Coexistence and relative abundance in annual plant assemblages: The roles of competition and colonization

Although an interspecific trade-off between competitive and colonizing ability can permit multispecies coexistence, whether this mechanism controls the structure of natural systems remains unresolved. We used models to evaluate the hypothesized importance of this trade-off for explaining coexistence and relative abundance patterns in annual plant assemblages. In a nonspatial model, empirically derived competition-colonization trade-offs related to seed mass were insufficient to generate coexistence. This was unchanged by spatial structure or interspecific variation in the fraction of seeds dispersing globally. These results differ from those of the more generalized competition-colonization models because the latter assume completely asymmetric competition, an assumption that appears unrealistic considering existing data for annual systems. When, for heuristic purposes, completely asymmetric competition was incorporated into our models, unlimited coexistence was possible. However, in the resulting abundance patterns, the best competitors/poorest colonizers were the most abundant, the opposite of that observed in natural systems. By contrast, these natural patterns were produced by competition-colonization models where environmental heterogeneity permitted species coexistence. Thus, despite the failure of the simple competition-colonization trade-off to explain coexistence in annual plant systems, this trade-off may be essential to explaining relative abundance patterns when other processes permit coexistence

Automatic tuning of hydrogen masers

Varying the density of the atoms in the cavity changes the Q of the atoms. When the cavity is mistuned, the density variation causes a frequency variation proportional to the degree of cavity mistuning

Effects of temporal variability on rare plant persistence in annual

Traditional conservation biology regards environmental fluctuations as detrimental to persistence, reducing long-term average growth rates and increasing the probability of extinction. By contrast, coexistence models from community ecology suggest that for species with dormancy, environmental fluctuations may be essential for persistence in competitive communities. We used models based on California grasslands to examine the influence of interannual fluctuations in the environment on the persistence of rare forbs competing with exotic grasses. Despite grasses and forbs independently possessing high fecundity in the same types of years, interspecific differences in germination biology and dormancy caused the rare forb to benefit from variation in the environment. Owing to the buildup of grass competitors, consecutive favorable years proved highly detrimental to forb persistence. Consequently, negative temporal autocorrelation, a low probability of a favorable year, and high variation in year quality all benefited the forb. In addition, the litter produced by grasses in a previously favorable year benefited forb persistence by inhibiting its germination into highly competitive grass environments. We conclude that contrary to conventional predictions of conservation and population biology, yearly fluctuations in climate may be essential for the persistence of rare species in invaded habitats

Development of a ferromagnetic rotary vacuum sealed spacecraft spin fixture

A number of successful spacecraft tests were conducted on an environmental spin fixture which utilizes a ferrofluidic rotary vacuum seal. The 27 cm (10.5 inch) diameter fixture drive shaft supports and spins communications satellites during flight acceptance testing in a thermal vacuum chamber. The drive shaft rotary seal serves to maintain the canned drive system electro-mechanical components at ambient pressure within the space simulator. The ferromagnetic fluid seal was chosen over conventional mechanical sealing devices for its zero-leakage, zero-wear, and minimum friction drag characteristics, as well as its high reliability potential

The radar cross section of dielectric disks

A solution is presented for the backscatter (nonstatic) radar cross section of dielectric disks of arbitrary shape, thickness and dielectric constant. The result is obtained by employing a Kirchhoff type approximation to obtain the fields inside the disk. The internal fields induce polarization and conduction currents from which the scattered fields and the radar cross section can be computed. The solution for the radar cross section obtained in this manner is shown to agree with known results in the special cases of normal incidence, thin disks and perfect conductivity. The solution can also be written as a product of the reflection coefficient of an identically oriented slab times the physical optics solution for the backscatter cross section of a perfectly conducting disk of the same shape. This result follows directly from the Kirchhoff type approximation without additional assumptions

Review of measurements of the RF spectrum of radiation from lightning

Measurements reported in the literature of the spectrum of electromagnetic radiation from lightning in the frequency range from 1 kHz to 1 GHz are reviewed. Measurements have been made either by monitoring the power received at individual frequencies using a narrow bandwidth recording device tuned to the frequencies under investigation or by recording the transient (time dependent) radiation with a wide bandwidth device and then Fourier transforming the waveform to obtain a spectrum. Measurements of the first type were made extensively in the 1950's and 1960's and several composite spectra have been deduced by normalizing the data of different investigators to common units of bandwidth and distance. The composite spectra tend to peak near 5 kHz and then decrease roughly as (frequency) to the -1, up to nearly 100 MHz where scatter in the data make the behavior uncertain. Measurements of the second type have been reported for return strokes, the stepped leader and for some intracloud processes. The spectrum of first return strokes obtained in this manner is very similar to the composite spectra obtained from the narrow-band measurements

Sources of the strongest RF radiation from lightning

Experiments performed at the Kennedy Space Center, Florida during TRIP-78 identified sources of the strongest RF radiation from lightning in the HF-VHF frequency range. Measurements were made of electric field changes associated with RF radiation using a field change system triggered on the output of an RF detector. The field changes associated with the strongest RF radiation are very fast (10 - 20 microseconds), bipolar pulses having an initial negative going half-cycle followed by a positive overshoot. These fast pulses consistently produced more RF radiation than was associated with return strokes, and their shape was remarkably consistent, independent of frequency

The temporal structure of RF radiation from lightning

Radiation from lightning in the RF band from 3-300 MHz has been examined. Simultaneous measurements were made of the RF radiation from lightning together with records of fast and slow field changes. Continuous analogue recordings were made with a system having 300 kHz of bandwidth in the RF channels. The temporal history of RF radiation of these frequencies consists of a sequence of discrete pulses. The data reveal a distinct pattern in the radiation which is independent of frequency and depends on the type of lightning flash: Cloud-to-ground flashes are characterized by an abrupt beginning associated with the stepped leader, whereas cloud-to-cloud flashes begin with a slower train of noise pulses more typical of the end of both types of flash. An exception to this pattern is cloud-to-ground flashes preceded by a breakdown phase, in which case the radiation begins like a cloud-to-cloud flash