22 research outputs found

    Predominant-Period Site Classification for Response Spectra Prediction Equations in Italy

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    Abstract We propose a site classification scheme based on the predominant period of the site, as determined from the average horizontal-to-vertical (H/V) spectral ratios of ground motion. Our scheme extends Zhao et al. (2006) classifications by adding two classes, the most important of which is defined by flat H/V ratios with amplitudes less than 2. The proposed classification is investigated by using 5%-damped response spectra from Italian earthquake records. We select a dataset of 602 three-component analog and digital recordings from 120 earthquakes recorded at 214 seismic stations within an hypocentral distance of 200 km. Selected events are in the moment-magnitude range 4.0 ≤ Mw ≤ 6.8 and focal depths from a few kilometers to 46 km. We computed H/V ratios for these data and used these to classify each site into one of six classes. We then investigate the impact of this classification scheme on empirical ground-motion prediction equations by comparing its performance with that of the conventional rock/soil classification. Although the adopted approach results in a only a small reduction of overall standard deviation, the use of H/V spectral ratios in site classification does capture the signature of sites with flat frequency-response, well as deep and shallow soil profiles, characterized 2 C:\di_alessandro\site_classification_italy\final_version_of_paper\bssa-d-11-00084_di_alessandro_etal_final_revisions.doc by long-and short-period resonance, respectively; in addition, the classification scheme is relatively quick and inexpensive, which is an advantage over schemes based on measurements of shear-wave velocity

    The mitochondrial genome of Sinentomon erythranum (Arthropoda: Hexapoda: Protura): an example of highly divergent evolution

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    <p>Abstract</p> <p>Background</p> <p>The phylogenetic position of the Protura, traditionally considered the most basal hexapod group, is disputed because it has many unique morphological characters compared with other hexapods. Although mitochondrial genome information has been used extensively in phylogenetic studies, such information is not available for the Protura. This has impeded phylogenetic studies on this taxon, as well as the evolution of the arthropod mitochondrial genome.</p> <p>Results</p> <p>In this study, the mitochondrial genome of <it>Sinentomon erythranum </it>was sequenced, as the first proturan species to be reported. The genome contains a number of special features that differ from those of other hexapods and arthropods. As a very small arthropod mitochondrial genome, its 14,491 nucleotides encode 37 typical mitochondrial genes. Compared with other metazoan mtDNA, it has the most biased nucleotide composition with T = 52.4%, an extreme and reversed AT-skew of -0.351 and a GC-skew of 0.350. Two tandemly repeated regions occur in the A+T-rich region, and both could form stable stem-loop structures. Eighteen of the 22 tRNAs are greatly reduced in size with truncated secondary structures. The gene order is novel among available arthropod mitochondrial genomes. Rearrangements have involved in not only small tRNA genes, but also PCGs (protein-coding genes) and ribosome RNA genes. A large block of genes has experienced inversion and another nearby block has been reshuffled, which can be explained by the tandem duplication and random loss model. The most remarkable finding is that <it>trnL2(UUR) </it>is not located between <it>cox1 </it>and <it>cox2 </it>as observed in most hexapod and crustacean groups, but is between <it>rrnL </it>and <it>nad1 </it>as in the ancestral arthropod ground pattern. The "<it>cox1</it>-<it>cox2</it>" pattern was further confirmed in three more representative proturan species. The phylogenetic analyses based on the amino acid sequences of 13 mitochondrial PCGs suggest <it>S</it>. <it>erythranum </it>failed to group with other hexapod groups.</p> <p>Conclusions</p> <p>The mitochondrial genome of <it>S. erythranum </it>shows many different features from other hexapod and arthropod mitochondrial genomes. It underwent highly divergent evolution. The "<it>cox1</it>-<it>cox2</it>" pattern probably represents the ancestral state for all proturan mitogenomes, and suggests a long evolutionary history for the Protura.</p

    Predominant-Period Site Classification for Response Spectra Prediction Equations in Italy

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    We propose a site-classification scheme based on the predominant period of the site, as determined from the average horizontal-to-vertical (H/V) spectral ratios of ground motion. Our scheme extends Zhao et al. (2006) classifications by adding two classes, the most important of which is defined by flat H/V ratios with amplitudes less than 2. The proposed classification is investigated by using 5%-damped response spectra from Italian earthquake records. We select a dataset of 602 three-component analog and digital recordings from 120 earthquakes recorded at 214 seismic stations within a hypocentral distance of 200 km. Selected events are in the momentmagnitude range 4:0 d Mw d 6:8 and focal depths from a few kilometers to 46 km.We computed H/V ratios for these data and used them to classify each site into one of six classes. We then investigate the impact of this classification scheme on empirical ground-motion prediction equations (GMPEs) by comparing its performance with that of the conventional rock/soil classification. Although the adopted approach results in only a small reduction of the overall standard deviation, the use of H/V spectral ratios in site classification does capture the signature of sites with flat frequency-response, as well as deep and shallow-soil profiles, characterized by long- and short-period resonance, respectively; in addition, the classification scheme is relatively quick and inexpensive, which is an advantage over schemes based on measurements of shearwave velocity

    High divergence across the whole mitochondrial genome in the "pan-Antarctic" springtail Friesea grisea: Evidence for cryptic species?

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    Collembola are one of the few hexapod groups adapted to live in the harsh environmental conditions of Antarctic terrestrial ecosystems. Diversity is limited to a few species that can be very abundant in coastal deglaciated sites. A remarkable lack of overlap in Collembola species composition is evident between Western and Eastern Antarctica, and Friesea grisea is currently the only species whose distribution is thought to span these two main regions of the continent. However, our analysis of the complete sequences of the mitochondrial genomes from specimens obtained from each of the two regions showed unexpected genetic divergence, well above the average levels observed between populations belonging to the same species, and so indicating that these are actually separate species, despite their lack of distinguishing morphology. Detailed analysis of the two genomes showed the presence of a non-coding region observed between trnS (uga) and nad1. Other features of these mitochondrial genomes, such as base compositional bias, secondary structure features of tRNAs and the presence of regulatory elements in the control region, are described and discussed from an evolutionary standpoint. (C) 2009 Elsevier B.V. All rights reserved

    Response to comment on "Hexapod origins: monophyletic or paraphyletic?"

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    Hexapod origins: Monophyletic or paraphyletic?

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    Recent morphological and molecular evidence has changed interpretations of ar- thropod phylogeny and evolution. Here we compare complete mitochondrial ge- nomes to show that Collembola, a wingless group traditionally considered as basal to all insects, appears instead to constitute a separate evolutionary lineage that branched much earlier than the separation of many crustaceans and insects and independently adapted to life on land. Therefore, the taxon Hexapoda, as com- monly defined to include all six-legged arthropods, is not monophyletic

    Domestication of olive fly through a multi-regional host shift to cultivated olives: Comparative dating using complete mitochondrial genomes

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    The evolutionary history of the olive fly, Bactrocera oleae, was reconstructed in a phylogenetic and coa- lescent framework using full mitochondrial genome data from 21 individuals covering the entire world- wide distribution of the species. Special attention was given to reconstructing the timing of the processes under study. The early subdivision of the olive fly reflects the Quaternary differentiation between Olea europea subsp. europea in the Mediterranean area and the two lineages of Olea europea subsp. cuspidata in Africa and Asia, pointing to an early and close association between the olive fly and its host. The geo- graphic structure and timing of olive fly differentiation in the Mediterranean indicates a clear connection with the post-glacial recolonization of wild olives in the area, and is irreconcilable with the early histor- ical process of domestication and spread of the cultivated olive from its Levantine origin. Therefore, we suggest an early co-history of the olive fly with its wild host during the Quaternary and post-glacial peri- ods and a multi-regional shift of olive flies to cultivated olives as these cultivars gradually replaced wild olives in historical times
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