Acknowledgments The authors wish to thank all landowners for permission to sample to populations; J. Squirrell, J. Droop, C. Horsfall, S. Aeschlimann, E. Crothall for fieldwork assistance; J. Squirrell, A. Forrest and M. Ruhsam for laboratory assistance; M. Ruhsam, A. Jump and A. Taylor for comments on an earlier version of this manuscript. RJC was supported by a Natural Environment Research Council studentship, a Scottish Natural Heritage Grant and an EU ATANS Grant (Fp6 506004) for travel and fieldwork expenses at the Abisko Scientific Research Station. The Royal Botanic Garden Edinburgh is supported by the Scottish Government’s Rural and Environment Science and Analytical Services Division.Peer reviewe
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