Orbital forcing and role of the latitudinal insolation/temperature gradient

Orbital forcing of the climate system is clearly shown in the Earths record of glacial-interglacial cycles, but the mechanism underlying this forcing is poorly understood. Traditional Milankovitch theory suggests that these cycles are driven by changes in high latitude summer insolation, yet this forcing is dominated by precession, and cannot account for the importance of obliquity in the Ice Age record. Here, we investigate an alternative forcing based on the latitudinal insolation gradient (LIG), which is dominated by both obliquity (in summer) and precession (in winter). The insolation gradient acts on the climate system through differential solar heating, which creates the Earths latitudinal temperature gradient (LTG) that drives the atmospheric and ocean circulation. A new pollen-based reconstruction of the LTG during the Holocene is used to demonstrate that the LTG may be much more sensitive to changes in the LIG than previously thought. From this, it is shown how LIG forcing of the LTG may help explain the propagation of orbital signatures throughout the climate system, including the Monsoon, Arctic Oscillation and ocean circulation. These relationships are validated over the last (Eemian) Interglacial, which occurred under a different orbital configuration to the Holocene. We conclude that LIG forcing of the LTG explains many criticisms of classic Milankovitch theory, while being poorly represented in climate model

Palaeolimnology and holocene environmental change from endoreic lakes in the Ebro Basin, North-East Spain

PhD ThesisLake sediments from inland endoreic (saline) lakes in the semi-arid Ebro Basin, NE Spain have been analysed to provide a history of lake level, vegetation, catchment erosion and anthropogenic activity over the last 10,000 years. Analysis was undertaken for pollen, macrofossils (seeds, Cladocera ephippia, Chironomid head capsules etc), charcoal, geochemistry (total cations/trace metals, sulphate, carbonate & LOI) and sediment composition. Fourteen AMS radiocarbon dates provide dating control. Seven cores were investigated from 4 seasonal playa lakes, I shallow (<1.5m) semi-permanent salt lake, 1 deep (5.0m) permanent salt lake and 1 Medieval-age reservoir. Over 40 surface samples were also taken to investigate modern analogue environments. A review of the use and interpretation of saline lake macrophytes (seeds & pollen), Cladocera and Chironomids in palaeolimnology is provided, with special emphasis on those found in Spanish salt lakes. Taphonomic problems and nearshore-offshore facies were also investigated using a surface sample transect across a small playa lake. A surface sample pollen data set from 30 lakes in the Ebro Basin is presented and the implications for palaeo-interpretation discussed. The sensitivity of the pollen record as a climate indicator is investigated using 6 sites across a climatic gradient from sub-humid to semi-arid. Lake level reconstruction is based on an 8 stage semi-quantitative palaeohydrological model, developed from a surface sample data set from 32 lakes ranging from temporary to permanent, and hypersaline to freshwater. A simple hydrological model for groundwater fed lakes is also discussed which can be used to quantify these palaeohydrological changes. The early Holocene (<9.3-8.6Kyr BP) was characterised by semi-arid extreme continental conditions in the Ebro Basin, with an extensive Juniper thurifera woodland. Lake levels rose to their highest point in the Holocene between 8.6-7.2Kyr BP when evergreen oak and pine forest dominated. This is interrupted by a short recession in lake level between 8.2-7.6Kyr BP. A distinct regional early-Neolithic fire and clearance event occurs between 7.7-7.3Kyr BP. A drop in lake level and development of a monospecific pine forest (P. halepensis) indicates warmer and drier conditions in the mid Holocene (7.2-5.4Kyr BP). Evergreen oak reappears as forest cover declines after 5.4Kyr BP, although this is not marked by any increase in charcoal or cultivation indicators. Low groundwater but moderate lake levels (4.0-2.7Kyr BP) may be linked to high summer storm frequency and low winter rainfall. This coincides with catchment erosion and valley floor alluviation. Lake level rises again significantly between 2.7-1.8Kyr BP during Iberian and Roman times when archaeological evidence indicates a peak in population. Agriculture changes from small scale pastoral to small scale arable without any further decline in woodland cover. A rapid fall in lake level at ca. 1.8Kyr BP is followed by complete forest recovery (P. halepensis) and depopulation, until major deforestation around 1.4Kyr BP marks the arrival of the Visigoths/Arabs and extensive nomadic pastoralism. Intensive grazing pressure or lower temperatures resulted in Juniperus increasing again between 1.4-0.4Kyr BP. This also coincides with a second period of catchment erosion and valley floor alluviation. After ca. 0.4Kyr BP, lake levels have increased along with extensive olive cultivation and the development of modern (irrigated) arable agriculture.Natural Environment Research Counci

Poynting Vector Flow in a Circular Circuit

A circuit is considered in the shape of a ring, with a battery of negligible size and a wire of uniform resistance. A linear charge distribution along the wire maintains an electrostatic field and a steady current, which produces a constant magnetic field. Earlier studies of the Poynting vector and the rate of flow of energy considered only idealized geometries in which the Poynting vector was confined to the space within the circuit. But in more realistic cases the Poynting vector is nonzero outside as well as inside the circuit. An expression is obtained for the Poynting vector in terms of products of integrals, which are evaluated numerically to show the energy flow. Limiting expressions are obtained analytically. It is shown that the total power generated by the battery equals the energy flowing into the wire per unit time.Comment: 19 pages, 8 figure

Role of Platelet-Activating Factor and Hypoxia in Persistent Pulmonary Hypertension of the Newborn — Studies with Perinatal Pulmonary Vascular Smooth Muscle Cells

Platelet-activating factor (PAF) plays an important physiological role of maintaining a high vasomotor tone in fetal pulmonary circulation. At birth, endogenous vasodilators such as nitric oxide and prostacyclin are released and facilitate pulmonary vasodilation via cAMP-dependent protein kinase (cAMP/PKA) and cGMP-dependent protein kinase (cGMP/PKG) pathways. Interaction between the cyclic nucleotides and PAF receptor (PAFR)-mediated responses in pulmonary arterial smooth muscle is not well understood. To further understand the interactions of PAF-PAFR pathway and the cyclic nucleotides in ovine fetal pulmonary arterial smooth muscle cells (FPASMC), effects of cAMP and cGMP on PAFR-mediated responses in pulmonary arterial smooth muscle cells (PASMC) were studied. Ovine FPASMC were incubated with 10μM cAMP or cGMP in normoxia (5% CO2 in air, pO2~100 Torr) or hypoxia (2% O2, 5% CO2, pO2~30-40 Torr). Proteins were prepared and subjected to Western blotting. Effect of cell permeable cAMP and cGMP on PAFR binding was also studied and effect of cAMP on cell proliferation was also studied by RNAi to PKA-Cα. cAMP and cGMP significantly decreased PAFR binding and protein expression in normoxia and hypoxia, more so in hypoxia, when PAFR expression was usually high. PKA-Cα siRNA demonstrated that inhibition of PAFR-mediated responses by the cyclic nucleotides occurred through PKA. These data suggest that the normally high levels of cyclic nucleotides in the normoxic newborn pulmonary circulation assist in the downregulation of postnatal PAFR-mediated responses and that under hypoxic conditions, increasing the levels of cyclic nucleotides will abrogate PAF-mediated vasoconstriction thereby ameliorating PAF-induced persistent pulmonary hypertension of the newborn

Holocene global mean surface temperature, a multi-method reconstruction approach

An extensive new multi-proxy database of paleo-temperature time series (Temperature 12k) enables a more robust analysis of global mean surface temperature (GMST) and associated uncertainties than was previously available. We applied five different statistical methods to reconstruct the GMST of the past 12,000 years (Holocene). Each method used different approaches to averaging the globally distributed time series and to characterizing various sources of uncertainty, including proxy temperature, chronology and methodological choices. The results were aggregated to generate a multi-method ensemble of plausible GMST and latitudinal-zone temperature reconstructions with a realistic range of uncertainties. The warmest 200-year-long interval took place around 6500 years ago when GMST was 0.7 °C (0.3, 1.8) warmer than the 19 th Century (median, 5 th , 95 th percentiles). Following the Holocene global thermal maximum, GMST cooled at an average rate −0.08 °C per 1000 years (−0.24, −0.05). The multi-method ensembles and the code used to generate them highlight the utility of the Temperature 12k database, and they are now available for future use by studies aimed at understanding Holocene evolution of the Earth system

STRADITIZE: An open-source program for digitizing pollen diagrams and other types of stratigraphic data

In an age of digital data analysis, gaining access to data from the pre-digital era – or any data that is only available as a figure on a page – remains a problem and an under-utilized scientific resource. Whilst there are numerous programs available that allow the digitization of scientific data in a simple x-y graph format, we know of no semi-automated program that can deal with data plotted with multiple horizontal axes that share the same vertical axis, such as pollen diagrams and other stratigraphic figures that are common in the Earth sciences. STRADITIZE (Stratigraphic Diagram Digitizer) is a new open-source program that allows stratigraphic figures to be digitized in a single semi-automated operation. It is designed to detect multiple plots of variables analyzed along the same vertical axis, whether this is a sediment core or any similar depth/time series. The program is written in python and supports mixtures of many different diagram types, such as bar plots, line plots, as well as shaded, stacked, and filled area plots. The package provides an extensively documented graphical user interface for a point-and-click handling of the semi-automatic process, but can also be scripted or used from the command line. Other features of STRADITIZE include text recognition to interpret the names of the different plotted variables, the automatic and semi-automatic recognition of picture artifacts, as well an automatic measurement finder to exactly reproduce the data that has been used to create the diagram. Evaluation of the program has been undertaken comparing the digitization of published figures with the original digital data. This generally shows very good results, although this is inevitably reliant on the quality and resolution of the original figure

On Exchange of Orbital Angular Momentum Between Twisted Photons and Atomic Electrons

We obtain an expression for the matrix element for a twisted (Laguerre-Gaussian profile) photon scattering from a hydrogen atom. We consider photons incoming with an orbital angular momentum (OAM) of $\ell \hbar$, carried by a factor of $e^{i \ell \phi}$ not present in a plane-wave or pure Gaussian profile beam. The nature of the transfer of $+2\ell$ units of OAM from the photon to the azimuthal atomic quantum number of the atom is investigated. We obtain simple formulae for these OAM flip transitions for elastic forward scattering of twisted photons when the photon wavelength $\lambda$ is large compared with the atomic target size $a$, and small compared the Rayleigh range $z_R$, which characterizes the collimation length of the twisted photon beam.Comment: 16 page

Evaluation of the functional role of the maize Glossy2 and Glossy2-like genes in cuticular lipid deposition

Plant epidermal cells express unique molecular machinery that juxtapose the assembly of intracellular lipid components and the unique extracellular cuticular lipids that are unidirectionally secreted to plant surfaces. In maize (Zea mays L.), mutations at the glossy2 (gl2) locus affect the deposition of extracellular cuticular lipids. Sequence-based genome scanning identified a novel gl2 homolog in the maize genome, Gl2-like. Sequence homology identifies that both the Gl2-like and Gl2 genes are members of the BAHD superfamily of acyltransferases, with close sequence homology to the Arabidopsis CER2 gene. Transgenic experiments demonstrate that Gl2-like and Gl2 functionally complement the Arabidopsis cer2 mutation, with differential impacts on the cuticular lipids and the lipidome of the plant, particularly affecting the longer alkyl chain acyl lipids, particularly at the 32-carbon chain length. Site-directed mutagenesis of the putative BAHD catalytic HXXXDX-motif indicates that Gl2-like requires this catalytic capability to fully complement the cer2 function, but Gl2 can accomplish this without the need for this catalytic motif. These findings demonstrate that both Gl2 and Gl2-like overlap in their cuticular lipid function, however the two genes have evolutionary diverged to acquire non-overlapping functions