Zur Fauna der Zikaden, Wanzen und Augenfliegen des Kaiserstuhls: (Hemiptera: Auchenorrhyncha et Heteroptera; Diptera: Pipunculidae)

On the fauna of hemipteran bugs (Hemiptera: Auchenorrhyncha et Heteroptera) and big-headed flies (Diptera: Pipunculidae) of the Kaiserstuhl, southwestern Germany. – Field records of 157 Auchenorrhyncha species gathered on field excursionsto five sites in the Kaiserstuhl Hill during the 9th Auchenorrhyncha Meeting, June 14th - 15th 2002, are presented. Particularly noteworthy was the first German record of the leafhopper Arocephalus sagittarius Rib., which was so far only known from southwestern Europe, and the very rare Edwardsiana smreczynskii Dwor., which is known only from less than 10 localities in Poland, Slowakia, Germany, Switzerland and France. Two further species, which are known in Germany only from this locality, notably Cixidia pilatoi D'Urso & Gugl. and Phlepsius intricatus (H.-S.), were no more found

Calamister cougalensis Löcker, 2014, sp. nov.

<i>Calamister cougalensis</i> sp. nov. <p>(Figs 1, 3, 5, 6 D–F)</p> <p>Zoobank Registration: http://zoobank.org/urn:lsid:zoobank.org:act:826CB89F-150F-472A-A21F-8E8A58 E57891</p> <p> <b>Types.</b> <i>Holotype</i>, ♂, <b>AUSTRALIA, Qld:</b> Cougal Creek, Upper Tallebudgera, 80m, 28°12’S 153°21’E, rainforest, 12–14.x.1990 (G. Daniels) (UQIC55277 QM). <i>Paratypes</i>, <b>Qld:</b> 1 ♀, same data as holotype (QM); 1 ♂, Nagarijoon Falls, Lamington Nat. Park, S. Qld, 28.x.1955 (J. Stoodley) (QM); 1 ♀ and 1 ♂ or ♀ (abdomen missing, body length indicates female), Lamington Nat. Park, 29.x.1955 (F.A. Perkins) (QM).</p> <p> <b>Etymology.</b> Named after the type locality, Cougal Creek.</p> <p> <b>Diagnosis.</b> This species can be distinguished from <i>C. obscurus</i> by the presence of darkened areas around the pterostigma, along the hind margin of the forewing and in most cases on the mesonotum, a brown pterostigma and a phallotheca bearing two spines ventrally to left-laterally, one of which is almost twice as long as the other.</p> <p> <b>Colour.</b> Body light brownish-yellow (testaceous), sometimes with an orange or greenish tinge; lateral parts of mesonotum, abdominal sternites, eyes and antennae slightly to distinctly darker. Forewing hyaline with testaceous veins and tubercles; pterostigma mid to dark brown; posterior margin including apical veins and cells darker i.e. distad of apex of clavus.</p> <p> <b>Morphology.</b> Body length: ♂ 4.7–5.3 mm; ♀ 5.2–5.7 mm.</p> <p>Head: Vertex 1.5–1.8 x wider than long; apical carina v-shaped, subapical carina u- or v-shaped; median carina of vertex absent or if present, covering up ½ of basal compartment of vertex; basal compartment 2.3–5.3 x longer than apical compartment. Frons 1.4–2.0 x longer than wide. Postclypeus with median carina evanescent; lateral carinae absent or evanescent. Anteclypeus with median carina absent or evanescent; lateral carinae absent. Rostrum very short, reaching mid femur but not hind coxa; apical segment of rostrum shorter than subapical segment.</p> <p>Thorax: Forewing 3.3–3.4 x longer than wide; costal margin with about 3–7 tubercles; Sc+R+M near basal cell fused, forming a minute common stem Sc+R+M or Sc+R fused, M emerging separately from basal cell; fork of ScRA+ RP distad or at same level as fork CuA1+CuA2; position of r-m basad of fork MA+MP; MP apically bifid; vein delimiting subapical cell C4 distinctly distad of vein delimiting C5; subapical cell C5 shorter than C4; CuA2 ending well before it reaches margin of wing; 8 apical cells; 5 anteapical cells. Hind leg: 1st tarsomere with 7 apical teeth and no platellae; 2nd tarsomere with 8 apical teeth and 4 platellae.</p> <p>Male genitalia: Anal tube as in Figs 5 C,D. Pygofer and genital styles as in Figs 5 E,F. Aedeagus (Figs 5 A,B): Phallotheca ventrally to left-laterally with two large, curved spines, unequal in length, with spine (b) almost twice as long as spine (a); spine (a) arising near apex of aedeagus and spine (b) arising right underneath spine (a). Flagellum unarmed.</p> <p>Female genitalia: Ovipositor, wax plate and anal tube as in Figs 6 D–F: wax plate circular to oval shaped; anal tube trapezoid, apex much wider than base.</p>Published as part of <i>Löcker, Birgit, 2014, Taxonomic revision of Calamister Kirkaldy, 1906 (Fulgoromorpha: Cixiidae) with the description of a new species, pp. 89-96 in Zootaxa 3878 (1)</i> on pages 93-95, DOI: 10.11646/zootaxa.3878.1.7, <a href="http://zenodo.org/record/287356">http://zenodo.org/record/287356</a&gt

Aka sorellensis Löcker, 2015, sp. nov.

<i>Aka sorellensis</i> sp. nov. <p>(Figs 5 D–F, 15)</p> <p>Zoobank Registration: http://zoobank.org/ urn:lsid:zoobank.org:act:2256C517-8BCD-4962-9E45-F97059887F91</p> <p> <b>Types.</b> <i>Holotype</i>, ♂, <b>AUSTRALIA, Tas:</b> Port Sorell, 41.09S, 146.31E, tall wet heath, 4644.602.443, 6.xi.1990 (L. Hill) (TAIC 145564).</p> <p> <b>Etymology.</b> Named after the type locality, Port Sorell.</p> <p> <b>Diagnosis.</b> <i>Aka sorellensis</i> and <i>A. dobsonensis</i> are the only Australian species of <i>Aka</i> with a deeply excavated apex of the anal tube. These two species differ in the number of apical teeth on the first tarsomere (<i>A. sorellensis</i> with 7, <i>A. dobsonensis</i> with 6).</p> <p> <b>Colour.</b> Face dark brown; paler near vertex; frons with a whitish spot at each lateral end of frontoclypeal suture; lateral carinae of postclypeus whitish near anteclypeus. Vertex, pro- and mesonotum with a creamy, whitish band that runs across most of the width of vertex and the central part of pro- and mesonotum, lateral parts dark brown. Forewings light brown to whitish, mottled with darker spots; tubercles and veins concolorous with cells; pterostigma whitish. Legs mid brown with a few paler marks.</p> <p> <b>Morphology.</b> Body length: ♂ 4.3 mm.</p> <p>Head: Vertex 1.4 x wider than long; median carina of vertex covering about 1/2 of basal compartment of vertex. Frons 1.1 x longer than wide; maximum width no more than 2x apical width; position of maximum width slightly to moderately dorsad of centre of frontoclypeal suture; median carina forked in basal (near frontoclypeal suture) fourth of frons. Median ocellus present (indistinct). Anteclypeus lacking median carina. Subapical segment of rostrum 1.6 x longer than apical segment.</p> <p>Thorax: Mesonotum without sublateral carinae. Forewing 2.9 x longer than wide; costal margin with a few indistinct tubercles; tubercles never within cells; tubercles very small, concolorous with veins; tubercles not bearing setae; Sc+R+M near basal cell fused, forming a minute common stem Sc+R+M; fork of ScRA+ RP distinctly basad of fork CuA1+CuA2; distance tegula to ScR+M fork about as long as distance between this fork and ScRA+ RP fork; position of r-m distinctly basad of fork MA+MP; fork MA+MP situated around apical 1/8 of forewing; icu distinctly distad of apex of clavus; RP apically bifid; MA unforked; MP unforked; nodus of y-vein distinctly distad of centre of clavus; y-vein with A1 moderately elevated; 6 apical cells. Hind leg: 1st tarsomere with 7 apical teeth; 2nd tarsomere with 6 apical teeth and 3 fine setae.</p> <p>Male genitalia: Anal tube apically deeply excavated, forming two separate apical lobes as in Figs 15 C,D. Pygofer and genital styles as in Figs 15 E–G. Aedeagus (Figs 15 A,B): Phallotheca left laterally with a short, curved spine (a); ventrally with a long spine (b), crossing over from right to left lateral; right lateral with two short, curved spines (c,d). Flagellum unarmed, steadily widening towards apex.</p>Published as part of <i>Löcker, Birgit, 2015, Revision of the Australian species of Aka White, 1879 (Fulgoromorpha: Cixiidae) with the description of a new genus, pp. 199-223 in Zootaxa 3956 (2)</i> on pages 216-217, DOI: 10.11646/zootaxa.3956.2.3, <a href="http://zenodo.org/record/246048">http://zenodo.org/record/246048</a&gt

Aka balma Löcker, 2015, sp. nov

<i>Aka balma</i> sp. nov <p>(Figs 2 A–C, 8)</p> <p>Zoobank Registration: http://zoobank.org/urn:lsid:zoobank.org:act:5A0967D9-98A4-4087-B197-BF3D8924DA17</p> <p> <b>Types.</b> <i>Holotype</i>, ♂, <b>AUSTRALIA, Tas:</b> Big Sassy Creek, pyrethrum knockdown, Sassafras, 12.v.1989 (D. Rounsevell) (ASCU ASCT00179976). <i>Paratypes</i>, <b>Tas:</b> 1 ♂, same as holotype except for site 2 (P. Greenslade & D. Rounsevell) (ASCU); 6 ♂, Big Sassy Creek, u.v.l., 10.iv.1991 (T. Semmens) (TAIC).</p> <p> <b>Etymology.</b> The term „ balma “ means „fork“ in Gooniyandi, an Aboriginal language spoken in Western Australia (Thieberger & McGregor 1994). Named after the forked spine on the phallotheca.</p> <p> <b>Diagnosis.</b> This species can be distinguished from all other Australian <i>Aka</i> by the presence of a bifurcate spine left laterally on the phallotheca.</p> <p> <b>Colour.</b> Vertex mid brown, carinae paler, with a whitish spot where subapical carinae meet lateral borders of vertex. Face light to mid brown, sometimes with a few darker areas; with a whitish spot at each lateral end of frontoclypeal suture. Pro- and mesonotum brown with slightly paler carinae; lateral parts of pro- and mesonotum usually darker. Forewings light brown to whitish, mottled with darker marks; tubercles and veins concolorous with cells; costa with 3 or more mid to dark brown bands; pterostigma whitish. Abdominal sternites mid to dark brown. Legs light brown with a few darker marks.</p> <p> <b>Morphology.</b> Body length: ♂ 4.6–4.9 mm.</p> <p>Head: Vertex 1.2–1.3 x wider than long; median carina of vertex covering ½ to almost full length of basal compartment of vertex. Frons 1.1–1.2 x longer than wide; maximum width no more than 2x apical width; position of maximum width moderately to distinctly dorsad of centre of frontoclypeal suture; median carina forked in basal (near frontoclypeal suture) third to half of frons. Median ocellus absent or present (distinct or indistinct). Anteclypeus lacking median carina. Subapical segment of rostrum 1.6–1.7 x longer than apical segment.</p> <p>Thorax: Mesonotum with distinct sublateral carinae. Forewing 2.9–3.4 x longer than wide; costal margin with a few indistinct tubercles; tubercles never within cells; tubercles very small, concolorous with veins; tubercles not bearing setae; Sc+R+M near basal cell fused, forming a minute common stem; Sc+R+M fork of ScRA+ RP distinctly basad of fork CuA1+CuA2; distance tegula to ScR+M fork longer than distance between this fork and ScRA+ RP fork; position of r-m distinctly basad of fork MA+MP; fork MA+MP situated between apical 1/16 and 1/8 of forewing or M unforked; icu distinctly distad of apex of clavus; RP apically unforked; MA unforked, MP unforked or sometimes M not forked into MA and MP; nodus of y-vein distinctly distad of centre of clavus; y-vein with A1 slightly elevated; 4–5 apical cells. Hind leg: 1st tarsomere with 7 apical teeth; 2nd tarsomere with 7 apical teeth and 4 fine setae.</p> <p>Male genitalia: Anal tube (Figs 8 C,D) in dorsal view rounded, straight or very slightly excavated, starting to form two separate lobes; lobes in lateral view produced into rounded hooks. Pygofer and genital styles as in Figs 8 E–G. Aedeagus (Figs 8 A,B): Phallotheca left laterally with a forked spine (a); right laterally with a very long spine (b), reaching down as far as flagellum and a much shorter spine (c). Flagellum unarmed.</p>Published as part of <i>Löcker, Birgit, 2015, Revision of the Australian species of Aka White, 1879 (Fulgoromorpha: Cixiidae) with the description of a new genus, pp. 199-223 in Zootaxa 3956 (2)</i> on pages 208-209, DOI: 10.11646/zootaxa.3956.2.3, <a href="http://zenodo.org/record/246048">http://zenodo.org/record/246048</a&gt

Aka tasmani Muir 1931

<i>Aka tasmani</i> Muir, 1931 <p>(Figs 6 A–C, 16, 18)</p> <p> <i>Aka tasmani</i> Muir, 1931: 63.</p> <p> <b>Types.</b> <i>Neotype</i>, here designated, ♂, <b>AUSTRALIA, Tas:</b> Hobart, 5.vii. 1913 /43 (G.H. Hardy) (BMNH K45391). <i>Paratypes.</i> <b>AUSTRALIA, Tas:</b> 2 ♂, Hobart, 20.vii. 1913 /39 (G.H. Hardy) (BMNH K45388) (originally on one mount, now mounted on two separate mounts).</p> <p> <b>Other material examined. AUSTRALIA, Tas:</b> 1 ♂, Thumbs Parallel Gullies, Sassafras, outer 3, 1.ix.1989 (H. Elliott) (ASCU).</p> <p> <b>Notes:</b> Muir (1931) based this species on 4 females and 2 males from Hobart, Tasmania (G.H. Hardy, May, July, 1913) with „ type in Australian Museum, paratype in British Museum“.Three paratypes were found in the BMNH (Fig. 18). Enquiries for the types in the AMS and BPBM were unsuccessful which means part of the type series (3 specimens), including the holotype, is lost. Interestingly all of the paratypes in the BMNH are males, although Muir stated he had only 2 males. Mick Webb (BMNH) was able to match the paratypes with illustrations and the description given in this paper of a specimen from Thumbs Parallel Gully. A neotype has been designated to provide a diagnostic reference for the species.</p> <p>Muir 1931 lists another female from the same location as the types, which was slightly larger and may also represent this species. This specimen could not be found and must have been lost together with parts of the type series.</p> <p> <b>Diagnosis.</b> This species can be distinguished from all other Australian <i>Aka</i> by the following combination of characters: first tarsomere with 7 and second tarsomere with 6–7 apical teeth; lobes of male anal tube in lateral view produced into hooks; aedeagus right lateral with a medium sized spine (about half as long as flagellum) with its tip directed towards the flagellum; absence of a bifurcate spine on the aedeagus.</p> <p> <b>Colour.</b> Vertex mid brown, with a whitish spot where subapical carinae meet lateral borders of vertex and near basal emargination. Face mid brown with a whitish spot at each lateral end of frontoclypeal suture; lateral carinae of postclypeus whitish near anteclypeus. Pro- and mesonotum brown with slightly paler carinae. Forewings light brown to whitish, with a few darker marks; costa with several mid brown bands; tubercles concolorous with cells; veins in general slightly darker than cells; pterostigma whitish. Legs light brown with a few darker marks.</p> <p> <b>Morphology.</b> Body length: ♂ 4.4 mm.</p> <p>Head: Vertex 1.1 x wider than long; median carina of vertex covering about 3/4 of basal compartment of vertex. Frons 1.1 x longer than wide; maximum width no more than 2x apical width; position of maximum width moderately dorsad of centre of frontoclypeal suture; median carina forked in basal (near frontoclypeal suture) third of frons. Median ocellus present. Anteclypeus lacking median carina. Subapical segment of rostrum 1.5 x longer than apical segment.</p> <p>Thorax: Mesonotum with distinct sublateral carinae. Forewing 3.3 x longer than wide; costal margin with a several indistinct tubercles; tubercles never within cells; tubercles very small, concolorous with veins; tubercles not bearing setae; Sc+R+M near basal cell fused, forming a minute common stem; Sc+R+M fork of ScRA+ RP distinctly basad of fork CuA1+CuA2; distance tegula to ScR+M fork much longer than distance between this fork and ScRA+ RP fork; icu distinctly distad of apex of clavus; RP apically unforked; M not forked into MA and MP; nodus of y-vein slightly to moderately distad of centre of clavus; y-vein with A1 slightly elevated; 4 apical cells. Hind leg: 1st tarsomere with 7 apical teeth; 2nd tarsomere with 6–7 apical teeth and 3 fine setae.</p> <p>Male genitalia: Anal tube (Figs 16 C,D, 18E,F) apically slightly excavated, starting to form two separate apical lobes; lobes in lateral view produced into hooks. Pygofer and genital styles as in Figs 16 E–G, 18E,F. Aedeagus (Figs 16 A,B, 18E,F): Phallotheca left laterally with a curved spine (a), right laterally with two spines (b,c). Flagellum unarmed, about the same width throughout.</p> <p> <b>Remarks.</b> The male genitalia of the specimen from Thumbs Parallel Gullies matches those illustrated by Muir (1931) of <i>A. tasmani</i>, apart from a few minor details discussed below. Further it matches the description apart from the following character states. Muir lists, “No spines on the hind tibiae”. The specimen examined showed 3–4 medium sized lateral spines. The colouration deviates slightly from the description, with lighter coloured areas (marks) situated near the frontoclypeal suture, at the posterior end of the vertex and the anterior part of the pronotum. However, the light basal half of the costa, with three or four lighter marks on the apical half of the costal margin as described by Muir, can be observed in the specimen from Thumbs Parallel Gullies.</p> <p>There is a certain degree of variation in the curvature, length and position of spines on the aedeagus. In two out of three specimens in the type series spines (b) and (c) are more pincer-like (Fig. 18 E) than in the third specimen (Fig. 18 F) and in the non-type-specimen from Thumbs Parallel Gully (Fig. 16 B) (pers. comm. Mick Webb, 2014). Because the pincer-like condition more closely resembles that of Muir’s illustration the neotype has been chosen from those two paratypes. Spine (b) is inserted closer to the ventral side in one of the paratypes (Fig. 18 F) than in the neotype (Fig. 18 E) and other paratype. The specimen from Thumbs Parallel Gully (Fig. 16 B) shows an intermediate state.</p> <p> The ASCU collection holds a specimen, collected on the 4 April, 1989 in yellow pan traps at Mt Mangana on Bruny Island by J. Diggle and P. Greenslade, that has a configuration of aedeagal spines similar to <i>A. tasmani</i> and <i>A. hardyi</i>. Further it shares the presence of 7 apical teeth on the first and second tarsomere with the specimen from Thumbs Parallel Gullies. However, it differs from that one, and the original description of <i>A. tasmani</i>, in the absence of well developed triangular spines on the anal tube (the lobes are only slightly produced), the forking of M, the evanescent sublateral (medio-lateral) carinae on the mesonotum and the colouration. The specimen from Bruny Island is very pale in colour with clearly contrasting light and dark brown marks. It has very distinct dark tubercles which are not present in the Thumbs Parallel Gullies specimen and which are not mentioned in Muir’s description. Further studies are needed to clarify whether this specimen belongs to <i>A. tasmani</i>, <i>A. hardyi</i> or represents a new species closely related to those two.</p>Published as part of <i>Löcker, Birgit, 2015, Revision of the Australian species of Aka White, 1879 (Fulgoromorpha: Cixiidae) with the description of a new genus, pp. 199-223 in Zootaxa 3956 (2)</i> on pages 217-219, DOI: 10.11646/zootaxa.3956.2.3, <a href="http://zenodo.org/record/246048">http://zenodo.org/record/246048</a&gt

Aka kuraka Löcker, 2015, sp. nov.

<i>Aka kuraka</i> sp. nov. <p>(Figs 4 D–G, 7B,C, 13)</p> <p>Zoobank Registration: http://zoobank.org/ urn:lsid:zoobank.org:act:1BD1C04B-C09B-4785-A647-002DDEED8777</p> <p> <b>Types.</b> <i>Holotype</i>, ♂, <b>AUSTRALIA, Tas:</b> Balfour Track F.R., 41.08S, 144.57E, 80m, uvl, <i>Nothofagus</i>, 8.xi.1992 (P.B. McQuillan) (TAIC 145563). <i>Paratypes</i>, <b>Tas:</b> 1 ♂, 2km W Commonwealth Hill, via Rennison Bell, 40.49S, 145.24E, ex ethanol, 9.xii.1981 (I.D. Naumann) (ANIC).; 1 ♀, Corinna, UVL, 11.iv.1986 (L. Hill) (TAIC); 1 ♂, Waratah (Lea) (SAMA).</p> <p> <b>Etymology.</b> The term „ kuraka “ means „duck“ in Nyungar, an Aboriginal language spoken in Western Australia (Thieberger & McGregor 1994). Named after the shape of the genital style in lateral view which resembles the head of a duck.</p> <p> <b>Diagnosis.</b> This species can be distinguished from all other Australian species of <i>Aka</i> by the angle formed by the caudal border of the vertex, which is rectangular to acute in <i>A. kuraka</i>, as opposed to broadly obtuse.</p> <p> <b>Colour.</b> Vertex light brown, carinae paler, disc and other concave areas between apical and subapical carinae often dark brown; sometimes with a whitish spot where subapical carinae meet lateral borders of vertex. Face light to mid brown; carinae concolorous with face; frons with a whitish spot at each lateral end of frontoclypeal suture. Pro- and mesonotum brown with slightly paler carinae; lateral parts of pro- and mesonotum darker. Forewings light brown to whitish, with a few mid to dark brown marks; 3 or more darker bands on costa; usually with 2–3 darker marks in clavus; tubercles dark, in distinct contrast to cells; veins concolorous with cells, sometimes slightly darker; pterostigma whitish. Abdominal sternites mid to dark brown. Legs light to mid brown sometimes with a few darker marks.</p> <p> <b>Morphology.</b> Body length: ♂ 5.3–5.4 mm; ♀ 5.7 mm.</p> <p>Head: Vertex 0.9–1.0 x as wide as long; median carina of vertex covering about 1/3 of basal compartment of vertex. Frons 1.4 x longer than wide; maximum width more than 2x apical width, steadily broadening; position of maximum width distinctly dorsad of centre of frontoclypeal suture; median carina forked in basal (near frontoclypeal suture) fourth of frons. Median ocellus absent or present (indistinct). Anteclypeus with very indistinct median carina. Subapical segment of rostrum 2.0 x longer than apical segment.</p> <p>Thorax: Mesonotum with or without indistinct sublateral carinae. Forewing 3.2–3.5 x longer than wide; costal margin with 13–18 distinct tubercles; tubercles aligned alongside veins, but reaching to some extent into cell area; tubercles dark, underlaid by a larger dark circle, therefore appearing large and distinctly in contrast to cells; tubercles not bearing setae; Sc+R fused, M emerging separately from basal cell; fork of ScRA+ RP distinctly basad of fork CuA1+CuA2; distance tegula to ScR+M fork distinctly (about ½ to ¾) shorter than distance between this fork and ScRA+ RP fork; position of r-m distinctly to moderately basad of fork MA+MP; fork MA+MP situated between apical 1/8 and 1/3 of forewing; icu at same level or slightly distad of apex of clavus; RP apically bifid; MA bifid; MP bifid; nodus of y-vein distinctly distad of centre of clavus; y-vein with A1 slightly elevated; 6–9 apical cells. Hind leg: 1st tarsomere with 6 apical teeth; 2nd tarsomere with 6 apical teeth and 3 fine setae.</p> <p>Male genitalia: Anal tube (Figs 13 C,D) apically slightly indented, forming two rounded lobes. Pygofer as in Fig. 13 E. Genital styles apically rounded, laterally produced resembling the beak of a duck as in Figs 13 F,G. Aedeagus (Figs 13 A,B): Phallotheca left laterally with a strongly curved spine (a); ventrally with a thick, long spine (b); right laterally with a short, straight spine (c). Base of phallotheca ventrally with 2 small sclerotised teeth. Flagellum unarmed, near base laterally extended.</p> <p> <b>Remarks.</b> <i>Aka kuraka</i> and <i>A. issidopsis</i> share the same lateral extension near the base of the flagellum, however in <i>A. issidopsis</i> the flagellum is widened twice as much as in <i>A. kuraka</i>.</p>Published as part of <i>Löcker, Birgit, 2015, Revision of the Australian species of Aka White, 1879 (Fulgoromorpha: Cixiidae) with the description of a new genus, pp. 199-223 in Zootaxa 3956 (2)</i> on pages 214-215, DOI: 10.11646/zootaxa.3956.2.3, <a href="http://zenodo.org/record/246048">http://zenodo.org/record/246048</a&gt

Yanganaka ariecornia Löcker, 2015, sp. nov

<i>Yanganaka ariecornia</i> sp. nov <p>(Figs 6 D–F, 17)</p> <p>Zoobank Registration: http://zoobank.org/ urn:lsid:zoobank.org:act:A0256F01-D5B6-44BA-BADE-A86E9DCD90BD</p> <p> <b>Types.</b> <i>Holotype</i>, ♂, <b>AUSTRALIA, Tas:</b> McPartlan Pass, 5256300N 0438000E, +/- 25m, Grid/Sample: ARE 2, Method: S, 10.viii.1999 (M. Driessen) (AMS K.437130). <i>Paratypes</i>, <b>Tas:</b> 1 ♂, same data as holotype except for Grid/Sample: ARW6, Method: P, 23.vii.2000.</p> <p> <b>Etymology.</b> The Latin term „aries“ means „ram“ and „cornus“ means „horn“. Named after a thick spine on the aedeagus which resembles the horn of a ram.</p> <p> <b>Colour.</b> Vertex mid to dark brown, carinae sometimes paler. Face including carinae mid brown; slightly darker near carinae. Pro- and mesonotum displaying various shades of brown with paler carinae. Forewings light brown to whitish, with a dark brown spot anterior to pterostigma; tubercles and veins dark, in distinct contrast to cells; pterostigma whitish. Abdominal sternites light to mid brown. Legs light brown sometimes with a few darker marks.</p> <p> <b>Morphology.</b> Body length: ♂ 4.1–4.2 mm.</p> <p>Head: Vertex 1.1–1.2 x wider than long; median carina of vertex covering full length of basal compartment of vertex; angle formed by caudal border of vertex broadly obtuse. Frons 1.3 x longer than wide; maximum width of frons no more than 2 x apical width; position of maximum width around frontoclypeal suture or slightly dorsad of centre of frontoclypeal suture; median carina forked in basal (near frontoclypeal suture) third to half of frons. Median ocellus absent. Anteclypeus with very indistinct median carina. Head including eyes slightly narrower than pronotum. Subapical segment of rostrum 1.7–1.9 x longer than apical segment.</p> <p>Thorax: Mesonotum with distinct or indistinct sublateral carinae. Forewing short, curved to fit body, only slightly raised in posterior part of thorax. Forewing 2.8 x longer than wide; costal margin with 12–14 very distinct tubercles; tubercles aligned alongside veins, but reaching to some extent into cell area; tubercles dark, some underlaid by a larger dark circle, therefore appearing large and distinctly in contrast to cells; tubercles not bearing setae; Sc+R+M near basal cell fused, forming a minute common stem Sc+R+M or Sc+R fused, M emerging separately from basal cell; fork of ScRA+ RP about same level as fork CuA1+CuA2; distance tegula to ScR+M fork distinctly (about ½ to ¾) shorter than distance between this fork and ScRA+ RP fork; position of r-m distinctly basad of fork MA+MP; fork MA+MP situated between apical 1/8 and 1/4 of forewing; icu at same level or slightly distad of apex of clavus; RP apically bifid or trifid; MA unforked, bifid of trifid; MP unforked or bifid; nodus of yvein more or less in centre of clavus; y-vein with A1 only very slightly elevated; 7–9 apical cells. Hind leg: 1st tarsomere with 8 apical teeth and no platellae but with 4 fine setae; 2nd tarsomere with 7–8 apical teeth and no platellae but with 4–5 fine setae.</p> <p>Male genitalia: Anal tube rounded, forming only one lobe as in Figs 17 C,D. Pygofer and genital styles as in Figs 17 E–G. Aedeagus (Figs 17 A,B): Phallotheca left laterally with an almost straight spine (a), ventrally with a very thick, curved spine (b) and right laterally with a thick, s-shaped spine (c). Flagellum unarmed, pear-shaped.</p> <p> <b>Remarks.</b> <i>Yanganaka ariecornia</i> slightly resembles <i>A. issidopsis</i> in the shape and curvature of the forewing. However, in <i>Y. ariecornia</i> the forewing is much less raised than in <i>A. issidopsis</i>.</p>Published as part of <i>Löcker, Birgit, 2015, Revision of the Australian species of Aka White, 1879 (Fulgoromorpha: Cixiidae) with the description of a new genus, pp. 199-223 in Zootaxa 3956 (2)</i> on pages 220-221, DOI: 10.11646/zootaxa.3956.2.3, <a href="http://zenodo.org/record/246048">http://zenodo.org/record/246048</a&gt

Calamister obscurus Kirkaldy

<i>Calamister obscurus</i> Kirkaldy <p>(Figs 2, 4, 6 A–C)</p> <p> <i>Calamister obscurus</i> Kirkaldy, 1906: 403.</p> <p> <b>Types.</b> <i>Lectotype</i> (designated here, examined), ♂, <b>AUSTRALIA, Qld:</b> Bundaberg, ix.–xii.1904 (Koebele) (BPBM). <i>Paralectotypes</i>, 2 ♀, same data as holotype (BPBM).</p> <p> <b>Notes:</b> The original description does not mention the designation of a holotype, therefore these specimens are regarded as syntypes. A lectotype is designated to provide a diagnostic reference for the species.</p> <p> <b>Other material examined. AUSTRALIA, Qld:</b> 11 ♂, 6 ♀, 1 ♂ or ♀ (abdomen missing, body length indicates male), Bundaberg, 20.viii.1919 (F. Muir) (BPBM); 1 ♂, 1 ♀, Coolum, 22.ix.1956 (Kirkpolnik) (QM). <b>NSW:</b> 2 ♀, Bullsground [state forest], Port Macquarie, beating understorey plants, xi. 2005 (M.L. Moir MLM01135-6).</p> <p> <b>Diagnosis.</b> This species can be distinguished from <i>C. cougalensis</i> by its colouration which is entirely yellow with hyaline forewings and a whitish or yellowish pterostigma. Further the phallotheca bears two spines ventrally to right-laterally, about equal in length.</p> <p> <b>Colour.</b> Body light brownish-yellow (testaceous), eyes and antennae darker; rarely lateral parts of mesonotum slightly darker. Forewing hyaline with testaceous veins and tubercles; pterostigma testaceous.</p> <p> <b>Morphology.</b> Body length: ♂ 4.3–4.5 mm; ♀ 4.7–5.1 mm.</p> <p>Head: Vertex 1.5–1.7 x wider than long; apical carina v-shaped, subapical carina u- shaped; median carina of vertex absent or if present, covering up ¼ of basal compartment of vertex; basal compartment 3.7–9.0 x longer than apical compartment. Frons 1.1–1.4 x longer than wide. Postclypeus with median carina absent or evanescent; lateral carinae absent. Anteclypeus with median and lateral carinae absent. Rostrum very short, reaching mid coxa or mid femur; apical segment of rostrum shorter than or as long as subapical segment.</p> <p>Thorax: Forewing 2.7–3.5 x longer than wide; costal margin with about 1–4 tubercles; Sc+R+M near basal cell fused, forming a minute common stem Sc+R+M; fork of ScRA+ RP basad or at same level as fork CuA1+CuA2; position of r-m at same level as or slightly basad of fork MA+MP; MP apically unforked or bifid; vein delimiting subapical cell C4 more or less at same level as vein delimiting C5; subapical cell C5 distinctly longer than C4; CuA2 ending well before it reaches margin of wing or fading (reaching margin of wing with reduced thickness); 7–8 apical cells; 3 anteapical cells. Hind leg: 1st tarsomere with 7 apical teeth (rarely 6 or 8) and no platellae; 2nd tarsomere with 8 (rarely 9) apical teeth and 4 (rarely 5) platellae (Fig. 2 E).</p> <p>Male genitalia: Anal tube as in Figs 4 C,D. Pygofer and genital styles as in Figs 4 E,F. Aedeagus (Figs 4 A,B): Phallotheca ventrally to right-laterally with two large, curved spines (a, b), about equal in length; spine (a) arising near apex of aedeagus and spine (b) arising right underneath spine (a). Flagellum unarmed.</p> <p>Female genitalia: Ovipositor, wax plate and anal tube as in Figs 6 A–C: wax plate oval-shaped; anal tube trapezoid, widening towards apex.</p> <p> <b>Remarks.</b> In general <i>C. obscurus</i> has 7 apical teeth and no platellae on the 1st tarsomere and 8 apical teeth and 4 platellae on the 2nd tarsomere. The following deviations from this formula appear to be aberrations as they only occurred on one leg of the specimen: One male from Bundaberg had only 6 and another male from the same location 8 apical teeth on the 1st tarsomere; the female from Coolum had 9 apical teeth and 5 platellae on the 2nd tarsomere.</p> <p> Kirkaldy’s original descriptions of the genus and species are not very specific and lack illustrations. Muir (1908) provided a dichotomous key to the genera of Cixiidae which includes <i>Calamister</i>. Based on this identification key Jacobi (1928) identified a female specimen from the 'Blackal Range’ (this appears to be the ‘Blackall Range’ in South East Queensland) to be <i>C. obscurus</i>. Jacobi mentions features such as a minute stem of Sc+R+M; a long common stem of Sc+R and the fact that fork Sc+R and fork CuA1+CuA2 are at about the same level, situated in the basal half of the forewing; and transverse carina of vertex slightly u-shaped. These features can be found in the type specimens of <i>C. obscurus</i> and/or the other material examined. Jacobi (1928) noted that apical veins (towards the margin) and crossveins were darker brown in the Blackall Range specimen. In <i>C. cougalensis</i> sp. nov. the apical veins are darker but not the crossveins. In <i>C. obscurus</i> all veins are uniformously coloured throughout (Figs 2 A,B). Jacobi included an illustration of the forewing in his redescription of <i>C. obscurus</i> Kirkaldy. The specimen depicted varies considerably from the wing venation found in <i>Calamister</i>, e.g. the specimen pictured has a trifid RP, MA and MP, however in <i>Calamister</i> these veins are bifid (Figs 1, 3 A), or even in some cases MP unforked (Fig. 2 B). Further, Jacobi’s specimen shows CuA1 and CuA2 rejoining which does not occur in any of the types. This means there is considerable doubt that the specimen from the Blackall Range is <i>C. obscurus</i> —examination of the specimen is needed to clarify this.</p>Published as part of <i>Löcker, Birgit, 2014, Taxonomic revision of Calamister Kirkaldy, 1906 (Fulgoromorpha: Cixiidae) with the description of a new species, pp. 89-96 in Zootaxa 3878 (1)</i> on pages 91-93, DOI: 10.11646/zootaxa.3878.1.7, <a href="http://zenodo.org/record/287356">http://zenodo.org/record/287356</a&gt

Innobindus multimaculatus Jacobi 1928

<p> <i>multimaculatus group</i></p> <p> Vertex lacking median carina; lateral carinae of frons strongly elevated (distinctly higher elevated than median carina of frons); lateral carinae of frons continuous with those of clypeus (Fig. 6C; Löcker <i>et al.</i> 2007: figs 3c, 4b, f, 5c); phallotheca ventrally with a fringed ridge or a bifurcate ventral process; phallotheca laterally with medium-sized or very long spine; flagellum with or without spines.</p>Published as part of <i>Löcker, Birgit, 2023, Six new species in the genus Innobindus Jacobi 1928 (Hemiptera: Auchenorrhyncha: Fulgoromorpha: Cixiidae: Brixiini), pp. 108-134 in European Journal of Taxonomy 908</i> on page 123, DOI: 10.5852/ejt.2023.908.2341, <a href="http://zenodo.org/record/10184129">http://zenodo.org/record/10184129</a&gt