral angles rounded, unarmed in adults. Telson mid-dorsal surface with distinct MD carina and 4 longitudinal carinae either side of midline (double accessory MD; anterior SM; carina of inner IM denticle) in addition to carinae of primary teeth. Uropodal exopod proximal distinctly longer than distal segment; outer margin with 10–12 movable spines, distalmost evenly tapering (juveniles) to spatulate with blunt or minute spinular apex (adults).</p> Material examined <p> MOZAMBIQUE • 1 ♀ (TL 110 mm); north of Beira; 19°49′00.0″ S, 36°05′00.0″ E; 54 m depth; 14 Jun. 1994; RV <i>Algoa</i> Mozambique Scad Survey; bottom trawl; SFI; SAMC – A041710.</p> Colour in alcohol <p>Faded to creamy yellow with telson still retaining salmon colour of live specimen. Eyes green. Uropod exopod distal segment dark and proximal segment with distal third dark. Uropodal protopod, exopod proximal segment and endopod covered in dark speckles.</p> Colour in life <p>Overall colour salmon (Ahyong 2001) or males’ bright salmon red; females’ salmon red anteriorly and blue, green posteriorly (Manning 1967b). A2 scale salmon proximally, pink distally. Uropods yellow; exopod with outer movable spines yellow orange with blue posterior margin; endopod and distal segment of exopod with red setae.</p> Measurements <p>Female (n = 1) TL 110 mm. CI 404. A1 peduncle 0.60CL. A2 scale 0.98CL. Uropod exopod distal segment length 1.69 times proximal segment length. The largest specimen is reported from Australia at TL 175 mm (Ahyong 2001).</p> Distribution and habitat <p>Indo-West Pacific; Australia and Japan to Western Indian Ocean from Seychelles, Madagascar, and now southern Mozambique [Sofala]. Associated with flat sandy or shelly substrata; 30–100 m depth.</p> Remarks <p> Although this is a new record for the Mozambique Channel, <i>O. japonicus</i> has been reported from Madagascar, not far from the locality recorded herein. The Madagascan specimen described by Manning (1967b) was collected from Ambovombe’s southern coast in shallow water at around 60 m, a similar depth to the female specimen collected from the Mozambique Channel at 54 m. The present specimen agrees in all aspects with Manning’s (1967b) diagnosis, as well as the most recent report by Ahyong (2001) of <i>O. japonicus</i> from Australia.</p> <p> Only <i>O. scyllarus</i> (Linnaeus, 1758) and <i>O. hawaiiensis</i> Manning, 1967 share the appressed ocular scales and the double accessory MD carinae either side of MD carina of telson with <i>O. japonicus</i>. <i>Odontodactylus japonicus</i> is distinguished from these similar species in having the longitudinal carina extending anteriorly from the inner IM denticle of the telson (Fig. 9E), as well as the absence of a posterolateral spine on AS 5 in adults (Fig. 9C). In addition, the number of teeth on the dactylus of the raptorial claw differs between the species; while <i>O. scyllarus</i> has two or three teeth, <i>O. hawaiiensis</i> and <i>O. japonicus</i> have more than five teeth. <i>Odontodactylus hawaiiensis</i> has not yet been recorded off the mainland of southern Africa.</p> <p> Postlarvae settle at sizes of 19–20 mm. Along with the juveniles of <i>O. japonicus,</i> the postlarvae differ from adult specimens in the following characteristics diagnostic for the species: the antennal scale bears setae on its anterior margin and a posterolateral spine on AS (3)4–5, the distal movable spines on the proximal segment of the uropodal exopod are spiniform instead of spatulate and the second accessory MD carina of the telson is underdeveloped (Manning 1967b; Ahyong 2001). Specimens above TL 60 mm are considered adult and exhibit ‘adult’ diagnostic features (Fig. 9A–F). No juvenile specimens were available to the present study.</p>Published as part of <i>Brokensha, Rouane, Landschoff, Jannes & Griffiths, Charles, 2023, New and notable stomatopods (Crustacea, Stomatopoda) from Mozambique, pp. 148-179 in European Journal of Taxonomy 900 (1)</i> on pages 166-168, DOI: 10.5852/ejt.2023.900.2309, <a href="http://zenodo.org/record/10082989">http://zenodo.org/record/10082989</a&gt

Manningia australiensis Manning 1970

<i>Manningia australiensis</i> Manning, 1970 new record <p>Fig. 3</p> <p> <i>Manningia australiensis</i> Manning, 1970a: 78–81, fig. 1 [type locality off Gillett Cay, Swain Reefs, Australia].</p> <p> <i>Manningia vinogradovi</i> Makarov, 1978: 183, fig. 4.</p> <p> <i>Manningia thorsoni</i> Naiyanetr, 1987: 239, figs 2–3.</p> <p> <i>Manningia australiensis</i> – Makarov 1978: 183. — Moosa 1991: 154. — Manning 1995: 19, 34. — Ahyong 1997: 331, 332; 2001: 23–24, fig. 11. — Debelius 1999: 291. — Liu 2008: (list).</p> Diagnosis <p>A2 protopod with 1 ventral papilla; with blunt dorsal tooth and anteriorly-directed ventral spine. Rostral plate cordiform to subpentagonal; apex acute; rounded laterally. Raptorial claw merus without outer inferodistal spine. TS6–8 without carinae. AS 1–5 with MG carina and submarginal sulcus; unarmed posterolaterally. AS 6 with armed SM, IM, and LT carinae. Telson dorsolateral surface with distinct MD carina and 3 longitudinal carinae in addition to MG carina as follows: accessory MD carina interrupted, composed of 4 or 5 posteriorly directed spines; anterior IM carina uninterrupted, armed posteriorly; LT carina sinuous and recurved proximally terminating in 1 or 2 spines. Telson posterior margin with 3 broad horizontal lobes between SM and IM teeth, inner lobe with inner IM denticle on margin; with 2 broad horizontal lobes between IM and LT teeth. Telson ventral surface with depressed outer IM and LT denticles only. Uropodal protopod inner margin with 8–10 slender spines; exopod outer margin with 6–8 movable spines; endopod without carinae.</p> Material examined <p> MOZAMBIQUE • 1 ♂ (TL 30 mm); Sofala Province, Mozambique Channel; 20°13′10.0″ S, 35°55′23.4″ E; 66–67 m depth; 22 Oct. 2017; RV <i>Agulhas II</i>; dredge; stn IIOE2 M205 INV264A; SAMC IIOE2 MOZ125.</p> Colour in alcohol <p>Mostly faded, but with dorsal mottling of dark-brown pigment. Telson carinae with traces of dark colouration and dark-brown along lateral margins of primary teeth. The holotypic account (Manning 1970a) of preserved material agrees with the present material. Colour in life unknown.</p> Measurements <p>Male (n = 1) TL 30 mm. A1 peduncle 0.94CL. A2 scale 0.43CL. Largest known specimen at TL 32 mm from Queensland, Australia (Ahyong 2001).</p> Distribution and habitat <p>Vietnam, Thailand, New Caledonia, Papua New Guinea, Australia and now the Western Indian Ocean from Mozambique [Sofala]. Fine to coarse sand or shell substrates at depths 20– 93 m.</p> Remarks <p> This is the first record from the Western Indian Ocean, as well as the first record of the family Eurysquillidae from southern Africa. The depth and habitat profile matches that described for specimens from Australia and New Caledonia (Manning 1970a; Moosa 1991; Ahyong 2001). Makarov’s (1978) specimen of <i>M. vinogradovi</i> from Vietnam and Naiyanetr’s (1987) <i>M. thorsoni</i> described from Thailand have subsequently been found to be indistinguishable from <i>M. australiensis</i> and are now considered synonyms (Ahyong 2001). These species synonyms expanded the species range, which is now further extended across the Indian Ocean to the Mozambique Channel.</p> <p> <i>Manningia australiensis</i> is distinguished from the other eleven species of <i>Manningia</i> by the shape of the rostral plate being cordiform to sub-pentagonal with rounded lateral margins and a small apical spine (Fig. 3B). The present specimen agrees well with previous accounts. The number of spines of the accessory median carina, as well as the outer margin of the uropodal exopod and the inner margin of the uropodal protopod for the specimen herein lie within the range of previously noted variation in telson and uropodal spination (Moosa 1991; Ahyong 2001). The present specimen exhibits eight movable spines on the outer margin of the uropodal exopod, as well as eight spines on the inner margin of the uropodal protopod (Fig. 3F). Four spines adorn the accessory median carina (Fig. 3E).</p>Published as part of <i>Brokensha, Rouane, Landschoff, Jannes & Griffiths, Charles, 2023, New and notable stomatopods (Crustacea, Stomatopoda) from Mozambique, pp. 148-179 in European Journal of Taxonomy 900 (1)</i> on pages 152-154, DOI: 10.5852/ejt.2023.900.2309, <a href="http://zenodo.org/record/10082989">http://zenodo.org/record/10082989</a&gt

Gonodactylus botti Manning 1975

<i>Gonodactylus botti</i> Manning, 1975 new record <p>Fig. 8</p> <p> <i>Gonodactylus botti</i> Manning, 1975: 289, fig. 1 [type locality Jakarta, Indonesia].</p> <p> <i>Gonodactylus botti</i> – Manning & Lewinsohn 1986: 5, 15 (list), fig. 3. — Manning 1990: 97, 104 (key). — Cappola & Manning 1995: 274–275. — Ahyong 2005: 159.</p> <p> non <i>Gonodactylus chiragra</i> – Holthuis 1967: 26, 41 (list), fig. 7a. — Tirmizi & Manning 1968: 21, fig. 7. [not <i>G. chiragra</i> (Fabricius, 1781)].</p> <p> non <i>Gonodactylellus affinis</i> – Moosa 1991: 155. [not <i>Gonodactylellus affinis</i> (de Man, 1902)].</p> Diagnosis <p>Ocular scales broad, flattened, separate, together slightly broader than 0.50 rostral plate width, anterior margins transverse. Rostral plate basal portion with anterior margins slightly concave; anterolateral angles rounded; lateral margins divergent anteriorly; apical spine just longer than base. Lateral margin of TS6 broader than TS7. Telson with LT tooth indicated by shallow notch in margin of telson between anterolateral angle and apex of IM tooth; mid-dorsal carinae blunt, neither sharp nor crested dorsally and strongly inflated in adults; MD carina unarmed posteriorly; accessory MD carinae forming ‘anchor’; with 8–13 SM denticles. Uropodal exopod distal segment outer margin with 12 movable spines.</p> Material examined <p>MOZAMBIQUE • 1 ♁ (TL 53 mm), 1 ♀ (TL 40 mm); Coconut Bay; 24°00′21.6″ S, 35°30′28.7″ E; depth unknown; 17 May 1973; event no. 19730517; SAMC – A019434.</p> <p>SOUTH AFRICA • 1 ♂ (TL 63 mm), 8 ♀♀ (TL 10–66 mm); Jesser Point, Sodwana Bay; 27°32′33″ S, 32°40′50.0″ E; rock pool at 13.5 m depth; 2Aug. 1976; event no. RW 26; R. Winterbottom leg; S.Ahyong det.; SAMC – A015632.</p> Colour in alcohol <p>Faded yellow, but with scattered blue pigment on all limbs and uropods. Dactylus of raptorial claw white. Raptorial claw with ‘meral spot’ also white. Colour in life unknown.</p> Measurements <p>Male (n = 1) TL 53 mm. A1 peduncle 6.0CL. A2 scale 0.58CL. AWCLI 755.</p> Distribution and habitat <p>Indonesia to Western Indian Ocean from Pakistan, Red Sea, Persian Gulf, Somalia, South Africa and now Mozambique [Inhambane]. Found in southern Africa by Ahyong (2005) from the coral reef systems in both sandy and rocky tidal pools at the base of dead coral in shallow water; shore to 13.5 m depth.</p> Remarks <p> As <i>G. botti</i> is presently understood, historical species records are restricted to localities west of Pakistan (Ahyong 2005). It was first recorded from southern Africa by Ahyong (2005) in the coral reefs of Sodwana Bay, South Africa. This single male specimen from Coconut Bay is the first account of the species from Mozambique.</p>Published as part of <i>Brokensha, Rouane, Landschoff, Jannes & Griffiths, Charles, 2023, New and notable stomatopods (Crustacea, Stomatopoda) from Mozambique, pp. 148-179 in European Journal of Taxonomy 900 (1)</i> on page 164, DOI: 10.5852/ejt.2023.900.2309, <a href="http://zenodo.org/record/10082989">http://zenodo.org/record/10082989</a&gt

Clorida undetermined

<i>Clorida</i> sp., undescribed species <p>Fig. 4</p> Diagnosis <p>A1 somite dorsal processes with short, triangular apices. A2 peduncle segment 1 extending anteriorly beyond eyes. Carapace with anterolateral spines. Raptorial claw dactylus with 5 teeth; outer proximal margin with basal notch. Mandibular palp 3-segmented. TS5 lateral process a short slender lobe, apex spiniform, with small ventral spine. TS6 lateral process broadly rounded. TS7 lateral process subtruncate; anterolateral and posterolateral angles obtusely rounded. TS7–8 and AS (1)2–5 with low SM carinae. AS 6 with distinct SM carinae. Abdominal carinae spined as follows: SM6, IM 5–6, LT 5–6, MG (4)5. Telson dorsolateral surface with widely-spaced rows of tubercles; margin of IM and LT teeth faintly crenulate to strongly tuberculate; denticles SM 4–6, IM 5–8, LT 1; ventral surface with long postanal carina, extending beyond half distance between anal pore and posterior margin. Uropodal protopod outer margin crenulate to serrate (smooth in one example); inner margin with 3–6 slender spines. Uropodal exopod proximal segment outer margin with 6–8 movable spines; distal segment longer than proximal segment; with black patch at articulation of exopod segments.</p> Material examined <p>MOZAMBIQUE • 1 ♂ (TL 64 mm), 2 ♀♀ (TL 47, 53 mm); Mozambique (exact location data unavailable); depth unknown; 1924; J.D.F. Gilchrist leg; SAMC – A006796.</p> <p> SOUTH AFRICA • 2 ♀♀ (TL 47, 53 mm); KwaZulu-Natal; 29°10.9′ S, 31°42.8′ E; 40 m depth; 25 May 1976; RV <i>Meiring Naude</i>; beam trawl; stn SM114; SAMC – A017939.</p> Measurements <p>Male (n = 1) TL 64 mm, female (n = 2) TL 47–53 mm.A1 peduncle 1.05–1.18CL. A2 scale 0.46–0.49CL.</p> Colour in alcohol <p>Mostly faded from dark-brown to pale amber. Uropodal exopod with darker patch at articulation of proximal and distal segments. Colour in life unknown.</p> Distribution and habitat <p>Recorded for the first time from Mozambique [unknown location] and South Africa.</p> Remarks <p> The Mozambican sample was collected a century ago and it is only after the recent taxonomic study of the ‘ <i>Clorida</i> complex’ by Ahyong (2000) that this species could be compared to the similar species <i>C. latreillei</i> Eydoux & Souleyet, 1842 and <i>C. albolitura</i> Ahyong & Naiyanetr, 2000. All specimens examined herein were previously identified as <i>C. latreillei</i> in the museum collection. The present specimens are most similar to <i>C. albolitura</i> described from the Gulf of Thailand (Ahyong & Naiyanetr 2000) and known from the Indian Ocean and most recently in the Mediterranean as an invasive species (Ahyong & Galil 2006). <i>Clorida albolitura</i> is morphologically distinguished from <i>C. latreillei</i> by the length of the postanal carina of the telson extending beyond half the distance between the anal pore and posterior margin and this characteristic is shared by the present specimens (Fig. 4G–H). But the degree of inflation in the adult male specimen is significantly greater than that exhibited by <i>C. albolitura</i>, and instead resembles that of an adult male of <i>C. latreillei</i>. Furthermore, an unusual feature of the present material is the crenulated or serrate outer margin of the uropodal protopod which is always smooth in <i>C. latreillei</i> and <i>C. albolitura.</i> This feature coupled with the degree of inflation of the telson teeth carinae as well as the long post-anal keel suggests that the specimens represent an undescribed species.</p>Published as part of <i>Brokensha, Rouane, Landschoff, Jannes & Griffiths, Charles, 2023, New and notable stomatopods (Crustacea, Stomatopoda) from Mozambique, pp. 148-179 in European Journal of Taxonomy 900 (1)</i> on pages 154-156, DOI: 10.5852/ejt.2023.900.2309, <a href="http://zenodo.org/record/10082989">http://zenodo.org/record/10082989</a&gt