The History of Pastoral Activities in S Italy Inferred from Palynology: A Long-Term Perspective to Support Biodiversity Awareness

The present-day Mediterranean landscape is a result of the long-term human\u2013environment\u2013climate interactions that have driven the ecological dynamics throughout the Holocene. Pastoralism had (and still has) an important role in shaping this landscape, and contributes to maintaining the mosaic patterns of the Mediterranean habitats. Palaeoecological records provide significant multi-proxy data on environmental changes during the Holocene that are linked to human activities. In such research, the palynological approach is especially useful for detailing the complexity of anthropogenically-driven landscape transformations by discriminating past land uses and pastoral/breeding activities. This paper focuses on the palynological evidence for the impact of centuries of grazing on the vegetation of Basilicata, a region of southern Italy where animal breeding and pastoralism have a long tradition. A set of 121 pollen samples from eight archaeological sites (dated from the 6th century BC to the 15th century AD) and five modern surface soil samples were analyzed. The joint record of pollen pasture indicators and spores of coprophilous fungi suggests that continuous and intense pastoral activities have been practiced in the territory and have highly influenced its landscape. The palaeoecological results of this study provide us with better knowledge of the diachronical transformations of the habitats that were exposed to continuous grazing, with a shift toward more open vegetation and increase of sclerophyllous shrubs. The palynological approach gives insights into the vocation and environmental sustainability of this southern Italy region on a long-term basis

Heterocysts of Rivularia Type for Interpreting a Palaeoenvironmental Context of the Late Quaternary in Northern Italy

This paper presents new results on a quali-quantitative analysis of the heterocysts of the Rivularia type as a key bioindicator informative on local eutrophic conditions. The Rivularia type is usually reported in palynological analyses due to the thick, multilayered envelope that ensures the preservation of heterocysts in sediments. Samples come from two continuous terrestrial cores (N-S3: 77 samples, C-S1: 20 samples) drilled in the area surrounding the Bronze Age site of the Terramara S. Rosa di Poviglio (Po Plain; N Italy) and spanning at least over the last 15 ka years. Concentrations and percentages of the Rivularia type, combined with pollen curves of wet environments, describe local variability and rapid changes in ecological conditions over the millennia of deposition. Given the abundance and diversity of heterocysts of the Rivularia type in the studied samples, this paper attempts to group these cells based on morphology (ellipsoidal or elongated) and the state of preservation of the sheaths (presence or absence). Actually, it is difficult to confirm a relationship between heterocyst morphologies and the presence of different cyanobacteria species. Increasingly accurate identification of heterocysts from biostratigraphical archives may improve the data available on these bioindicators for achieving more detailed decoding of wetland (and terrestrial) transformations. Since the Bronze Age and at the most recent levels, the Rivularia type may be a good indicator of the local presence of agriculture and livestock, which lead to trophic and water changes in the soil

Multiporate Pollen of Poaceae as Bioindicator of Environmental Stress: First Archaeobotanical Evidence from the Early–Middle Holocene Site of Takarkori in the Central Sahara

This paper reports on the most ancient unusual morphological trait of the apertures of Poaceae pollen found in archaeological layers. In Poaceae, high levels of hybridization, polyploidy, apomixis, and multiporate pollen are often related. Multiple genomes in polyploids are critical for the adaptation of plant species to stresses and could be revealed by anomalies in pollen development. Therefore, the paleoenvironmental research can gain great benefits from identifying polyploids in past contexts by observing anomalous pollen morphology during pollen counts. The occurrence of multiporate pollen in Poaceae has also been related to special features of the ecology of the species showing this anomaly, as well as to climatic and environmental stresses experienced by Poaceae living in a given region. Multiporate and bi- or tri-porate instead of monoporate pollen grains have been observed in samples taken from Takarkori rockshelter, an archaeological site in southwestern Libya (central Sahara) that has been occupied between ~10,200 and ~4650 cal BP. Multiporate pollen was found in organic sands and coprolites of ovicaprines. On the basis of archaeobotanical research, this work aims to investigate whether the presence of supernumerary pores in Poaceae pollen may be an effect of both climatic/hydrological changes and continued anthropogenic pressure on the wild grasses living in the region. The presence of multiporate pollen reveals that Poaceae that lived in central Sahara tackled several kinds of stress during the early and middle Holocene. The Takarkori pollen record suggests that climate change could have played a major role in the early Holocene, while human pressure became stronger during the middle Holocene. The change in environmental conditions determined adaptive responses of polyploid grasses even in the form of multiporate pollen

ECONOMY AND ENVIRONMENT OF THE GREEK COLONIAL SYSTEM IN SOUTHERN ITALY: POLLEN AND NPPS EVIDENCE OF GRAZING FROM THE RURAL SITE OF FATTORIA FABRIZIO (VI-IV CENT. BC; METAPONTO, BASILICATA)

this paper reports a study case showing integrated analyses of microscopic records from an archaeological site of southern italy. pollen and non pollen palynomorphs-Npps were found in archaeological layers and were basic in reconstructing both the past environment and the economic activities of the site. the site, Fattoria Fabrizio, is a modest 4th-cenury bc farmhouse of the chora (rural territory) of the Greek city (polis) of Metaponto (basilicata). pollen analyses in addition to the study of Npps have been particularly worthwhile for palaeoenvironmental and palaeoeconomical reconstructions of this site and others in the chora. the pollen spectra delineate an open plant landscape, with scanty woodlands and presence of local wet environments. Olea pollen is fairly well represented in all samples, suggesting that this tree was an important element of the agricultural economy of the chora. shrubby grasslands and a well-developed macchia characterized the territory, probably as result of grazing activities by sheep and goats. accordingly, the high percentages of poaceae and cichorieae pollen, together with coprophilous fungal spores (such as Sordaria type and Sporormiella type), suggest that pastoral activities were widely practiced. the low number of cerealia pollen grains suggests that the inhabitants’ fields covered small areas or were quite far from the farmhouse

Plant Responses to Climate Change: The Case Study of Betulaceae and Poaceae Pollen Seasons (Northern Italy, Vignola, Emilia-Romagna)

Aerobiological data have especially demonstrated that there is correlation between climate warming and the pollination season of plants. This paper focuses on airborne pollen monitoring of Betulaceae and Poaceae, two of the main plant groups with anemophilous pollen and allergenic proprieties in Northern Italy. The aim is to investigate plant responses to temperature variations by considering long-term pollen series. The 15-year aerobiological analysis is reported from the monitoring station of Vignola (located near Modena, in the Emilia-Romagna region) that had operated in the years 1990-2004 with a Hirst spore trap. The Yearly Pollen Index calculated for these two botanical families has shown contrasting trends in pollen production and release. These trends were well identifiable but fairly variable, depending on both meteorological variables and anthropogenic causes. Based on recent reference literature, we considered that some oscillations in pollen concentration could have been a main effect of temperature variability reflecting global warming. The duration of pollen seasons of Betulaceae and Poaceae, depending on the different species included in each family, has not unequivocally been determined. Phenological responses were particularly evident in Alnus and especially in Corylus as a general moving up of the end of pollination. The study shows that these trees can be affected by global warming more than other, more tolerant, plants. The research can be a contribution to the understanding of phenological plant responses to climate change and suggests that alder and hazelnut trees have to be taken into high consideration as sensible markers of plant responses to climate change

Humans and Water in Desert “Refugium” Areas: Palynological Evidence of Climate Oscillations and Cultural Developments in Early and Mid-Holocene Saharan Edges

Saharan anthropic deposits from archaeological sites, located along wadis or close to lakes, and sedimentary sequences from permanent and dried basins demonstrate that water has always been an attractive environmental feature, especially during periods of drought. This paper reports on two very different examples of Holocene sites where “humans and water” coexisted during dry periods, as observed by stratigraphic, archaeological and palynological evidence. Independent research was carried out on the Jefara Plain (Libya, 32°N) and the Gobero area (Niger, 17°N), at the extreme northern and southern limits of the Sahara, respectively. The histories of the Jefara and Gobero areas, as revealed by the archaeological and palaeoenvironmental reconstructions, suggest that these areas were likely to have been visited and exploited for a long time, acting as anthropic refugia, and therefore they have been profoundly transformed. Human presence and actions have conditioned the local growing of plants and selected a more or less synanthropic flora. Today, modern conservation strategies should take into consideration that water reservoirs, which are crucial for the long-term conservation of biodiversity, have provided refugia in the past just as they presently do under global warming conditions

The representativeness of Olea pollen from olive groves and the Late Holocene landscape reconstruction in central Mediterranean

Modern pollen spectra are an invaluable reference tool for paleoenvironmental and cultural landscape reconstructions, but the importance of knowing the pollen rain released from orchards remains underexplored. In particular, the role of cultivated trees is in past and current agrarian landscapes has not been fully investigated. Here, we present a pollen analysis of 70 surface soil samples taken from 12 olive groves in Basilicata and Tuscany, two regions of Italy that exemplify this cultivation in the Mediterranean basin. This study was carried out to assess the representativeness of Olea pollen in modern cultivations. Although many variables can influence the amount of pollen observed in soils, it was clear that most of the pollen was deposited below the trees in the olive groves. A rapid decline in the olive pollen percentages (c. 85% on average) was found when comparing samples taken from IN vs. OUT of each grove. The mean percentages of Olea pollen obtained from the archaeological sites close to the studied orchards suggest that olive groves were established far from the Roman farmhouses of Tuscany. Further south, in the core of the Mediterranean basin, the cultivation of Olea trees was likely situated approximately 500\u20131000 m from the rural sites in Basilicata, and dated from the Hellenistic to the Medieval period

Palaeoenvironment, Settlement, and Land Use in the Late Neolithic—Bronze Age Site of Colombare di Negrar di Valpolicella (N Italy, On-Site)

Palynological and archaeobotanical analyses have been carried out as part of the interdisciplinary project of Colombare di Negrar, a prehistoric site in the Lessini Mountains (northern Italy). The palaeoenvironmental and economic reconstruction from the Late Neolithic to the beginning of the Early Bronze Age was based on 16 pollen samples and three samples of macroremains taken from two contiguous trenches. The landscape reconstruction shows the presence of natural clearings in the wood. Forest cover was characterised by oak wood, with Ulmus and Tilia. The intermediate morphology of size and exine of Tilia cordata/platyphyllos pollen may be regarded as the first palynological evidence of lime hybrids in palaeorecords. Hygrophilous trees and Vitis vinifera testify to the presence of riparian forests and moist soils. Among trees supplying fruits, in addition to the grapevine, hazelnut (Corylus avellana) and walnut (Juglans regia) were present. A mixed economy based on animal breeding and cultivation of cereals (Hordeum vulgare, Triticum monococcum, T. dicoccum, T. timopheevii) emerged from the data. The combined analysis of pollen and plant macroremains suggests that different activities were carried out simultaneously in Colombare and a relationship between natural resources and the socio-economic and cultural evolution of the territory