Little owl (Athene noctua): nuclear and mitochondrial DNA analysis reveals divergence of soutwestern and central european subspecies

We utilized mitochondrial and nuclear DNA to study the phylogeography of little owls (Athene noctua) across Europe. Data from 326 individuals distributed among 22 sites were used to assess geographical distribution of the European subspecies, i.e., the widely accepted vidalii, noctua and indigena, and the questioned sarda-species. On the basis of the sequences of two mtDNA markers (CRI D-Loop 494 bp and COI 679 bp), we found a major difference between a first haplogroup, distributed in the North-West (from Iberia to Denmark and Czech Republic) and three other clades, distributed in the South-Eastern part of the study area. In particular we found a second group distributed in the Balkans and a third located in Sardinia, while in Italy there was a mixture between a fourth haplogroup of South-Italy and both the North-West and the Balkan clades. All individuals sampled in Sardinian differed clearly from other haplogroups. The combined analyses of the 13 nuclear microsatellite loci corroborated these results. Both mitochondrial and nuclear markers showed that interaction across contact zones generate a mixture of genotypes not only in Italy, but also in Central Europe. The apparent position of the contact zones differs substantially from the subspecies distribution reported in literature. DNA data are consistent with the phylogeographic hypothesis that this species survived the Pleistocene glaciations in three major refugia located in Iberia, South Italy, and the Balkans. Expansion patterns indicate that little owls from the Iberian refugium were the predominant source of postglacial colonization of northern Europe, while expansion out of South-Italy and Balkans had more limited effects

Combination of Measurements of Inclusive Deep Inelastic e±pe^{\pm}p Scattering Cross Sections and QCD Analysis of HERA Data

A combination is presented of all inclusive deep inelastic cross sections previously published by the H1 and ZEUS collaborations at HERA for neutral and charged current $e^{\pm}p$ scattering for zero beam polarisation. The data were taken at proton beam energies of 920, 820, 575 and 460 GeV and an electron beam energy of 27.5 GeV. The data correspond to an integrated luminosity of about 1 fb$^{-1}$ and span six orders of magnitude in negative four-momentum-transfer squared, $Q^2$, and Bjorken $x$. The correlations of the systematic uncertainties were evaluated and taken into account for the combination. The combined cross sections were input to QCD analyses at leading order, next-to-leading order and at next-to-next-to-leading order, providing a new set of parton distribution functions, called HERAPDF2.0. In addition to the experimental uncertainties, model and parameterisation uncertainties were assessed for these parton distribution functions. Variants of HERAPDF2.0 with an alternative gluon parameterisation, HERAPDF2.0AG, and using fixed-flavour-number schemes, HERAPDF2.0FF, are presented. The analysis was extended by including HERA data on charm and jet production, resulting in the variant HERAPDF2.0Jets. The inclusion of jet-production cross sections made a simultaneous determination of these parton distributions and the strong coupling constant possible, resulting in $\alpha_s(M_Z)=0.1183 \pm 0.0009 {\rm(exp)} \pm 0.0005{\rm (model/parameterisation)} \pm 0.0012{\rm (hadronisation)} ^{+0.0037}_{-0.0030}{\rm (scale)}$. An extraction of $xF_3^{\gamma Z}$ and results on electroweak unification and scaling violations are also presented.A combination is presented of all inclusive deep inelastic cross sections previously published by the H1 and ZEUS collaborations at HERA for neutral and charged current $e^{\pm}p$ scattering for zero beam polarisation. The data were taken at proton beam energies of 920, 820, 575 and 460 GeV and an electron beam energy of 27.5 GeV. The data correspond to an integrated luminosity of about 1 fb$^{-1}$ and span six orders of magnitude in negative four-momentum-transfer squared, $Q^2$, and Bjorken $x$. The correlations of the systematic uncertainties were evaluated and taken into account for the combination. The combined cross sections were input to QCD analyses at leading order, next-to-leading order and at next-to-next-to-leading order, providing a new set of parton distribution functions, called HERAPDF2.0. In addition to the experimental uncertainties, model and parameterisation uncertainties were assessed for these parton distribution functions. Variants of HERAPDF2.0 with an alternative gluon parameterisation, HERAPDF2.0AG, and using fixed-flavour-number schemes, HERAPDF2.0FF, are presented. The analysis was extended by including HERA data on charm and jet production, resulting in the variant HERAPDF2.0Jets. The inclusion of jet-production cross sections made a simultaneous determination of these parton distributions and the strong coupling constant possible, resulting in $\alpha_s(M_Z)=0.1183 \pm 0.0009 {\rm(exp)} \pm 0.0005{\rm (model/parameterisation)} \pm 0.0012{\rm (hadronisation)} ^{+0.0037}_{-0.0030}{\rm (scale)}$. An extraction of $xF_3^{\gamma Z}$ and results on electroweak unification and scaling violations are also presented.A combination is presented of all inclusive deep inelastic cross sections previously published by the H1 and ZEUS collaborations at HERA for neutral and charged current $e^{\pm }p$ scattering for zero beam polarisation. The data were taken at proton beam energies of 920, 820, 575 and 460 GeV and an electron beam energy of 27.5 GeV. The data correspond to an integrated luminosity of about 1 fb$^{-1}$ and span six orders of magnitude in negative four-momentum-transfer squared, $Q^2$ , and Bjorken x. The correlations of the systematic uncertainties were evaluated and taken into account for the combination. The combined cross sections were input to QCD analyses at leading order, next-to-leading order and at next-to-next-to-leading order, providing a new set of parton distribution functions, called HERAPDF2.0. In addition to the experimental uncertainties, model and parameterisation uncertainties were assessed for these parton distribution functions. Variants of HERAPDF2.0 with an alternative gluon parameterisation, HERAPDF2.0AG, and using fixed-flavour-number schemes, HERAPDF2.0FF, are presented. The analysis was extended by including HERA data on charm and jet production, resulting in the variant HERAPDF2.0Jets. The inclusion of jet-production cross sections made a simultaneous determination of these parton distributions and the strong coupling constant possible, resulting in $\alpha _s(M_Z^2)=0.1183 \pm 0.0009 \mathrm{(exp)} \pm 0.0005\mathrm{(model/parameterisation)} \pm 0.0012\mathrm{(hadronisation)} ^{+0.0037-0.0030}\mathrm{(scale)}$ . An extraction of $xF_3^{\gamma Z}$ and results on electroweak unification and scaling violations are also presented