Molecular phylogeny of brachiopods and phoronids based on nuclear-encoded small subunit ribosomal RNA gene sequences

Brachiopod and phoronid phylogeny is inferred from SSU rDNA sequences of 28 articulate and nine inarticulate brachiopods, three phoronids, two ectoprocts and various outgroups, using gene trees reconstructed by weighted parsimony, distance and maximum likelihood methods. Of these sequences, 33 from brachiopods, two from phoronids and one each from an ectoproct and a priapulan are newly determined. The brachiopod sequences belong to 31 different genera and thus survey about 10% of extant genus-level diversity. Sequences determined in different laboratories and those from closely related taxa agree well, but evidence is presented suggesting that one published phoronid sequence (GenBank accession UO12648) is a brachiopod-phoronid chimaera, and this sequence is excluded from the analyses. The chiton, Acanthopleura, is identified as the phenetically proximal outgroup; other selected outgroups were chosen to allow comparison with recent, non-molecular analyses of brachiopod phylogeny. The different outgroups and methods of phylogenetic reconstruction lead to similar results, with differences mainly in the resolution of weakly supported ancient and recent nodes, including the divergence of inarticulate brachiopod sub-phyla, the position of the rhynchonellids in relation to long- and short-looped articulate brachiopod clades and the relationships of some articulate brachiopod genera and species. Attention is drawn to the problem presented by nodes that are strongly supported by non-molecular evidence but receive only low bootstrap resampling support. Overall, the gene trees agree with morphology-based brachiopod taxonomy, but novel relationships are tentatively suggested for thecideidine and megathyrid brachiopods. Articulate brachiopods are found to be monophyletic in all reconstructions, but monophyly of inarticulate brachiopods and the possible inclusion of phoronids in the inarticulate brachiopod clade are less strongly established. Phoronids are clearly excluded from a sister-group relationship with articulate brachiopods, this proposed relationship being due to the rejected, chimaeric sequence (GenBank UO12648). Lineage relative rate tests show no heterogeneity of evolutionary rate among articulate brachiopod sequences, but indicate that inarticulate brachiopod plus phoronid sequences evolve somewhat more slowly. Both brachiopods and phoronids evolve slowly by comparison with other invertebrates. A number of palaeontologically dated times of earliest appearance are used to make upper and lower estimates of the global rate of brachiopod SSU rDNA evolution, and these estimates are used to infer the likely divergence times of other nodes in the gene tree. There is reasonable agreement between most inferred molecular and palaeontological ages. The estimated rates of SSU rDNA sequence evolution suggest that the last common ancestor of brachiopods, chitons and other protostome invertebrates (Lophotrochozoa and Ecdysozoa) lived deep in Precambrian time. Results of this first DNA-based, taxonomically representative analysis of brachiopod phylogeny are in broad agreement with current morphology-based classification and systematics and are largely consistent with the hypothesis that brachiopod shell ontogeny and morphology are a good guide to phylogeny

Deep near-IR observations of the Globular Cluster M4: Hunting for Brown Dwarfs

We present an analysis of deep HST/WFC3 near-IR (NIR) imaging data of the globular cluster M4. The best-photometry NIR colour-magnitude diagram (CMD) clearly shows the main sequence extending towards the expected end of the Hydrogen-burning limit and going beyond this point towards fainter sources. The white dwarf sequence can be identified. As such, this is the deepest NIR CMD of a globular cluster to date. Archival HST optical data were used for proper-motion cleaning of the CMD and for distinguishing the white dwarfs (WDs) from brown dwarf (BD) candidates. Detection limits in the NIR are around F110W approx 26.5 mag and F160W approx27 mag, and in the optical around F775W approx 28 mag. Comparing our observed CMDs with theoretical models, we conclude that we have reached beyond the H-burning limit in our NIR CMD and are probably just above or around this limit in our optical-NIR CMDs. Thus, any faint NIR sources that have no optical counterpart are potential BD candidates, since the optical data are not deep enough to detect them. We visually inspected the positions of NIR sources which are fainter than the H-burning limit in F110W and for which the optical photometry did not return a counterpart. We found in total five sources for which we did not get an optical measurement. For four of these five sources, a faint optical counterpart could be visually identified, and an upper optical magnitude was estimated. Based on these upper optical magnitude limits, we conclude that one source is likely a WD, one source could either be a WD or BD candidate, and the remaining two sources agree with being BD candidates. For only one source no optical counterpart could be detected, which makes this source a good BD candidate. We conclude that we found in total four good BD candidates.Comment: ApJ accepted, 28 pages including 16 figure

The experience of children with disabilities and their families during the COVID-19 pandemic; what lessons can we learn?

COVID-19 has had serious, negative impacts on children’s health and development. But the impacts have not been felt equally; disabled children and their families have been hit particularly hard. In this paper we summarise UK legislation to limit the spread of the virus and describe how services to disabled children changed as a result. We discuss the long-term deleterious impacts of changes in service provision on the health and wellbeing of disabled children and the parent carers supporting them. We close with lessons learned for resetting services to support the ongoing recovery of children and their families and recommendations for delivering services better in future emergencies to ensure that disabled children’s health and wellbeing is maintained

Light scattering from three-level systems: The T-matrix of a point-dipole with gain

We present an extension of the T-matrix approach to scattering of light by a three-level system, using a description based on a Master equation. More particularly, we apply our formalism to calculate the T-matrix of a pumped three-level atom, providing an exact and analytical expression describing the influence of a pump on the light scattering properties of an atomic three-level system

Bond percolation on a class of correlated and clustered random graphs

We introduce a formalism for computing bond percolation properties of a class of correlated and clustered random graphs. This class of graphs is a generalization of the Configuration Model where nodes of different types are connected via different types of hyperedges, edges that can link more than 2 nodes. We argue that the multitype approach coupled with the use of clustered hyperedges can reproduce a wide spectrum of complex patterns, and thus enhances our capability to model real complex networks. As an illustration of this claim, we use our formalism to highlight unusual behaviors of the size and composition of the components (small and giant) in a synthetic, albeit realistic, social network.Comment: 16 pages and 4 figure

Multiplicity of Nearby Free-floating Ultra-cool Dwarfs: a HST-WFPC2 search for companions

We present HST/WFPC2 observations of a sample of 134 ultra-cool objects (spectral types later than M7) coming from the DENIS, 2MASS and SDSS surveys, with distances estimated to range from 7 pc to 105 pc. Fifteen new ultra-cool binary candidates are reported here. Eleven known binaries are confirmed and orbital motion is detected in some of them. We estimate that the closest binary systems in this sample have periods between 5 and 20 years, and thus dynamical masses will be derived in the near future. For the calculation of binary frequency we restrict ourselves to systems with distances less than 20 pc. After correction of the binaries bias, we find a ratio of visual binaries (at the HST limit of detection) of around 10%, and that ~15% of the 26 objects within 20 parsecs are binary systems with separations between 1 and 8 A.U. The observed frequency of ultra-cool binaries is similar than that of binaries with G-type primaries in the separation range from 2.1 A.U. to 140 A.U. There is also a clear deficit of ultra-cool binaries with separations greater than 15 A.U., and a possible tendency for the binaries to have mass ratios near unity. Most systems have indeed visual and near-infrared brightness ratios between 1 and 0.3. We discuss our results in the framework of current scenarios for the formation and evolution of free-floating brown dwarfs.Comment: 67 pages, 14 figures, Accepted for publication in AJ, September 2003. First submission to AJ: august 2002, 5 submission

Temperature constraints on the coldest brown dwarf known WISE 0855-0714

Context. Nearby isolated planetary mass objects are beginning to be discovered, but their individual properties are poorly constrained because their low surface temperatures and strong molecular self-absorption make them extremely faint. Aims. We aimed to detect the near infrared emission of the coldest brown dwarf (BD) found so far, WISE0855$-$0714, located $\sim$2.2 pc away, and to improve its temperature estimate (T$_{\rm eff}$= 225-260 K) from a comparison with state-of-the-art models of BD atmospheres. Methods. We observed the field containing WISE0855-0714 with HAWK-I at the VLT in the $Y$ band. For BDs with T$_{\rm eff}<$500\,K theoretical models predict strong signal (or rather less molecular absorption) in this band. Results. WISE0855-0714 was not detected in our Y-band images, thus placing an upper limit on its brightness to Y>24.4 mag at 3-$\sigma$ level, leading to Y-[4.5]>10.5. Combining this limit with previous detections and upper limits at other wavelengths, WISE0855$-$0714 is confirmed as the reddest BD detected, further supporting its status as the coldest known brown dwarf. We applied spectral energy distribution fitting with collections of models from two independent groups for extremely cool BD atmospheres leading to an effective temperature of T$_{\rm eff}<$250\,K,.Comment: 4 pages, 4 figures. A&A letter Accepte

The Substellar Mass Function in sigma Orionis

We combine results from imaging searches for substellar objects in the sigma Orionis cluster and follow-up photometric and spectroscopic observations to derive a census of the brown dwarf population in a region of 847 arcmin^2. We identify 64 very low-mass cluster member candidates in this region. We have available three color (IZJ) photometry for all of them, spectra for 9 objects, and K photometry for 27% of our sample. These data provide a well defined sequence in the I vs I-J, I-K color magnitude diagrams, and indicate that the cluster is affected by little reddening despite its young age (~5 Myr). Using state-of-the-art evolutionary models, we derive a mass function from the low-mass stars (0.2 Msol) across the complete brown dwarf domain (0.075 Msol to 0.013 Msol), and into the realm of free-floating planetary-mass objects (<0.013 Msol). We find that the mass spectrum (dN/dm ~ m^{-alpha}) increases toward lower masses with an exponent alpha = 0.8+/-0.4. Our results suggest that planetary-mass isolated objects could be as common as brown dwarfs; both kinds of objects together would be as numerous as stars in the cluster. If the distribution of stellar and substellar masses in sigma Orionis is representative of the Galactic disk, older and much lower luminosity free-floating planetary-mass objects with masses down to about 0.005 Msol should be abundant in the solar vicinity, with a density similar to M-type stars.Comment: Accepted for publication in ApJ. 19 pages, 3 figures include

Infrared Colors at the Stellar/Substellar Boundary

We present new infrared photometry for 61 halo and disk stars around the stellar/substellar boundary. These data are combined with available optical photometry and astrometric data to produce color--color and absolute magnitude--color diagrams. The disk and halo sequences are compared to the predictions of the latest model atmospheres and structural models. We find good agreement between observation and theory except for known problems in the V and H passbands probably due to incomplete molecular data for TiO, metal hydrides and H$_2$O. The metal--poor M subdwarfs are well matched by the models as oxide opacity sources are less important in this case. The known extreme M subdwarfs have metallicities about one--hundredth solar, and the coolest subdwarfs have T$_{eff}\sim 3000$ K with masses $\sim$0.09M/M$_{\odot}$. The grainless models are not able to reproduce the flux distributions of disk objects with T$_{eff} <$ 2500 K, however a preliminary version of the NextGen--Dusty models which includes homogeneous formation and extinction by dust grains {\it is} able to match the colors of these very cool objects. The least luminous objects in this sample are GD165B, three DENIS objects --- DBD0205, DBD1058 and DBD1228 --- and Kelu-1. These have T$_{eff}\sim$ 2000 K and are at or below the stellar limit with masses $\leq$0.075M/M$_{\odot}$. Photometry alone cannot constrain these parameters further as the age is unknown, but published lithium detections for two of these objects (Kelu-1 and DBD1228) imply that they are young (aged about 1 Gyr) and substellar (mass $\leq$0.06M/M$_{\odot}$).Comment: ApJ, in press. 18 pages. Also available at ftp://ftp.jach.hawaii.edu/pub/ukirt/skl/dM_preprint