Zmeny spoločenstiev bystruškovitých rôznych typov habitatov poľnohospodárskej krajiny v závislosti od vybraných environmentálnych faktorov vrátane pôdnych vlastností

The variations in ground beetles (Coleoptera: Carabidae) assemblages across the three types of farmland habitats, arable land, meadows and woody vegetation were studied in relation to vegetation cover structure, intensity of agrotechnical interventions and selected soil properties. Material was pitfall trapped in 2010 and 2011 on twelve sites of the agricultural landscape in the Prešov town and its near vicinity, Eastern Slovakia. A total of 14,763 ground beetle individuals were entrapped. Material collection resulted into 92 Carabidae species, with the following six species dominating: Poecilus cupreus, Pterostichus melanarius, Pseudoophonus rufipes, Brachinus crepitans, Anchomenus dorsalis and Poecilus versicolor. Studied habitats differed significantly in the number of entrapped individuals, activity abundance as well as representation of the carabids according to their habitat preferences and ability to fly. However, no significant distinction was observed in the diversity, evenness neither dominance. The most significant environmental variables affecting Carabidae assemblages species variability were soil moisture and herb layer 0-20 cm. Another best variables selected by the forward selection were intensity of agrotechnical interventions, humus content and shrub vegetation. The other from selected soil properties seem to have just secondary meaning for the adult carabids. Environmental variables have the strongest effect on the habitat specialists, whereas ground beetles without special requirements to the habitat quality seem to be affected by the studied environmental variables just little.Zmeny v spoločenstvách bystruškovitých (Coleoptera: Carabidae) troch typov habitatov poľnohospodárskej krajiny, t.j. ornej pôdy, trvalo trávnych porastov a mimolesnej krovinovej vegetácie boli sledované v závislosti od štruktúry vegetačnej pokrývky, intenzity agrotechnických zásahov a vybraných pôdnych vlastností. Materiál bol zbieraný metódou formalínových zemných pascí v rokoch 2010 a 2011 v rámci 12-tich stanovíšť poľnohospodárskej krajiny v urbánnej zóne mesta Prešov a jeho blízkeho okolia na východnom Slovensku. V rámci uvedeného zberu bolo odchytených celkovo 14 763 jedincov a determinovaných 92 druhov bystruškovitých. Dominantnými druhmi boli: Poecilus cupreus, Pterostichus melanarius, Pseudoophonus rufipes, Brachinus crepitans, Anchomenus dorsalis and Poecilus versicolor. Spoločenstvá sledovaných biotopov sa signifikantne líšili v počte odchytených jedincov, epigeickej aktivite ako aj zastúpení bystrušiek vo vzťahu k ich habitatovým preferenciám a letovým schopnostiam. Výskum však nepotvrdil signifikantné rozdiely v diverzite, ekvitabilite ani dominancii. Z nami sledovaných environmentálnych premenných kompozíciu spoločenstva bystruškovitých najvýznamnejšie ovplyvňuje pôdna vlhkosť a vegetačný kryt s výškou 0-20 cm. Ďalšími významnými premennými, ktoré vplývajú na kompozíciu spoločenstva sú intenzita agrotechnických zásahov, obsah pôdneho humusu a krovinová vegetácia. Ostatné zo sledovaných pôdnych vlastností majú na dospelé jedince bystrušiek len sekudárny vplyv. Rovnako, environmentálne premenné najvýraznejšie ovplyvňujú habitatových špecialistov, zatiaľ čo druhy bystrušiek bez vyhranených nárokov na podmienky prostredia sú ovplyvnené uvedenými faktormi len málo

Coat Polymorphism in Eurasian Lynx: Adaptation to Environment or Phylogeographic Legacy?

We studied the relationship between the variability and contemporary distribution of pelage phenotypes in one of most widely distributed felid species and an array of environmental and demographic conditions. We collected 672 photographic georeferenced records of the Eurasian lynx throughout Eurasia. We assigned each lynx coat to one of five phenotypes. Then we fitted the coat patterns to different environmental and anthropogenic variables, as well as the effective geographic distances from inferred glacial refugia. A majority of lynx were either of the large spotted (41.5%) or unspotted (uniform, 36.2%) phenotype. The remaining patterns (rosettes, small spots and pseudo-rosettes) were represented in 11.0%, 7.4%, and 3.9% of samples, respectively. Although various environmental variables greatly affected lynx distribution and habitat suitability, it was the effect of least-cost distances from locations of the inferred refugia during the Last Glacial Maximum that explained the distribution of lynx coat patterns the best. Whereas the occurrence of lynx phenotypes with large spots was explained by the proximity to refugia located in the Caucasus/Middle East, the uniform phenotype was associated with refugia in the Far East and Central Asia. Despite the widely accepted hypothesis of adaptive functionality of coat patterns in mammals and exceptionally high phenotypic polymorphism in Eurasian lynx, we did not find well-defined signs of habitat matching in the coat pattern of this species. Instead, we showed how the global patterns of morphological variability in this large mammal and its environmental adaptations may have been shaped by past climatic change.publishedVersio

Lomechusoides amurensis Wasmann 1897

Lomechusoides amurensis (Wasmann, 1897) (Figs. 14 –27, 49) Lomechusa amurensis Wasmann, 1897: 246 Lomechusa Ganglbaueri Bernhauer, 1936: 323. Type data: Ostsibirien: Irkutsk (im British Museum und in meiner eigenen Sammlung), Mongolei: Shangai (in der Sammlung des naturhistorischen Staatsmuseum in Wien), part, synonymy in Hlaváč et al., 2011: 14. Lomechusa ganglbaueri ssp. uralensis Bernhauer, 1936: 323. Type data: ein weiteres Exemplar vom Ural (Slatoust) [=Zlatousť], synonymy in Smetana, 2004: 457, synonymy confirmed. Type material studied: Lomechusoides amurensis Wasmann: HOLOTYPE, 13: (h) yellow parabolic label: Amur land / (h) Lomechusa n. sp. nec inflata Thoms. ?? L. sibirica Motsch. / (h) Lomechusa amurensis Wasm. n. sp. Type / round label with red margin: (p) Type / red label: (p) Holotypus, (h) Lomechusa amurensis Wasmann, W. Schillow des. 981 / (p) Sharp Coll 1905 - 323 / red label (p) HOLOTYPE 3 Lomechusoides amurensis (Wasmann), P. Hlaváč det. 2007. BMNH. Lomechusoides ganglbaueri uralensis Bernhauer: HOLOTYPE, 13: (h) Ural, Slatoust [=Zlatousť]/ (h) ganglbaueri Brh. (p) det. Bernh. / orange label (h) ssp. uralensis Bernh. Typus unic. / (p) Chicago NHMus, M. Bernhauer Collection / red label (p) Lomechusa ganglbaueri ssp. uralensis (Bernhauer), P. Hlaváč des., 2009 / Lomechusoides amurensis (Wasmann) = Lomechusa ganglbaueri ssp. uralensis (Bernhauer) P. H l av á č det., 2009. FMNH. Other material. 1 Ƥ: in Russian characters: Vost. [=eastern] Sayan 1500m, verkh [=top] Kyngarga, 18– 23.07. 95 / iz mkhov [from moss], Shavrin, A. / Lomechusa amurensis Wasm. Shavrin der. 97, in toto in euparal. CPHK. 1 3: in Russian characters: promysel Ozerpach liman Amura, 16 – 12.vi. 1915, Chernavin / L. amurensis W. Schilov det. 19. ZIN. 1 Ƥ: in Russian characters: Syutszukte, yu-b. [south-east], Kenteys. Z, Urgi, Kozlov, 1.vi. 925. ZIN. Redescription. Body lighter reddish-brown (Fig. 49); head and narrow basal margin of tergites darker, slightly shining. Head (Fig. 15) quadrate, longer than wide (HW/HL= 0.79), with deep median, frontal, triangular depression which is sparsely microsculptured, anterior part shining; length of eyes 0.26 times of length of head; temples slightly convergent posteriorly, posterior corners of temples rounded, posterior margin of head rounded, postoccipital proccess well-defined, protuberant. Antennae (Fig. 16) shorter, scape 1.35 times as long as wide and 1.95 times as long as pedicel, all antennomeres elongate except II which is as long as wide, III 1.10 and IV 1.27 as long as wide; antennomeres IV–X elongate, oval, terminal segment thin and acute, 1.15 times as long as scape, relative length of antennomeres from base to apex: 21: 10: 13: 14: 17: 16: 16: 17: 16: 16: 24. Pronotum (Fig. 17) unicoloured, with clearly convex anterior margin, sides rounded with basal corners obtuse and with deep lateral depressions, surface of this depressions with microsculpture, disc smooth, with sparse, uneven puncturation, basal median lobe large; pronotum in hind corners as wide as humeral width of elytra; ratio: PW/PL = 1.42. Metaventrite (Fig. 18 a) with metaventral process (Fig. 18 b) with anterior corners sharply angled, anterior margin straight, lateral margins convex, metaventral process in anterior part with lens-like depression bounded with rounded groove, surface unevenly punctured and with short setae, setae not reaching margin of next puncture, metaventral disc lacking microsculpture, unevenly punctured, in places lacking punctures. Elytra finely punctured, distance between punctures greater than puncture diameter, 1.15 times shorter than pronotum measured on median line, ratio: EW/EL=2.00. Abdomen with tergite II densely punctured and setaceous, distance between punctures on posterior margin greater than puncture diameter or even double of diameter of puncture, tergites III–IV with fine puncturation and setation on posterior margin, tergites V–VII sparsely but finely punctured at base, densely punctured on lateral parts, tergites II–VIII with visible microsculpture on lateral sides, tergite and sternite VIII as in Figs. 19 a, 19 b, 20 a, 20 b. Aedeagus as in Figs. 21, 23, 24, median lobe robust, basal bulb wide with well-developed distal crest and basal bridge. Internal structures weakly-defined, apical lobe tube-like, apical part sharp, with many pseudopores. Paramere (Fig. 22) with triangular paramerite, velum about as large as paramerite, apical lobe subparallel, hookshaped bent backwards, apex with about four setae. Spermatheca as in Figs. 25–27, basal part V-shaped, apical part as long as basal part, sligthly broader and 2.5 –3.0 times as long as wide, internal wall in apical part roughly wrinkled. Measurements. (Holotype), body length 4.85 mm; forebody length 2.40 mm; head length 0.95 mm; head width 0.75 mm; antennal length 2.60 mm; pronotal length 1.10 mm; pronotal width 1.57 mm; elytral length on sutura 0.85 mm; elytral width 1.70 mm; foretibial length 1.00 mm; midtibial length 1.20 mm; hindtibial length 1.50 mm. Differential diagnosis. L. amurensis can be distinguished from other species of the group by the characteristic shape of the metaventral process with deep median depression and by the shape of spermatheca. Host ant. Unknown Distribution. Russia (Ural, Siberia and Far East) and Mongolia Lomechusoides schneideri Maruyama et Hlavá č, 2004 (Figs. 28 –31, 50) Lomechusoides schneideri Maruyama & Hlaváč, 2004: 110. Type data: China m., 19 –VII– 1992, Sichuan, 3,600 m, Kangding env. Type material studied. HOLOTYPE, 1 Ƥ: (p) China m., 19.7. 1992, Sichuan, 3600m, Ken ding env., J. Schneider legit. / (p) ex coll. M. Dvořák, National Museum Prague, Czech Republik/ red label (p) Holotype, Lomechusoides schneideri des. Maruyama & Hlaváč, 2004. Supplementary description. The species was recently adequately described by Maruyama & Hlaváč (2004: 110). Head (Fig. 28), pedicel as long as wide, antennomere III about 1.4 times as long as wide, both equal in length, relative length of antennomeres from base to apex: 22: 11: 12: 14: 15: 16: 15: 15: 15: 14: 20. Pronotum (Fig. 29) with sides and medium line with dense microsculpture, dull, pronotal disc near medium line with fine microsculpture, glossy. Pronotum in hind corners wider than humeral width of elytra. Metaventrite with metaventral process with anterior corners sharply angled, anterior margin slightly convex, lateral margins convex (Fig. 30), metaventral process in anterior part lacking shalow depression and rounded groove, whole surface of metaventrite unevenly but densely punctured and with short setae, setae reaching margin of next puncture, metaventral disc with sparse microsculpture. Basal part of elytra densely punctured, distance betwen puntures shorter than diameter of puncture, densely and markedly microsculptured, dull, rest of surface of elytra coarsely punctured, lacking microsculpture, glosy. Abdomen with tergite II densely punctured and setaceous, distance between punctures on posterior margin as long or slightly longer than puncture diameter, posterior margin of tergites III–IV with fine, sparse puncturation and setation, tergite IV much sparsely punctured than III, tergites V–VI sparsely, finely and evenly punctured, tergites II–VI in basal part with fine microsculpture, tergite and sternite VIII as in Figs 31 a, 31 b. Diferencial diagnosis. L. schneideri can be easily distinguished from the other species of the L. amurensis group by the long and quadrate head and long, subparallel temples. Host ant. Myrmica cf. ruginodis Nylander, 1846 Distribution. China (Sichuan)Published as part of Jászay, Tomáš & Hlaváč, Peter, 2013, A taxonomic revision of the myrmecophilous genus Lomechusoides Tottenham, 1939 (Coleoptera: Staphylinidae: Aleocharinae) Part I. Redescription of the genus, definition of species groups and the revision of the amurensis Wasmann 1897 species group, pp. 65-81 in Zootaxa 3683 (1) on pages 71-75, DOI: 10.11646/zootaxa.3683.1.4, http://zenodo.org/record/28413

Lomechusoides Tottenham 1939

Genus Lomechusoides Tottenham, 1939 Lomechusoides Tottenham, 1939: 226. Type species: Staphylinus strumosus Fabricius, 1793, original designation Goniodes Stephens, 1829: 260. Type species: Staphylinus strumosus Fabricius, 1793, virtual monotypy (attributed to Kirby; preoccupied, not Nitzsch, 1818) Lomechusa Gravenhorst: Erichson, 1839 (redescription); Ganglbauer, 1895 (redescription); Wasmann, 1897 (revision, key); Wasmann, 1888 (biology); Feynes, 1920: 304 (redescription, catalogue, distribution); Bernhauer, 1936 (key to known species) Lomechusoides Tottenham: Maruyama & Hlaváč, 2004 (catalogue, new combinations, synonyms) Diagnosis. The genus Lomechusoides can be readily separated from between three genera of the subtribe Lomechusina by the combination of following characters: 1) antennae and legs stout, 2) pronotum with lateral part dull, finely microsculptured and 3) lacinia without hook or spine. Redescription. Body yellowish-brown (Fig. 48), pronotum partly darker, elytra and tergites lighter, some tergites with basal part darker. Head: (Fig. 1) quadrate, longer than wide, or as long as wide, eyes weakly protuberant, shorter than temples, temples slightly convergent posteriorly or parallel, surface with sparse puncturation and setation, with microsculpture, matt with frontal triangular depression between well developed antennal protuberances. Labrum bilobed, wider than long, with some setae. Mandibles (Fig. 2) symmetric, angularly bent, lacking internal teeth, basal dorsal part in middle with numerous dentiform protuberances, lateral edge with dense setation. Maxillae (Fig. 3) with galea longer than lacinia, both of about same width, galea evenly expanded in basal part, in apical part narrowed, palpifer and stipes with setae, maxillary palpi 4 -segmented, palpomere I small, subquadrate, II expanded apically, widest, III slightly shorter and slender, IV as long as I, needle-like. Labium (Fig. 4) with prementum slighty longer than wide in basal part, anterior part with numerous setae, basal part with numerous pseudopores, glossy, indistinctly divided in two rounded lobes, lacking sensillae, labial palpi with palpomere I barrel-like, 1.5 as long as wide, about three times as long as II, palpomere II shortest, trapezoidal, wider than long, palpomere III narrowest, subparallel, three times as long as wide, as long as II, palpomere I and II with setae, III with some pseudopores. Mentum (Fig. 5) rhomboidal, 1.3 times as wide as long, widest at base, narrowed anteriorly, anterior margin deeply excavated, in lateral anterior and posterior corners with about seven pseudopores, in middle with about four pseudopores. Gular suture widely separated, closest in middle, suddenly narrowed anteriorly and evenly narrowed posteriorly. Antennae (Fig. 6) long, when bent backwards usually reaching or exceeding base of elytra, all antennomeres more or less asymmetric, scape large, larger than all other antenommeres individually, pedicel smallest, II–III rhomboid, both slightly pedunculate, IV–X barrel-like shaped, terminal antennomere pointed apically, longest, all antennomeres matt, with setation, antennomeres III–IV in males with bunch of lateral setae (absent in L. suensoni). Pronotum (Figs. 7, 8) variable in shape and size, transverse, widest at base, anterior angles rounded, posterior angles well-defined, sharp, surface sometimes with tubercles, basal margin with strong median lobe, surface with various depressions, pronotal disc with uneven puncturation, with or without microsculpture, hypomeron large and matt. Mesoventrite wide, five times as wide as mid-line length, with short and fine setae, surface with microsculpture, posterior mesoventral process truncated. Metaventrite (Fig. 9) about four times as long as mesoventrite, surface unevenly punctured, setaceous, with or without microsculpture, macrosetation species-characteristic, shape of metaventral process species-characteristic, with or without depression. Elytra as long as or shorter than pronotum, elytron about as long as wide or slightly shorter, finely punctured with setation, in basal and humeral part with macrosetae, posterior margin of elytra in front of posterior corner excavated, angle of sutura rounded, wings well-developed. Abdomen robust, evenly expanded, tergite V widest, from tergite V slightly convergent posteriorly, abdomen longer than head, pronotum and elytra combined. Paratergites III–V with trichomes, lateral margins of tergites II– IV, VIII–X as well as lateral margins of sternites II–IV and metepimeron with dense golden setae, tergites V–VII in posterior corners with short setae, surface with or without weak microsculpture, sternites evenly punctured with erect macrosetae. Legs long, with setation, apical part of femora with bunch of long setae (Fig. 10) (groups L. amurensis and L. strumosus) or lacking any long setae (groups L. minor and L. straneoi), tarsal formula 4-5 - 5, trochanters similarly shaped, semi-triangular, procoxae about three times as long as wide, mesocoxae 1.3 times as long as wide and metacoxae about as long as wide, tibiae slightly shorter than femora, slightly convergent posteriorly, coxae, trochanters and ventral part of femora with few macrosetae. Sexual dimorphism. setae on antenommeres III–IV longer in males (Fig. 6), tergites and sternites VIII differently shaped in males and females (Figs. 11 a, 11 b, 12 a, 12 b). Distribution. Almost the whole Palaearctic region, from Spain to Siberia and Russian Far East, China and Japan. Biology. Lomechusoides is a strictly myrmecophilous genus with symphilic way of life which so far has been recorded only with ants of the genus Formica Linnaeus, 1758 and Myrmica Latreille, 1804. The following ants have been found as hosts of Lomechusoides so far: Formica cunicularia Latreille, 1798; F. f u s c a Linnaeus, 1758, F. pratensis Retzius, 1783; F. r u f a Linnaeus, 1761; F. rufibarbis Fabricius, 1793; F. sanguinea Latreille, 1798; Myrmica rubra Linnaeus, 1758 and M. scabrinodis Nylander, 1846.Published as part of Jászay, Tomáš & Hlaváč, Peter, 2013, A taxonomic revision of the myrmecophilous genus Lomechusoides Tottenham, 1939 (Coleoptera: Staphylinidae: Aleocharinae) Part I. Redescription of the genus, definition of species groups and the revision of the amurensis Wasmann 1897 species group, pp. 65-81 in Zootaxa 3683 (1) on pages 66-67, DOI: 10.11646/zootaxa.3683.1.4, http://zenodo.org/record/28413

Lomechusoides amurensis Jászay & Hlaváč, 2013, sp. nov.

The amurensis species group Head narrow, longer than wide, eyes short, shorter than half and longer than one-third of temples, frontal, triangular depression sparsely microsculptured or microsculpture lacking, with equally dense setation. Antennomere III slightly longer than pedicel and shorter or at most as long as IV, all antennomeres matt, with dense microsculpture. Pronotum trapezoidal with straight anterior margin, in anterior third with well-defined medial impression and welldefined microsculpture, median groove weak, disc with tubercules bearing long setae and with variable microsculpture. Metaventral process with lens-like depression, better or weaker defined, surface with puncturation, median line lacking punctures, posterior margin of disc with punctures, only anterior part with microsculpture, lacking macrosetae. Tergites III–IV on posterior lateral margin finely punctured and with fine macrosetae (8–10), tergites V–VIII with sparse but coarse punctures, especially VI, surface of tergite VII with uneven microsculpture, tergite III–V with macrosetae, VI–VII lacking macrosetae. Sternites with short setae, only on sternite III exceeding posterior margin of sternite, on other sternites setae not exceeding its posterior margin. List of species. L. amurensis (Wasmann), L. kozlovi sp. nov., L. schneideri Maruyama & Hlaváč, L. suensoni (Bernhauer). Distribution: Eastern Palaearctic region, from Ural to China, Russian Far East and Japan.Published as part of Jászay, Tomáš & Hlaváč, Peter, 2013, A taxonomic revision of the myrmecophilous genus Lomechusoides Tottenham, 1939 (Coleoptera: Staphylinidae: Aleocharinae) Part I. Redescription of the genus, definition of species groups and the revision of the amurensis Wasmann 1897 species group, pp. 65-81 in Zootaxa 3683 (1) on page 70, DOI: 10.11646/zootaxa.3683.1.4, http://zenodo.org/record/28413

Lomechusoides kozlovi Jászay & Hlaváč, 2013, sp. n.

Lomechusoides kozlovi sp. n. (Figs. 42 –47, 52) Etymology. The species is named after Russian explorer of Mongolia and Tibet, Pjotr Kuzmič Kozlov (1863– 1935), the collector of the species. Type material. HOLOTYPE, 13: in Russian characters: (p) „р. Сэрг-чю 13800´б. Желтой, Тибетъ Козловъ кон. v 01 ” [river Serg-čju, 13 800 feet (= 4 140 m), right confluent of Yellow river (= Chuang-che)], Tibet, Kozlov, V.01] / (h) Lomechusa prope amurensis (? sp.n?) W. Schilow, 981 / red label (p) Holotype Lomechusoides kozlovi sp. nov. T. Jászay & P. Hlaváč des. 2012. ZIN. Description. Body lighter reddish-brown (Fig. 52), head, pronotum and wide basal margin of tergites darker. Head quadrate, only slightly longer than wide (HW/HL= 0.94), with deep median, densely microsculptured frontal triangular depression, anterior part dull, length of eyes 0.32 that of head; temples slightly convergent, postoccipital process large (Fig. 42). Antennae (Fig. 43) shorter, scape 1.63 times as long as wide and 2.21 times as long as pedicel, all antennomeres elongate except transverse pedicel, antennomere III 1.45 and IV 1.40 as long as wide; antennomeres V–X elongate, oval, terminal antennomere thin and acute, about as long as scape, relative length of antennomeres from base to apex: 20: 10: 11: 13: 14: 14: 14: 14: 14: 14: 21. Pronotum (Fig. 44) bicoloured, pronotal disc clearly darker, with clearly straight anterior margin, sides straight, extended posteriorly, with lateral deep depressions, surface of these depressions with microsculpture, disc dull, with dense, uneven puncturation, basal median lobe broadly rounded, pronotum in hind corners narrower than humeral width of elytra; ratio: PW/PL = 1.36. Metaventrite with metaventral process with anterior corners rectangular, anterior margin straight, lateral margins subparallel (Fig. 45), metaventral process in anterior part with weakly defined lens-like depression but lacking rounded groove, whole surface of metaventrite unevenly punctured and setaceous, setae reaching margin of next puncture, metaventral disc with dense microsculpture. Elytra densely punctured, distance between punctures less than puncture diameter, 1.20 times shorter than pronotum measured along median line, ratio: EW/EL= 1.78. Abdomen with tergite II densely punctured and setaceous, distance between punctures on posterior margin less than puncture diameter, tergites III–IV on posterior margin with fine, sparse puncturation and setation, tergites V– VII sparsely but roughly punctured at base, densely punctured on lateral parts, tergites II–VIII with uneven microsculpture, only tergites V–VI with microsculpture on lateral sides, tergite and sternite VIII as in Fig. 46 a, 46 b. Aedeagus unknown. Spermatheca as in Fig. 47, basal part U-shaped, apical part slightly shorter and about as wide as basal part, about three times as long as wide, internal wall in apical part sparsely wrinkled. Measurements. (Holotype), body length 4.70 mm; forebody length 2.45 mm; head length 0.79 mm; head width 0.75 mm; antennal length 2.40 mm; pronotal length 1.10 mm; pronotal width 1.50 mm; elytral length 0.95 mm; elytral width 1.70 mm; foretibial length 0.95 mm; midtibial length 1.15 mm; hindtibial length 1.40 mm Diferencial diagnosis. L. kozlovi differs from other species of the amurensis group by its deep median triangular depresssion on head, metavenral process rectangular, elytra densely and coarsely punctured and by the characteristic shape of spermatheca. Host ant. Unknown Distribution. China (Tibet)Published as part of Jászay, Tomáš & Hlaváč, Peter, 2013, A taxonomic revision of the myrmecophilous genus Lomechusoides Tottenham, 1939 (Coleoptera: Staphylinidae: Aleocharinae) Part I. Redescription of the genus, definition of species groups and the revision of the amurensis Wasmann 1897 species group, pp. 65-81 in Zootaxa 3683 (1) on pages 77-81, DOI: 10.11646/zootaxa.3683.1.4, http://zenodo.org/record/28413

A taxonomic revision of the myrmecophilous genus Lomechusoides Tottenham, 1939 (Coleoptera: Staphylinidae: Aleocharinae) Part I. Redescription of the genus, definition of species groups and the revision of the amurensis Wasmann 1897 species group

Jászay, Tomáš, Hlaváč, Peter (2013): A taxonomic revision of the myrmecophilous genus Lomechusoides Tottenham, 1939 (Coleoptera: Staphylinidae: Aleocharinae) Part I. Redescription of the genus, definition of species groups and the revision of the amurensis Wasmann 1897 species group. Zootaxa 3683 (1): 65-81, DOI: 10.11646/zootaxa.3683.1.

Lomechusoides

Key to the species groups of the genus Lomechusoides 1 Apical half of femora with a bunch of dense and long setae (Fig. 10)............................................ 2 - Apical half of femora with normal setation (setae not longer than one fourth of width of femora) or in whole glabrous, lacking any setation.......................................................................................... 3 2 Setae on posterior margin of elytra directed towards posterior corners of elytra (Fig. 13), pronotum more rectangular, anterior margin with uneven microsculpture.................................................. species group L. strumosus - Setae on posterior margin of elytra in internal, sutural half directed against each other and in external half directed towards posterior corners of elytra (Fig. 14), pronotum trapezoidal with broadly rounded anterior corners, pronotum in anterior fifth matt, with strong microsculpture..................................................... species group L. amurensis 3 Setae on posterior margin of elytra directed towards posterior corners of elytra, pronotum and elytra lacking macrosetae..................................................................................... species group L. straneoi - Setae on posterior margin of elytra in internal, sutural half directed against each other and in external half directed towards posterior corners of elytra, pronotum and elytra with macrosetae...............................species group L. minorPublished as part of Jászay, Tomáš & Hlaváč, Peter, 2013, A taxonomic revision of the myrmecophilous genus Lomechusoides Tottenham, 1939 (Coleoptera: Staphylinidae: Aleocharinae) Part I. Redescription of the genus, definition of species groups and the revision of the amurensis Wasmann 1897 species group, pp. 65-81 in Zootaxa 3683 (1) on page 69, DOI: 10.11646/zootaxa.3683.1.4, http://zenodo.org/record/28413

Staphylinus strumosus

The strumosus species group Head wide, about as wide as long, eyes slightly longer than temples, frontal, triangular depression with sparser puncturation and setation as on rest of head. Antennomere III longer than pedicel and longer or at most as long as IV, all antennomeres matt, with dense microsculpture. Pronotum strongly transverse, more rectangular than trapezoidal in shape, with excavation on anterior margin, lateral margin matt, with tubercles and setae and with well-defined microsculpture, medial groove well-defined, pronotal disc with tubercles with erect setae, microsculpture variable, with setation. Metaventral process lacking depression, punctures with long setae, medial groove punctured and setose, surface with microsculpture and macrosetae. Tergites III–IV finely punctured on posterior margin with fine setae and macrosetae (14–18), tergite V–VIII lacking puncturation, surface with uneven microsculpture and setae. Sternites with long setae, on sternite III–IV exceeding posterior margin of sternite, on other sternites setae not exceeding its posterior margin. List of species. L. inflatus (Zetterstedt), L. mongolicus (Wasmann), L. sibiricus (Motschulsky), L. strumosus siculus (Fiori), L. strumosus strumosus (Fabricius), L. teres (Eppelsheim). Distribution. Almost the whole Palaearctic region, from Spain, central and northern Europe, Caucasus, Mongolia, China and Russia Far East.Published as part of Jászay, Tomáš & Hlaváč, Peter, 2013, A taxonomic revision of the myrmecophilous genus Lomechusoides Tottenham, 1939 (Coleoptera: Staphylinidae: Aleocharinae) Part I. Redescription of the genus, definition of species groups and the revision of the amurensis Wasmann 1897 species group, pp. 65-81 in Zootaxa 3683 (1) on page 70, DOI: 10.11646/zootaxa.3683.1.4, http://zenodo.org/record/28413

Lomechusoides amurensis

Key to the species of the amurensis species group 1 Head long, widest just before posterior margin, temples much longer than eyes, eyes small, 0.22 of length of head.................................................................................................... L. schneideri - Head shorter, widest accross eyes, temples longer than eyes, eyes larger, at least 0.30 of length of head.................. 2 2 Pronotum in posterior corners narrower than elytra, elytra coarsely and densely punctured.................... L. kozlovi - Pronotum in posterior corners as wide than elytra, elytra finely and sparsely punctured............................... 3 3 Antenomere II as long as wide, apical part of spermatheca long (Fig. 25), about as wide as basal part, 2.5 –3.0 times as long as wide...................................................................................... L. amurensis - Antenomere II wider than long, apical part of spermatheca short (Fig. 33), about 1.5 times as wide as basal part, about 2.0 times as long as wide......................................................................... L. suensoniPublished as part of Jászay, Tomáš & Hlaváč, Peter, 2013, A taxonomic revision of the myrmecophilous genus Lomechusoides Tottenham, 1939 (Coleoptera: Staphylinidae: Aleocharinae) Part I. Redescription of the genus, definition of species groups and the revision of the amurensis Wasmann 1897 species group, pp. 65-81 in Zootaxa 3683 (1) on page 71, DOI: 10.11646/zootaxa.3683.1.4, http://zenodo.org/record/28413