34 research outputs found
ΠΠΠ₯ΠΠΠΠΠΠ«βΠ ΠΠΠΠΠ’ΠΠ―βΠ§ΠΠΠΠΠΠ§ΠΠ‘ΠΠΠΠβΠΠΠΠΠ’ΠΠΠ ΠβΠΠΠ ΠΠ ΠΠΠβΠ‘ΠΠΠ’ΠΠ Π
The publication considers problems and mechanisms of human capital assets development. The authors review the poll of rural people in order to estimate financial situation in concern of living conditions and development of human capital assets in rural areas. Studying of human capital assets is explained by new review of a person and his life as the main assets of Russian society and one of the main factors of economic progress. Modernization and technological re-equipment of the industrial sphere in Russia, agribusiness re-equipment is possible in case of sufficient human capital assets and its efficient application. Development of human capital assets and its quality increasing contributes to solving socio-economic problems and strengthens innovative development in rural areas. Human capital assets are the main source of various innovations and social changes; it is the factor of transition to new quality which is found in all spheres of social life. Nowadays, innovative development of rural areas depends on human capital assets. Bad financial situation is a strong factor restricting successful rural development. Insufficient level of human capital assets development in rural areas leads to higher requirements to qualifications, solving socio-economic problems of rural areas, new conception of personnel training and development of human capital assets. The publication suggests the main principles of conception of development of human capital assets (strengthening of state support in rural areas, development of entrepreneurship, investments in personal development, education, medicine and culture).Π Π°ΡΡΠΌΠ°ΡΡΠΈΠ²Π°ΡΡΡΡ ΠΏΡΠΎΠ±Π»Π΅ΠΌΡ, ΠΌΠ΅Ρ
Π°Π½ΠΈΠ·ΠΌΡ ΡΠ°Π·Π²ΠΈΡΠΈΡ ΡΠ΅Π»ΠΎΠ²Π΅ΡΠ΅ΡΠΊΠΎΠ³ΠΎ ΠΊΠ°ΠΏΠΈΡΠ°Π»Π°. ΠΡΠΎΠ²Π΅Π΄Π΅Π½ ΠΎΠΏΡΠΎΡ ΡΠ΅Π»ΡΡΠΊΠΎΠ³ΠΎ Π½Π°ΡΠ΅Π»Π΅Π½ΠΈΡ ΡΒ ΡΠ΅Π»ΡΡ ΠΎΡΠ΅Π½ΠΊΠΈ ΠΌΠ°ΡΠ΅ΡΠΈΠ°Π»ΡΠ½ΠΎΠ³ΠΎ ΠΏΠΎΠ»ΠΎΠΆΠ΅Π½ΠΈΡ, ΠΆΠΈΠ»ΠΈΡΠ½ΡΡ
ΠΈΒ Π΄ΡΡΠ³ΠΈΡ
ΡΡΠ»ΠΎΠ²ΠΈΠΉ ΡΠ°Π·Π²ΠΈΡΠΈΡ ΡΠ΅Π»ΠΎΠ²Π΅ΡΠ΅ΡΠΊΠΎΠ³ΠΎ ΠΊΠ°ΠΏΠΈΡΠ°Π»Π° Π½Π° ΡΠ΅Π»Π΅. ΠΠ·ΡΡΠ΅Π½ΠΈΠ΅ ΡΠ΅Π»ΠΎΠ²Π΅ΡΠ΅ΡΠΊΠΎΠ³ΠΎ ΠΊΠ°ΠΏΠΈΡΠ°Π»Π° Π½Π° ΡΠ΅Π³ΠΎΠ΄Π½ΡΡΠ½ΠΈΠΉ Π΄Π΅Π½Ρ ΠΎΠΏΡΠ΅Π΄Π΅Π»ΡΠ΅ΡΡΡ Π½Π΅ΠΎΠ±Ρ
ΠΎΠ΄ΠΈΠΌΠΎΡΡΡΡ Π½ΠΎΠ²ΠΎΠ³ΠΎ Π²Π·Π³Π»ΡΠ΄Π° Π½Π° ΡΠ΅Π»ΠΎΠ²Π΅ΠΊΠ° ΠΈΒ ΡΠ΅Π»ΠΎΠ²Π΅ΡΠ΅ΡΠΊΡΡ ΠΆΠΈΠ·Π½Ρ ΠΊΠ°ΠΊ Π³Π»Π°Π²Π½ΡΠ΅ Π΄ΠΎΡΡΠΎΡΠ½ΠΈΡ ΡΠΎΡΡΠΈΠΉΡΠΊΠΎΠ³ΠΎ ΠΎΠ±ΡΠ΅ΡΡΠ²Π° ΠΈΒ ΠΎΠ΄ΠΈΠ½ ΠΈΠ· Π²Π°ΠΆΠ½Π΅ΠΉΡΠΈΡ
ΡΠ°ΠΊΡΠΎΡΠΎΠ² Π΄ΠΎΡΡΠΈΠΆΠ΅Π½ΠΈΡ ΡΠΊΠΎΠ½ΠΎΠΌΠΈΡΠ΅ΡΠΊΠΎΠ³ΠΎ ΠΏΡΠΎΠ³ΡΠ΅ΡΡΠ°. ΠΠΎΠ΄Π΅ΡΠ½ΠΈΠ·Π°ΡΠΈΡ ΠΈΒ ΡΠ΅Ρ
Π½ΠΎΠ»ΠΎΠ³ΠΈΡΠ΅ΡΠΊΠΎΠ΅ ΠΎΠ±Π½ΠΎΠ²Π»Π΅Π½ΠΈΠ΅ Π²ΡΠ΅ΠΉ ΠΏΡΠΎΠΈΠ·Π²ΠΎΠ΄ΡΡΠ²Π΅Π½Π½ΠΎΠΉ ΡΡΠ΅ΡΡ ΡΡΡΠ°Π½Ρ, Π²Β ΡΠΎΠΌ ΡΠΈΡΠ»Π΅ ΠΠΠ, Π²ΠΎΠ·ΠΌΠΎΠΆΠ½ΠΎ ΠΏΡΠΈ ΡΡΠ»ΠΎΠ²ΠΈΠΈ ΡΠΎΡΠΌΠΈΡΠΎΠ²Π°Π½ΠΈΡ Π΄ΠΎΡΡΠ°ΡΠΎΡΠ½ΠΎ Π²ΡΡΠΎΠΊΠΎΠ³ΠΎ ΡΡΠΎΠ²Π½Ρ ΡΠ°Π·Π²ΠΈΡΠΈΡ ΡΠ΅Π»ΠΎΠ²Π΅ΡΠ΅ΡΠΊΠΎΠ³ΠΎ ΠΊΠ°ΠΏΠΈΡΠ°Π»Π° ΠΈΒ Π΅Π³ΠΎ ΡΡΡΠ΅ΠΊΡΠΈΠ²Π½ΠΎΠ³ΠΎ ΠΈΡΠΏΠΎΠ»ΡΠ·ΠΎΠ²Π°Π½ΠΈΡ. Π Π°Π·Π²ΠΈΡΠΈΠ΅ ΡΠ΅Π»ΠΎΠ²Π΅ΡΠ΅ΡΠΊΠΎΠ³ΠΎ ΠΊΠ°ΠΏΠΈΡΠ°Π»Π°, ΠΏΠΎΠ²ΡΡΠ΅Π½ΠΈΠ΅ Π΅Π³ΠΎ ΠΊΠ°ΡΠ΅ΡΡΠ² ΠΏΠΎΠ·Π²ΠΎΠ»ΡΠ΅Ρ ΡΠ΅ΡΠ°ΡΡ ΠΌΠ½ΠΎΠ³ΠΎΡΠΈΡΠ»Π΅Π½Π½ΡΠ΅ ΡΠΎΡΠΈΠ°Π»ΡΠ½ΠΎ-ΡΠΊΠΎΠ½ΠΎΠΌΠΈΡΠ΅ΡΠΊΠΈΠ΅ ΠΏΡΠΎΠ±Π»Π΅ΠΌΡ, ΡΡΠΊΠΎΡΡΠ΅Ρ ΠΈΠ½Π½ΠΎΠ²Π°ΡΠΈΠΎΠ½Π½ΠΎΠ΅ ΡΠ°Π·Π²ΠΈΡΠΈΠ΅ ΡΠ΅Π»ΡΡΠΊΠΈΡ
ΡΠ΅ΡΡΠΈΡΠΎΡΠΈΠΉ. Π§Π΅Π»ΠΎΠ²Π΅ΡΠ΅ΡΠΊΠΈΠΉ ΠΊΠ°ΠΏΠΈΡΠ°Π» ΡΠ²Π»ΡΠ΅ΡΡΡ ΠΎΡΠ½ΠΎΠ²Π½ΡΠΌ ΠΈΡΡΠΎΡΠ½ΠΈΠΊΠΎΠΌ ΠΌΠ½ΠΎΠ³ΠΎΡΠΈΡΠ»Π΅Π½Π½ΡΡ
Π½ΠΎΠ²Π°ΡΠΈΠΉ ΠΈΒ ΠΏΠ΅ΡΠ΅ΠΌΠ΅Π½ Π²Β ΠΎΠ±ΡΠ΅ΡΡΠ²Π΅, ΡΠ°ΠΊΡΠΎΡΠΎΠΌ Π΅Π³ΠΎ ΡΠΎΠ²Π΅ΡΡΠ΅Π½ΡΡΠ²ΠΎΠ²Π°Π½ΠΈΡ, ΠΏΠ΅ΡΠ΅Ρ
ΠΎΠ΄Π° ΠΊΒ Π½ΠΎΠ²ΠΎΠΌΡ ΠΊΠ°ΡΠ΅ΡΡΠ²Ρ ΠΈΒ ΠΏΠΎΠ»ΡΡΠ°Π΅Ρ Π²ΠΎΠΏΠ»ΠΎΡΠ΅Π½ΠΈΠ΅ Π²Β ΡΠ°Π·Π²ΠΈΡΠΈΠΈ ΡΡΠ΅Ρ ΠΆΠΈΠ·Π½ΠΈ ΠΎΠ±ΡΠ΅ΡΡΠ²Π°. ΠΒ ΡΠΎΠ²ΡΠ΅ΠΌΠ΅Π½Π½ΡΡ
ΡΡΠ»ΠΎΠ²ΠΈΡΡ
ΠΈΠ½Π½ΠΎΠ²Π°ΡΠΈΠΎΠ½Π½ΠΎΠ΅ ΡΠ°Π·Π²ΠΈΡΠΈΠ΅ ΡΠ΅Π»ΡΡΠΊΠΈΡ
ΡΠ΅ΡΡΠΈΡΠΎΡΠΈΠΉ Π²Β Π·Π½Π°ΡΠΈΡΠ΅Π»ΡΠ½ΠΎΠΉ ΡΡΠ΅ΠΏΠ΅Π½ΠΈ ΠΎΠΏΡΠ΅Π΄Π΅Π»ΡΠ΅ΡΡΡ Π½Π°ΠΊΠΎΠΏΠ»Π΅Π½Π½ΡΠΌ ΡΠ΅Π»ΠΎΠ²Π΅ΡΠ΅ΡΠΊΠΈΠΌ ΠΊΠ°ΠΏΠΈΡΠ°Π»ΠΎΠΌ. ΠΠΈΠ·ΠΊΠΈΠΉ ΡΡΠΎΠ²Π΅Π½Ρ ΠΌΠ°ΡΠ΅ΡΠΈΠ°Π»ΡΠ½ΠΎΠ³ΠΎ ΠΏΠΎΠ»ΠΎΠΆΠ΅Π½ΠΈΡ Π»ΡΠ΄Π΅ΠΉ ΡΠ²Π»ΡΠ΅ΡΡΡ ΠΌΠΎΡΠ½ΡΠΌ ΠΎΠ³ΡΠ°Π½ΠΈΡΠΈΡΠ΅Π»Π΅ΠΌ ΡΡΡΠΎΠΉΡΠΈΠ²ΠΎΠ³ΠΎ ΡΠ°Π·Π²ΠΈΡΠΈΡ ΡΠ΅Π»Π°, Π°Β Π΅Π³ΠΎ ΠΏΡΠ΅ΠΎΠ΄ΠΎΠ»Π΅Π½ΠΈΠ΅Β β Π²Π°ΠΆΠ½Π΅ΠΉΡΠ΅Π΅ ΡΡΠ»ΠΎΠ²ΠΈΠ΅ ΡΠ°ΠΊΠΎΠ³ΠΎ ΡΠ°Π·Π²ΠΈΡΠΈΡ. ΠΠ΅Π΄ΠΎΡΡΠ°ΡΠΎΡΠ½ΡΠΉ ΡΡΠΎΠ²Π΅Π½Ρ ΡΠ°Π·Π²ΠΈΡΠΈΡ ΡΠ΅Π»ΠΎΠ²Π΅ΡΠ΅ΡΠΊΠΎΠ³ΠΎ ΠΊΠ°ΠΏΠΈΡΠ°Π»Π° Π²Β ΡΠ΅Π»ΡΡΠΊΠΎΠΉ ΠΌΠ΅ΡΡΠ½ΠΎΡΡΠΈ ΡΡΠ΅Π±ΡΠ΅Ρ ΡΠ΅ΡΠ΅Π½ΠΈΡ ΠΏΡΠΎΠ±Π»Π΅ΠΌ ΡΠΎΡΠΈΠ°Π»ΡΠ½ΠΎ-ΡΠΊΠΎΠ½ΠΎΠΌΠΈΡΠ΅ΡΠΊΠΎΠ³ΠΎ ΡΠ°Π·Π²ΠΈΡΠΈΡ ΡΠ΅Π»Π°, ΠΏΠΎΠ²ΡΡΠ΅Π½ΠΈΡ ΡΡΠ΅Π±ΠΎΠ²Π°Π½ΠΈΠΉ ΠΊΒ ΠΊΠ²Π°Π»ΠΈΡΠΈΠΊΠ°ΡΠΈΠΈ ΡΡΡΠ΄Π°, ΡΠ°Π·ΡΠ°Π±ΠΎΡΠΊΠΈ Π½ΠΎΠ²ΠΎΠΉ ΠΊΠΎΠ½ΡΠ΅ΠΏΡΠΈΠΈ ΠΏΠΎΠ΄Π³ΠΎΡΠΎΠ²ΠΊΠΈ ΠΊΠ°Π΄ΡΠΎΠ². ΠΡΠ΅Π΄Π»ΠΎΠΆΠ΅Π½Ρ ΠΎΡΠ½ΠΎΠ²Π½ΡΠ΅ ΠΏΡΠΈΠ½ΡΠΈΠΏΡ ΠΊΠΎΠ½ΡΠ΅ΠΏΡΠΈΠΈ ΡΠ°Π·Π²ΠΈΡΠΈΡ ΡΠ΅Π»ΠΎΠ²Π΅ΡΠ΅ΡΠΊΠΎΠ³ΠΎ ΠΊΠ°ΠΏΠΈΡΠ°Π»Π° (ΡΡΠΈΠ»Π΅Π½ΠΈΠ΅ ΠΌΠ΅Ρ Π³ΠΎΡΡΠ΄Π°ΡΡΡΠ²Π΅Π½Π½ΠΎΠΉ ΠΏΠΎΠ΄Π΄Π΅ΡΠΆΠΊΠΈ ΡΠ΅Π», ΡΠ°Π·Π²ΠΈΡΠΈΠ΅ ΠΏΡΠ΅Π΄ΠΏΡΠΈΠ½ΠΈΠΌΠ°ΡΠ΅Π»ΡΡΡΠ²Π°, ΡΠ²Π΅Π»ΠΈΡΠ΅Π½ΠΈΠ΅ ΠΈΠ½Π²Π΅ΡΡΠΈΡΠΈΠΉ Π²Β ΡΠ°Π·Π²ΠΈΡΠΈΠ΅ ΡΠ΅Π»ΠΎΠ²Π΅ΠΊΠ°, ΠΎΠ±ΡΠ°Π·ΠΎΠ²Π°Π½ΠΈΡ, Π·Π΄ΡΠ°Π²ΠΎΠΎΡ
ΡΠ°Π½Π΅Π½ΠΈΡ ΠΈΒ ΠΊΡΠ»ΡΡΡΡΡ)
Characterization of the cork oak transcriptome dynamics during acorn development
Background: Cork oak (Quercus suber L.) has a natural distribution across western Mediterranean regions and is a keystone forest tree species in these ecosystems. The fruiting phase is especially critical for its regeneration but the molecular mechanisms underlying the biochemical and physiological changes during cork oak acorn development are poorly understood. In this study, the transcriptome of the cork oak acorn, including the seed, was characterized in five stages of development, from early development to acorn maturation, to identify the dominant processes in each stage and reveal transcripts with important functions in gene expression regulation and response to water.
Results: A total of 80,357 expressed sequence tags (ESTs) were de novo assembled from RNA-Seq libraries representative of the several acorn developmental stages. Approximately 7.6Β % of the total number of transcripts present in Q. suber transcriptome was identified as acorn specific. The analysis of expression profiles during development returned 2,285 differentially expressed (DE) transcripts, which were clustered into six groups. The stage of development corresponding to the mature acorn exhibited an expression profile markedly different from other stages. Approximately 22Β % of the DE transcripts putatively code for transcription factors (TF) or transcriptional regulators, and were found almost equally distributed among the several expression profile clusters, highlighting their major roles in controlling the whole developmental process. On the other hand, carbohydrate metabolism, the biological pathway most represented during acorn development, was especially prevalent in mid to late stages as evidenced by enrichment analysis. We further show that genes related to response to water, water deprivation and transport were mostly represented during the early (S2) and the last stage (S8) of acorn development, when tolerance to water desiccation is possibly critical for acorn viability.
Conclusions: To our knowledge this work represents the first report of acorn development transcriptomics in oaks. The obtained results provide novel insights into the developmental biology of cork oak acorns, highlighting transcripts putatively involved in the regulation of the gene expression program and in specific processes likely essential for adaptation. It is expected that this knowledge can be transferred to other oak species of great ecological value.Fundação para a CiΓͺncia e a Tecnologi