Revision of the genus Holocephalus (Coleoptera: Scarabaeidae: Scarabaeinae: Coprini)

The genus Holocephalus Hope is revised and now contains five species: H. cristatus (Gillet), H. eridanus (Olivier), H. julieni sp.nov. H. sculptus (Gillet), and H. simoni sp.nov. All taxa are described and illustrated. A key to speies of Holocephalus is included. Lectotypes are designated for H. eridanus and H. sculptus. Scarabaeus eridanus Olivier is designated as the type species for the generic name Atrichius Gillet (a junior synonym of Holocephalus). On révise le genre Holocephalus Hope qui regroupe maintenant cinq es-pèces, soit H. cristatus (Gillet), H. eridanus (Olivier), H. julieni sp.nov., H. sculptus (Gillet) et H. simoni sp.nov. Outre les descriptions et illustrations pour chaque espèces on propose un tableau de détermination pour les espèces maintenant comprises dans le genre Holocephalus. On désigne des lectypes pour H. eridanus et H. sculptus. Enfin, on désigne l’espèce Scarabaeus eridanus Olivier comme type du genre Atrichius Gillet (considéré comme synonymec junior du genre Holocephalus)

A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini)

Génier, François (2012): A new species and notes on the subgenus Deltochilum (Deltochilum) Eschscholtz, 1822 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 3357: 25-36, DOI: 10.5281/zenodo.21364

Ateuchus cujuchi n. sp., a new inquiline species of Scarabaeinae (Coleoptera: Scarabaeidae) from tuco-tuco burrows in Bolivia

Génier, François (2015): Ateuchus cujuchi n. sp., a new inquiline species of Scarabaeinae (Coleoptera: Scarabaeidae) from tuco-tuco burrows in Bolivia. Zootaxa 3946 (1): 146-148, DOI: http://dx.doi.org/10.11646/zootaxa.3946.1.

FIGURE 1 – 7. Ateuchus cujuchi n in Ateuchus cujuchi n. sp., a new inquiline species of Scarabaeinae (Coleoptera: Scarabaeidae) from tuco-tuco burrows in Bolivia

FIGURE 1 – 7. Ateuchus cujuchi n. sp. (Figs. 1 – 6. holotype; Fig. 7, paratype). 1. habitus, dorsal view; 2. habitus, ventral view; 3. habitus, lateral view; 4. head, dorsal view; 5. pygidium, frontal view; 6. aedeagus with internal sac in situ, lateral view; 7. internal sac extracted

Digitonthophagus bonasus

<i>Digitonthophagus bonasus</i> species group <p> 1. <i>Digitonthophagus bonasus</i> (Fabricius, 1775), <b>valid species</b> Distribution: Bangladesh, Cambodia, India, Laos, Malaysia, Myanmar, Nepal, Pakistan, Sri Lanka, Thailand, Vietnam</p> <p> 2. <i>Digitonthophagus uks</i> Génier, <b>new species</b></p> <p>Distribution: Afghanistan, India, Iran, Pakistan</p> <p> 3. <i>Digitonthophagus sahelicus</i> Moretto, <b>new species</b></p> <p>Distribution: Burkina Faso, Cameroon, Chad, Djibouti, Eritrea, Mali, Mauritania, Niger, Senegal, Sudan</p>Published as part of <i>Génier, François & Moretto, Philippe, 2017, Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae), pp. 1-110 in Zootaxa 4248 (1)</i> on page 74, DOI: <a href="http://zenodo.org/record/439444">10.5281/zenodo.439444</a&gt

Catharsius camillus Harold 1877

Catharsius camillus Harold, 1877 (Figs. 13–17) Catharsius vitulus Boheman, 1857: 223 Catharsius camillus Harold, 1877: 97 Type locality. Cap b. sp. (South Africa) As for C. pandion, the syntypic series consist of seven specimens (4 ♂♂, 3 ♀♀). In order to provide a primary type with the most diagnostic characters a large male is selected from the syntypic series as lectotype (present designation). This specimen bears the following labels (Fig. 15): [Krebs!] handwriting on green card; [Camillus / Harold/ Cap b. sp.] handwriting on green card; [9352] white paper; [Type] red card; [SYNTYPUS / Catharsius / camillus Harold, 1877 / labelled by MNHUB 2014] red card; [LECTOTYPE ♂/ Catharsius / camillus / Harold, 1877 / dés. F. Génier, 2014] red card. Remark. The male lectotype has been dissected and the aedeagus extracted and figured (Fig. 16–17). Catharsius camillus is currently a junior subjective synonym of Catharsius vitulus Boheman, 1857. It should be pointed out that Harold’s original description state Port Natal (=Durban) as the type locality which conflict with the original specimen labels. Because C. camillus is synonymous with C. vitulus, we believe that the proper type locality would be most likely Cap. b. sp., in its former sense which included all of western South Africa. Although inaccurate it would more or less coincide with the southern distribution range of C. vitulus (A. Davis pers. comm.). All 6 paralectotypes bear a “Capland” printed label further supporting this interpretation.Published as part of Génier, François & Josso, Jean-François, 2016, Notes on Afrotropical Catharsius species described by Edgar von Harold (Coleoptera: Scarabaeidae, Scarabaeinae), pp. 333-342 in Zootaxa 4072 (3) on pages 335-338, DOI: 10.11646/zootaxa.4072.3.3, http://zenodo.org/record/26181

Catharsius harpagus Harold 1877

Catharsius harpagus Harold, 1877 (Figs. 8–12) Catharsius harpagus Harold, 1877: 97 Type locality. Delagoa Bay (Mozambique) Three specimens are in the collection. A male and a female are from recent collecting (1963) and have been seen by M.C. Ferreira. The remaining specimen is a syntype. Harold’s description is clear in stating a length range of “ 22–30 mill.” in the original description excluding the possibility of a unique specimen studied. Harold also stated “Specimina typica in Museo Berolinensi asservantur” for the type deposition. It is possible that the remaining specimens studied by Harold are deposited in different collection(s). Fortunately, the syntype studied is a well-developed male. This specimens is designated as lectotype (present designation) and bears the following labels (Fig. 10): [Delagoa/ Bay] white paper; [72066] white paper; [Type] red card; [D. O. Africa/ 36] red handwriting on white paper; [O.-Afrika/ Delagoa boy/ Felsche] handwriting on blue paper; [SYNTYPUS / Catharsius / harpagus Harold, 1877 / labelled by MNHUB 2014] red card; [LECTOTYPE ♂/ Catharsius / harpagus / Harold, 1877 / dés. F. Génier, 2014] red card. Remark. The male lectotype has been dissected and the aedeagus extracted (Figs. 11–12). This species is currently valid with no synonyms. No change in the taxonomic status of this species is required.Published as part of Génier, François & Josso, Jean-François, 2016, Notes on Afrotropical Catharsius species described by Edgar von Harold (Coleoptera: Scarabaeidae, Scarabaeinae), pp. 333-342 in Zootaxa 4072 (3) on page 335, DOI: 10.11646/zootaxa.4072.3.3, http://zenodo.org/record/26181