Aspects of precommercial thinning in heterogeneous forests in southern Sweden

The overall objective of the work underlying this thesis was to suggest and evaluate possible strategies for the tending of young heterogeneous stands of Norway spruce, Scots pine and birch in southern Sweden. Heterogeneity was defined as variation in species composition, height distribution and spatial arrangement of the trees. The influence of stand density after precommercial thinning and timing of thinning on the diameter of the thickest branch was studied for naturally regenerated Scots pine. The branch diameter was found to decrease with increasing number of remaining stems after precommercial thinning. However, leaving very dense stands (> 3000 stems ha-1) resulted only in a minor reduction of the branch diameter. Late precommercial thinning, compared to early, reduced the branch diameter. The influence of the precommercial thinning regime on the crown ratio (living crown length/tree height) was also analysed. To be able to simulate the influence of different management options on the development of the young forest, single-tree growth models was developed for Scots pine, Norway spruce and birch. Height growth and diameter was estimated as a function of tree height, stand and site variables. Growth reduction due to competition was estimated using individual, distance independent indices as well as expressions of the overall stand density. In the third study the influence of stand structure after precommercial thinning on the development of mixtures between Norway spruce and silver birch was simulated. The aim was to identify mixtures that allowed both species to develop well until the first commercial thinning. By leaving birches with an average height slightly greater than spruce at precommercial thinning, a large proportion of competitive birches were available at first commercial thinning, at the same time as the relative diameter distribution of spruce in the mixture was equal to that of a pure spruce stand of the same density. The height difference between the species as well as the species proportion had a decisive impact on volume production. In the fourth study different precommercial thinning strategies were identified and applied to a heterogeneous stand including Scots pine, Norway spruce and birch. Stand development and economical returns over a rotation was estimated using a set of empirical models. The aim of the long-term strategies was: (i) a conifer dominated stand with focus on high production, (ii) a conifer dominated stand with focus on high timber quality, (iii) to preserve the heterogeneous stand structure, (iv) a mosaic pattern by tree species, (v) to reduce the precommercial thinning cost, without jeopardizing the future stand development. The difference in total volume production was found to be relatively small between the strategies. The lowest production was found for the strategies promoting species mixture at tree level (iii) and group level (iv). The net present value was highest for the strategy aiming at high production (ii) and lowest for the strategy aiming at preserved heterogeneity (iii). The minimal precommercial thinning (v) was a less profitable alternative, mainly because of an expensive first commercial thinning. Differences in timber quality were not considered in the simulations. The case study illustrates the possibilities for influencing the structure of a heterogeneous stand through precommercial thinning, as well as the limitations imposed by the initial stand structure

Select or Not? Comparing the Impact of Selective and Schematic Thinning on Scots Pine Tree Growth and Stand Structure

The first commercial thinning usually entails a high cost in harvest operations and a low resulting income. From a practical forestry perspective, a schematic spatial selection might be more efficient than a selective approach. Therefore, this study aimed to compare basal area, total and standing volumes, and periodic annual increment (PAI), as well as stand structure, between different thinning designs (selective and schematic thinning) and strategies (thinning once or more than once) over a long-term monitoring period of Scots pine (Pinus sylvestris) plantations in Sweden. We also evaluated the relevancy of distance-dependent competition indices (CIs) in individual tree growth models by comparing growth model predictions with the use of distance-dependent and distance-independent CIs. Despite higher heterogeneity in schematically thinned stands, there were no significant differences in standing and total volumes (m(3)& BULL;ha(-1)) among treatments in the short or long term. Although the inclusion of a distance-dependent CI improved the model slightly, distance-independent models predicted diameter growth just as effectively. Schematic thinning could be a viable option for a first commercial thinning or one-time thinning if, at least, one more thinning is included in the management plan, or if the motivating interest is mainly volume

Pre-Commercial Thinning Increases the Profitability of Norway Spruce Monoculture and Supports Norway Spruce-Birch Mixture over Full Rotations

Pre-commercial thinning (PCT) is a common measure in Norway spruce (Picea abies L. Karst.) stands but the profitability of doing PCT and timing of PCT has not been fully investigated over a full rotation. Further, limited knowledge is available for mixed forest management compared to monocultures. In this study, different PCT strategies were tested to investigate the effect of PCT and timing of PCT on the production and profitability of Norway spruce monocultures and mixed Norway spruce-birch stands. A forest decision support system was used to simulate stand development during the whole rotation. Our study findings show that there is a positive effect of PCT on Norway spruce plantations' long-term profitability but that the timing of PCT has little effect on profitability. However, site variation might influence the effect of PCT timing on the profitability of Norway spruce stands. Moreover, we also found that retaining 1000 Norway spruce ha(-1) and 1000 birch ha(-1) after PCT supports a mixture of Norway spruce and birch over a full rotation, with little or no economic loss compared to pure Norway spruce stands. Therefore, such a mixture can lead to profitable production while still providing other ecosystem services

Multivariate mixed-effects models for stand characteristics of hybrid aspen plantations in southern Finland and southern Sweden

Hybrid aspen, a hybrid between the European aspen and North American trembling aspen (Populus tremula L. × P. tremuloides Michx.), is a promising species because of its fast-growth and its suitability for multi-purpose use. However, models for predicting the age-dependent development of stand characteristics are still missing. The main objectives of this study were therefore to develop the models for predicting stand characteristics of hybrid aspen plantations and to validate the model applicability. The target response variables were stand basal area (BA), basal area-weighted mean diameter (DG) and basal area-weighted mean height (HG). Data were obtained from clonal hybrid aspen trials in southern Finland and southern Sweden. Multivariate mixed-effects modelling was used to estimate the parameters of seemingly unrelated regression for BA, DG, and HG. Model fit provided the following predictor variables: stand age (AGE), the number of trees per hectare (TPH), site index (SI), growing degree-days (GDD5), soil and site type, and thinning treatment. The chosen predictors differed slightly by response variable, but all parameters were highly significant (P < 0.0001), and model goodness-of-fit statistics presented high accuracy: RMSE of 2.59 m2 ha−1 for BA, 1.21 cm for DG, 1.05 m for HG in arithmetic scale. The applied simulations illustrated clear differences in the predicted development of stand characteristics when input variables SI, TPH or GDD5 changed. The developed models were assessed to be easily applicable and useful for predicting the stand and tree characteristics of clonal hybrid aspen plantations, especially for the stands with AGE ≤ 30 years and TPH ≤ 2000 trees ha−1

Production of genetically improved silver birch plantations in southern and central Sweden

Investing in planting genetically unproved silver birch (Betula pendula Roth) in Swedish plantations requires understanding how birch stands will develop over their entire rotation. Previous studies have indicated relatively low production of birch compared to Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.). This could result from using unrepresentative basic data, collected from unimproved, naturally-regenerated birch (Betula spp.) growing on inventory plots often located in coniferous stands. The objective of this study was to develop a basal area development function of improved silver birch and evaluate production over a full rotation period. We used data from 52 experiments including planted silver birch of different genetic breeding levels in southern and central Sweden. The experimental plots were established on fertile forest sites and on former agricultural lands, and were managed with different numbers of thinnings and basal area removal regimes. The model best describing total stand basal area development was a dynamic equation derived from the Korf base model. The analysis of the realized gain trial for birch showed a good stability of the early calculated relative differences in basal area between tested genotypes over time. Thus, the relative difference in basal area might be with cautious used as representation of the realized genetic gain. On average forest sites in southern Sweden, improved and planted silver birch could produce between 6-10.5 m(3) ha(-1) year(-1), while on fertile agriculture land the average productivity might be higher, especially with material coming from the improvement program. The performed analysis provided a first step toward predicting the effects of genetic improvement on total volume production and profitability of silver birch. However, more experiments arc needed to set up the relative differences between different improved material

Replacing monocultures with mixed-species stands: Ecosystem service implications of two production forest alternatives in Sweden

Whereas there is evidence that mixed-species approaches to production forestry in general can provide positive outcomes relative to monocultures, it is less clear to what extent multiple benefits can be derived from specific mixed-species alternatives. To provide such insights requires evaluations of an encompassing suite of ecosystem services, biodiversity, and forest management considerations provided by specific mixtures and monocultures within a region. Here, we conduct such an assessment in Sweden by contrasting even-aged Norway spruce (Picea abies)-dominated stands, with mixed-species stands of spruce and birch (Betula pendula or B. pubescens), or spruce and Scots pine (Pinus sylvestris). By synthesizing the available evidence, we identify positive outcomes from mixtures including increased biodiversity, water quality, esthetic and recreational values, as well as reduced stand vulnerability to pest and pathogen damage. However, some uncertainties and risks were projected to increase, highlighting the importance of conducting comprehensive interdisciplinary evaluations when assessing the pros and cons of mixtures

Stand structure and future development of a managed multi-layered forest in southern Sweden

A heterogeneously structured forest stand with pine overstory and naturally regenerated spruce and oak trees in different size classes was documented. The effect of target diameter cutting on stand structure and growth was analyzed as a case-study. Both, systematic sample plots and forest gaps were used to describe the stand structure after cutting. Target diameter cutting in different treatments reduced the standing volume from ca. 320 to 180 m³/ha. Forest canopy gaps were created on more than 15% of the stand area. The seedling number of advanced natural regeneration was low (less than 500-1000 individuals per ha). Based on the advanced regeneration in gaps, three different scenarios for future ingrowth into the tree layer were defined. The extreme minimum ingrowth scenario assumed about 10 cm annual height growth and rather high mortality reported in literature from other experiments (resulting in one tree annually reaching 5 cm DBH during the next 50 years). Two other scenarios assumed 20 and 30 cm annual height growth. While no mortality was presumed within the latter scenario, moderate mortality rates (reported in literature) were chosen for the intermediate scenario. The maximum scenario postulates ten trees per year and ha reaching 5 cm DBH (equal to ingrowth reported from boreal single-tree selection forests). The moderate scenario assumes four new trees per year and he. Additional scenarios after soil preparation in gaps were used, defined on the base of shelterwood experiments. To describe the future basal area growth and continued target diameter cutting in the next 50 years, a single-tree growth model was applied, using stand age-independent estimations of the age of single trees. Thereby, a mean annual increment of 0.53-0.64 m2/ha was projected, similar to 5.6-6.8 m³/ha volume. Some errors to estimate the standing volume in multi-layered stands were detected during the simulation process. Compared to an evenaged spruce stand planted on the same site, the expected growth of the study stand during the next 50 years was one third lower. In average, about 120 m3/ha standing volume was removed in 20-25 years-cutting cycles. To continue without longer harvest intervals after the 50 years-simulation period, soil preparation seems necessary to achieve a sustainable number of small trees. Beside timber production, profitability was also lower by selective cutting. But, the important advantage of target diameter cutting can be more equally distributed income over time, with investments costs that can be covered by profit from timber harvest at the same time. A regular income of 17000-28000 SEK per ha every 20-25 years seems possible from today´s perspective. An additional treatment with alternative target diameters to promote particular tree species did not affect the amount of removals and the length of cutting intervals substantially. But simulations with 5 cm reduced target diameters caused very heavy removals and 35-40 years to reach 300 m³/ha standing volume again. The study includes discussions of tree species composition and development as well as a sensitivity analysis of the applied growth model