New nomenclatural changes are presented for selected North American mordellid beetles (Coleoptera: Mordellidae). The following five genera are newly recorded fi.·om the United States: Mordellaria Ermisch 1950, Falsomordellistena Ermisch 1941, Mordellina Schilsky 1908, Mordellochroa Emery 1876, and

Pseudotolida Ermisch 1950. The following species are transferred from Tomoxia and represent new combinations: Mordellaria borealis (LeConte 1862), Mordellaria fascifera (LeConte 1878), Mordellaria latipalpis (Ray 1946), Mordellaria serval (Say 1835), and Mordellaria undulata (Melsheimer 1845). The following species are transferred fi.·om Mordellistena and represent new combinations: Falsomordellistena hebraica (LeConte 1862) and Falsomordellistena pubescens (Fabricius 1798); Mordellina blatchleyi (Liljeblad 1945), Mordellina pilosella (Ray 1947), Mordellina pustulata (Melsheimer 1845), Mordellina ustulata (LeConte 1862), and Mordellina wichhami (Liljeblad 1945); Mordellochroa scapularis (Say 1824); Pseudotolida

arida (LeConte 1862), Pseudotolida hnausi (Liljeblad 1945), and Pseudotolida lutea (Melsheimer 1845). Mordellina ustulata (LeConte 1862) represents a return to species rank after being treated as a subspecies of Mordellistena andreae LeConte 1862. There are a total of 17 genera of Mordellidae in America north of Mexico

Jackman, John A.

Lu, Wenhua

English

New nomenclatural changes are presented for selected North American mordellid beetles (Coleoptera: Mordellidae). The following five genera are newly recorded from the United States: Mordellaria Ermisch 1950, Falsomordellistena E rmisch 1941, Mordellaria Schilsky 1908, Mordellochroa Emery 1876, and Pseudotolida Ermisch 1950. The following species are transferred from Tornoxia and represent new combinations: Mordellaria borealis (LeConte 1862), Mordellaria fascifera (LeConte 1878), Mordellaria latipalpis (Ray 1946), Mordellaria serval  (Say 1835), and Mordellaria undulata (Melsheimer 1845). The following species are transferred from Mordellistena and represent new combinations: Falsomordellistena hebraica (LeConte 1862) and Falsomordellistena pubescens (Fabricius 1798); Mordellina blatchleyi (Liljeblad 1945), Mordellina pilosella (Ray 1947), Mordellina pustulata (Melsheimer 1845), Mordellina ustulata (LeConte 1862), and Mordellina wickhami  (Liljeblad 1945); Mordellochroa scapularis (Say 1824); Pseudotolida arida  (LeConte 1862), Pseudotolida knausi (Liljeblad 1945), and Pseudotolida lutea (Melsheimer 1845). Mordellina  ustulata (LeConte 1862) represents a return to species rank after being treated as a subspecies of Mordellistena andreae LeConte 1862. There are a total of 17 genera of Mordellidae in America north of Mexico

DigitalCommons@University of Nebraska

University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Insecta Mundi Center for Systematic Entomology, Gainesville, Florida March 2001 Nomenclatural changes for selected Mordellidae (Coleoptera) in North America John A. Jackman Texas A&M University, College Station, TX Wenhua Lu Jamestown, RI Follow this and additional works at: https://digitalcommons.unl.edu/insectamundi  Part of the Entomology Commons Jackman, John A. and Lu, Wenhua, "Nomenclatural changes for selected Mordellidae (Coleoptera) in North America" (2001). Insecta Mundi. 179. https://digitalcommons.unl.edu/insectamundi/179 This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. INSECTA MUNDI, Vol. 15, No. 1, March, 2001 31 Nomenclatural changes for selected Mordellidae (Coleoptera) in North America John A. Jackman Department of Entomology Texas A&M University College Station, Texas, 77843-2475, USA and Wenhua Lu The Conservation Agency 6 Swinburne Street Jamestown, RI 02835, USA A b s t r a c t .  New nomenclatural changes are presented for selected North American mordellid beetles (Coleoptera: Mordellidae). T h e  following five genera are newly recorded from the  United States: Moi-dellaria Ermisch 1950, Falsornordellistei~a Ermisch 1941, Mordelliiza Schilsky 1908, Mordellochroa Emery 1876, and Pseudotolida Ermisch 1950. T h e  following species are transferred from Tornoxia and represent new combinations: Mordellaria borealis (LeConte 1862), Mordellaria fascifera (LeConte 1878), Mordellaria la,tipalpis (Ray 1946), Mordellaria serval (Say  1835), and Mordellaria z~izdulata (Melsheimer 1845). T h e  following species are transferred from Mordellisteiza and represent new combinations: Falsornordellisteiza hebraica (LeConte 1862) and Falsoinordellisteiza pubesceizs (Fabricius 1798); Mordelliira blatchleyi (Liljeblad 1945), Mordelliira pilosella (Ray 1947), Mordellirza pustulata (Melsheimer 1845), Mordelliira ustulata (LeConte 1862), and Mordelliira wiclzharni (Liljeblad 1945); Mordellochroa scapularis (Say  1824); Psez~dotolida arida (LeConte 1862), Pseudotolida lzrrausi (Liljeblad 1945), and Pseudotolida lutea (Melsheimer 1845). Mordelliira ustzslata (LeConte 1862) represents a return t o  species rank after being treated as a subspecies o f  Mordellisteira aizdreae LeConte 1862. There are a total o f  17 genera o f  Mordellidae i n  America north o f  Mexico. Introduction The latest revision of the  Mordellidae i n  North America was published by Liljeblad (1945). The latest catalog of North American mordellids was completed by Bright (1986) and  was subsequently updated by Jackman (1991) resulting in  12 genera recognized i n  North America north of Mexico. Jack- m a n  (1991) suggested tha t  some of the  North Amer- ican species have not been correctly assigned to genera a s  European and Oriental authorities un- derstand them. Consequently, we continue to in- vestigate the generic placement of North American mordellids. Although various authors have added to the  work, Franciscolo (1957) prepared the most com- prehensive study for the world genera. I n  this paper we propose nomenclatural changes tha t  a re  justified based on our current  understanding of the genera. The characters used i n  the key by Fran- ciscolo (1957, 1965: 1967) provide the  basis for the nomenclatural changes. We transfer five species from Tolnoxia to Mordellaria. From Mordellistel~a, we transfer two species to Falsomordelliste7~a, five species to Mordelli7~a, one species to Mordello- chroa, and three to Pseudotolida. Our work in- creases the number of genera i n  America north of Mexico from 12 to 17 total. Further  study will likely justify additional nomenclatural reassignments. Mordellaria Errnisch 1950 Mordellaria contains about 14 species i n  Afri- ca, Japan,  Chile, Taiwan, and  now North America. No worldwide key to species is available. Takakuwa (1985) provided a key to species i n  J apan  and Taiwan. The keys to species by Liljeblad (1945) and Ray (1946) will separate species in  North America even though these spocies a re  treated in  Tomoxia. Mordellaria has  the scutellum triangular or more or less rounded a t  the apex. Male genitalia of the genus (e.g., Lu e t  al. 1997) are very different from those of Tolnoxia redefined by Franciscolo (1982). These characters separate it from Tomoxia, which has  the scutellum quadrilateral or trapezoi- dal. Based on these characters, five North Ameri- 3 2 Volume 15, No. 1, March, 2001, INSECTA MUNDI can species are here transferred from Torlzoxia: Mordellaria borealis (LeConte 1862); Mordellaria fascifera (LeConte 1878); Mordellaria latipalpis (Ray 1946); Mordellaria serual (Say 1835); and Mordellaria urtdulata (Melsheimer 1845), [NEW COMBINATIONS]. Falsomordellistena Ermisch 1941 Fa.lson~ol-dellistelta contains about 25 species and is found primarily in Japan,  Taiwan, China, and now North America. Hatayama (1985) provid- ed a key to species i n  Japan and Taiwan. Falsontordellisterta has the penultimate tar- someres of the fore and middle legs emarginate, more or less bilobed, and usually dilated; the apical palpomere of the maxilla is securiform or scalene in  both sexes; the eyes are small, oval shaped, and sometimes slightly emarginate behind the anten- na; eye facets are fine. Based on these characters, two North American species are here transferred from Mordelliste~ta: Falso~~zordell iste~~a hebraica (LeConte 1862); and Falso~~zordel l i s te~~apubesce~~s  (Fabricius 1798), [NEW COMBINATIONS]. Both of the North American species have the elytra patterned. Mordellina Schilsky 1908 This genus contains about 45 species found in  Europe, Africa, China, Japan, Taiwan, and now North America. Ermisch (1968) and Francisco10 (1967) provided keys to African species. Hatayama (1985) provided a key to species in  Japan and Taiwan. Batten (1976) provided a key to the species in  the Netherlands. Mordellirta has the penultimate tarsomeres of the fore and middle legs transversely truncate a t  the apex, or very moderately emarginate, neither dilated nor bilobed; the apical palpomere of the maxilla is securiform, scalene, or cultriform, with sharp or sometimes rounded angles in both sexes; the eyes are large, shape unusually broad, emar- ginate behind the antenna; eye facets are coarse, and never smaller than 0.022 mm in diameter. Based on these characters, five North American species are here transferred from Mordellisterta: Mordellirza blatchleyi (Liljeblad 1945); Mordellirta pilosella (Ray 1947); Mordelli~zapustulata (Melshe- imer 1845); Mordelli7~a ustulata (LeConte 1862); and Mordellina wickharni (Liljeblad 1945), [NEW COMBINATIONS]. This restores species rank to M. ustulata, which has been treated as a subspecies of Mordelliste7ta andreae LeConte 1862 (Liljeblad 1945). Mordellochroa Emery 1876 Mordellochroa has a t  least six species world- wide and is found in  Japan,  Mongolia, Poland, Hungary, and probably Russia. Batten (1977) pro- vided a key to the Palearctic species, Kaszab (1979) and Horhk (1993) provided keys for the species in Hungary, and Borowiec (1996) provided a key to species in Poland. Mordellochroa has  the penultimate tarsomeres of the fore and middle legs transversely truncate a t  the apex, or very moderately emarginate, neither dilated nor bilobed; the apical palpomere of the maxilla is malleiform i n  males and securiform or scalene in  the females. Based on these characters, one species is here transferred from Mordellisterta: Mordellochroa scapz~laris (Say 1824), [NEW COM- BINATION]. This species has blacli elytra with humeral spots reddish to yellow and the lateral ridges on the hind tibia quite short. Pseudotolida Ermisch 1950 Pseudotolida has fewer than 10 species in  Ja-  pan, China, Taiwan, Guatemala, and Brazil. Fran- ciscolo (1982) provided a key to some species. Shiy- ake (1995, 1997) compared the genus to other related genera. Pseudotolida has the penultimate tarsomeres of the fore and middle legs deeply emarginate, more or less bilobed and usually strongly dilated; the apical palpomere of the maxilla is malleiform in males and securiform in  females. Based on these characters, three North American species are here transferred from Mordellisterta: Pseudotolida ari- d a  (LeConte 1862); Pseudotolida Kl~ausi (Liljeblad 1945); and Pseudotolida lutea (Melsheimer 1845), [NEW COMBINATIONS]. Acknowledgments We thank J a n  Horhk for pointing out that  Mordelis- tens scap~~lal- is  (Say) belongs in  Mordellochroa. Bibliography Batten, R. 1976. De Nederlandse soorten van de keverfamilie Mordell idae.  Zoologische Bijdragen, 19: 3-37. INSECTA MUNDI, Vol. 15, No. 1, March, 2001 Batten, R. 1977. lMbrdellochroa species of the Western Palaearctic region (Coleoptera, Mordel- lidae). Entomologische Berichten, 37: 21-26. Borowiec, L. 1996. Mordellidae. Miastkowate (In- sects: Coleoptera). Fauna Polski, Tom 18, 189 pp. Polska Akademia Nauk. Muzeum I Instytut Zoologii, Warszawa. Bright, D. E. 1986. A catalog of the Coleoptera of America North of Mexico, Family: Mordellidae. USDA, ARS, Agriculture Handbook 529-125, 22 pp. Emery, M. C. 1876. Essai m ~ n o g ~ a p h i q u e  sur les Mordellides de L'Europe e t  des Contr6es Lim- itrophes. L'Abeille, 14: 1-128. Ermisch, K. 1941. Tribus Mordellistenini (Col. Mordell.) (8. Beitrag zur Kenntnis der Mordel- liden). Mitteilungen der Miinchener Entomol- ogischen Gesellschaft, 31: 710-726. Errnisch, K. 1950. Die Gattungen der Mordelliden der Welt (16. Beitrag zur Kenntnis der Mordel- liden). Entomologische Blatter, 46: 34-92. Ermisch, K. 1968. Contributions A la connaissance de la faune entomologique de la CBte-D'Ivoire (J. Decelle, 1961-1964). 19. Coleoptera Mordel- lidae. (55. Beitrag zur Kenntnis der Mordel- liden). Musee Royal de L-Afrique Centrale, Tervuren, Belgique Annales, Serie in-8 , Sci- ences Zoologiques, 165: 257-287. Fabricius, J. C. 1798. Supplementum entomologi- ae systematicae, Hafniae, 572 pp. Franciscolo, M. 1957. A monograph of the South African genera and species of Mordellidae. Part  1. Morphology, subfamily Ctenidiinae and tribe Stenaliini. I~L South African Animal Life, Re- sults of the Lunds University Expedition in 1950-1951, 4: 207-291. Franciscolo, M. 1965. A monograph of the South African genera and species of Mordellidae. Part 2. Tribe Mordellini. I~L South African Animal Life, Results of the Lunds University Expedi- tion in 1950-1951, 11: 244-458. Franciscolo, M. 1967. A monograph of the South African genera and species of Mordellidae. Part  3. Tribe Mordellistenini. Iit South African An- imal Life, Results of the Lunds University Ex- pedition in 1950-1951, 13: 67-203. Franeiscolo, M. 1982. Mordellidae (C~leop te~a :  Heteromera) from Rennell and Bellona Islands (Solomons). I~L T. Wolff (ed.). The Natural His- tory of Rennell Island, British Solomon Islands, University of Copenhagen, 8: 49-72. Hatay ama, T. 1985. Mordellidae: Tribe Mordellis- tenini. ITL Y. I<ui*osawa et al. (eds.), The Co- leoptera of Japan in Color. Hoikusha, Osaka, 387-397. Horak, J. 1993. Mordellidae, Scraptiidae. I r z  J .  Jel nek (ed.), Check-list of Czechoslovak Insects IV (Coleoptera). Seznam eskoslovensk ch brouk . Folia Heyrovskyana, Suppl. I, 111-112, 115. Jackman, J. A. 1991. Notes on the nomenclature of Mordellidae of America north of Mexico. Coleopterists Bulletin, 45(4): 323-330. Kaszab, Z. 1979. Felemhs lhbfejizes bogarak 11. Heteromera 11. Mordellidae. Magyarorszag Al- latvilaga, Fauna Hungaria, Akademiai Kiado, Budapest, 9(2): 1-100. Leconte, J. L. 1862. Synopsis of the Mordellidae of the United States. Proceedings of the Natural Sciences of Philadelphia, 14: 43-51. Leconte, J. L. 1878. Additional descriptions of new species, pp. 373-434. I~L E. A. Schwarz (ed.), The Coleoptera of Florida. Proceedings of the Amer- ican Philosophical Society, 17: 373-472. Liljeblad, E. 1945. Monograph of the family Mordel- lidae (Coleoptera) of North America, north of Mexico. Miscellaneous Publications, Museum of Zoology, University of Michigan, No. 62, 229 PP. Lu, W., J. A. Jackman and P. W. Johnson. 1997. Male genitalia and phylogenetic relationships in North American Moi-dellidae (C~leopte~a) .  Annals of the Entoinological Society of Ameri- ca, 90(6): 742-767. Melsheimer, F. E. 1845. Descriptions of new spe- cies of Coleoptera of the United States. Pro- ceedings of the Academy of Natural Sciences of Philadelphia, 2: 213-223, 302-318. Ray, E. 1946. Studies on North American Mordel- lidae, I11 (Coleoptera). Pan-Pacific Entomolo- gist. 22(3): 90-99. Ray, E. 1947. Studies on North American Mordel- lidae, V (Coleoptera). Pan-Pacific Entomolo- gist, 23(3): 121-131. Say, T. 1824. Descriptions of coleopterous insects collected in the late expedition to the Rocky Mountains, performed by order of Mr. Cal- houn, Secretai-y of War. under cominand of Major Long. Journal of the Academy of Natural Sciences of Philadelphia, 3(2): 238-282, 298- 331, 403-464; 4(1): 83-99. Say, T. 1835. Descriptions of North American co- leoptero-cts insects and observations on some already described. Boston Journal of Natural History, l(2): 151-203. 3 4 Volume 15, No. 1, March, 2001, INSECTA MUNDI Schilsky, J. 1908. XII. Beitrag zur deutschen Shiyake, S. 1997. Two new species of the genus Kaferfauna. Deutsche Entomologische Zeitschr- Tolidopalpz~s from the Philippines (Coleoptera, schrift, 604-608. Mordellidae). Entomological Review of Japan, Shiyake, S. 1995. A taxonomic study on the genus 52(1): 51-54. Tolidopalpus, with description of a new species Takakuwa, M. 1985. Mordellidae: Tribe Mordelli- (Coleoptera: Mordellidae). Bulletin of the Osa- ni. In Y. Kurosawa e t  al. (eds.), The Coleoptera lia Museum of Natural History, 49: 11-18. of J apan  in  Color. Hoikusha, Osaka: 376-387. 

Nomenclatural changes for selected Mordellidae (Coleoptera) in North America

Hochschulschriftenserver - Universität Frankfurt am Main

INSECTA MUNDI, Vol. 15, No.1, March, 2001 Nomenclatural changes for selected Mordellidae (Coleoptera) in North America John A. Jackman Department of Entomology Texas A&M University College Station, Texas, 77843-2475, USA and Wenhua Lu The Conservation Agency 6 Swinburne Street Jamestown, RI 02835, USA Abstract.  New  nomenclatural changes  are  presented  for  selected  North American  mordellid  beetles (Coleoptera: Mordellidae). The following five genera are newly recorded fi.·om the United States: Mordellaria Ermisch 1950, Falsomordellistena Ermisch 1941, Mordellina Schilsky 1908, Mordellochroa Emery 1876, and Pseudotolida  Ermisch  1950.  The  following  species  are  transferred  from  Tomoxia  and represent  new combinations:  Mordellaria  borealis  (LeConte  1862),  Mordellaria  fascifera  (LeConte  1878),  Mordellaria latipalpis  (Ray  1946),  Mordellaria serval  (Say 1835),  and Mordellaria undulata (Melsheimer 1845).  The following species are transferred fi.·om  Mordellistena and represent new combinations: Falsomordellistena hebraica (LeConte 1862) and Falsomordellistena pubescens (Fabricius 1798); Mordellina blatchleyi (Liljeblad 1945),  Mordellina  pilosella  (Ray  1947),  Mordellina  pustulata  (Melsheimer  1845),  Mordellina  ustulata (LeConte 1862), and  Mordellina wichhami (Liljeblad 1945); Mordellochroa scapularis (Say 1824); Pseudotolida arida (LeConte  1862),  Pseudotolida hnausi (Liljeblad  1945),  and Pseudotolida  lutea  (Melsheimer 1845). Mordellina ustulata (LeConte 1862) represents a return to species rank after being treated as a subspecies of Mordellistena andreae LeConte 1862. There are a total of 17 genera of Mordellidae in America north of Mexico. 31 Introduction The latest revision of the Mordellidae in North America was published by Liljeblad  (1945).  The latest catalog of North American mordellids was completed by Bright (1986) and was subsequently updated by Jackman (1991) resulting in 12 genera recognized in North America north of  Mexico. J ack-man (1991) suggested that some ofthe North  Amer-ican species have not been correctly assigned to genera as European and Oriental authorities un-derstand them. Consequently, we continue to in-vestigate the generic placement of North American mordellids. we transfer two species to Falsomordellistena, five species  to  Mordellina,  one  species  to  Mordello-chroa,  and three  to  Pseudotolida.  Our work in-creases the number of genera in America north of Mexico from 12 to 17 total. Further study will likely justify additional nomenclatural reassignments. Although various authors have added to the work, Franciscolo (1957) prepared the most com-prehensive  study  for  the  world  genera.  In this paper we propose nomenclatural changes that are justified based on our current understanding ofthe genera. The characters used in the key by Fran-cis  colo (1957, 1965, 1967) provide the basis for the nomenclatural changes. We transfer five  species from Tomoxia to Mordellaria. From Mordellistena, Mordellaria Ermisch 1950 Mordellaria contains about 14 species in Afri-ca, Japan, Chile, Taiwan, and now North America. No worldwide key to species is available. Takakuwa (1985)  provided  a  key  to  species in Japan and Taiwan. The keys to species by Liljeblad (1945) and Ray (1946) will separate species in North America even though these sp9cies are treated in Tomoxia. Mordellaria  has the scutellum triangular or more or less rounded at the apex. Male genitalia of the genus (e.g.,  Lu et al.  1997) are very different from  those  of Tomoxia  redefined by  Franciscolo (1982). These characters separate it from Tomoxia, which has the scutellum quadrilateral or trapezoi-dal. Based on these characters, five North Ameri-32  Volume 15, No.1, March, 2001, INSECTA MUNDI can species are  here transferred from  TOlnoxia: Mordellaria borealis (LeConte 1862); Mordellaria fascifera  (LeConte  1878);  Mordellaria  latipalpis (Ray  1946);  Mordellaria  serual  (Say  1835);  and Mordellaria undulata (Melsheimer  1845),  [NEW COMBINATIONS]. Falsomordellistena Ermisch 1941 Falsomordellistena contains about 25  species and is found primarily in Japan, Taiwan, China, and now North America. Hatayama (1985) provid-ed a key to species in Japan and Taiwan. Falsomordellistena has the penultimate tar-someres of the fore  and middle legs emarginate, more or less bilobed, and usually dilated; the apical palpomere ofthe maxilla is securiform or scalene in both sexes; the eyes are small, oval shaped, and sometimes slightly emarginate behind the anten-na; eye facets are fine. Based on these characters, two North American species are here transferred from  Mordellistena:  Falsomordellistena  hebraica (LeConte 1862); and Falsonwrdellistena pubescens (Fabricius 1798),  [NEW COMBINATIONS]. Both of the  North American  species  have  the  elytra patterned. Mordellina Schilsky 1908 This genus contains about 45 species found in Europe, Africa,  China, Japan, Taiwan,  and now North America.  Ermisch (1968)  and Franciscolo (1967) provided keys to African species. Hatayama (1985)  provided  a  key  to  species  in Japan and Taiwan. Batten (1976) provided a key to the species in the Netherlands. Mordellina has the penultimate tarsomeres of the fore and middle legs transversely truncate at the apex, or very moderately emarginate, neither dilated nor bilobed;  the apical palpomere of the maxilla is securiform, scalene, or cultriform, with sharp or sometimes rounded angles in both sexes; the eyes are large, shape unusually broad, emar-ginate behind the antenna; eye facets are coarse, and never smaller than 0.022  mm in diameter. Based on these characters, five  North American species are here transferred from Mordellistena: Mordellina blatchleyi (Liljeblad 1945); Mordellina pilosella (Ray 1947); Mordellina  pustulata (Melshe-imer 1845); Mordellina ustulata (LeConte 1862); and Mordellina wichhami (Liljeblad 1945), [NEW COMBINATIONS]. This restores species rank to M. ustulata, which has been treated as a subspecies of Mordellistena andreae LeConte 1862 (Liljeblad 1945). Mordellochroa Emery 1876 Mordellochroa has at least six species world-wide  and is  found  in Japan,  Mongolia,  Poland, Hungary, and probably Russia. Batten (1977) pro-vided a key to the Pale  arctic species, Kaszab (1979) and Horak (1993) provided keys for the species in Hungary, and Borowiec (1996) provided a  key to species in Poland. Mordellochroa has the penultimate tarsomeres of the fore and middle legs transversely truncate at the apex, or very moderately emarginate, neither dilated nor bilobed; the apical palpomere  of the maxilla is malleiform in males and securiform or scalene in the females. Based on these characters, one species is here transferred from Mordellistena: Mordellochroa scapularis (Say 1824), [NEW COM-BINATION].  This species has black elytra with humeral spots reddish to yellow  and the lateral ridges on the hind tibia quite short. Pseudotolida Ermisch 1950 Pseudotolida has fewer than 10 species in Ja-pan, China, Taiwan, Guatemala, and Brazil. Fran-cis  colo (1982) provided a key to some species. Shiy-ake  (1995,  1997)  compared  the  genus  to  other related genera. Pseudotolida has the penultimate tarsomeres of  the fore and middle legs deeply emarginate, more or less bilobed and usually strongly dilated; the apical palpomere of the maxilla is malleiform in males and securiform in females. Based on these characters, three North American species are here transferred from Mordellistena: Pseudotolida ari-da (LeConte 1862); Pseudotolida hnausi (Liljeblad 1945); and Pseudotolida lutea (Melsheimer 1845), [NEW COMBINATIONS]. Acknow  ledgments We thank Jan  Horak  for pointing out  thatMordelis-tena scapularis (Say) belongs in Mordellochroa. Bibliography Batten, R. 1976. De Nederlandse soorten van de keverfamilie  Mordellidae.  Zoologische Bijdragen, 19: 3-37. INSECTA MUNDI, Vol. 15, No.1, March, 2001 Batten,  R.  1977.  Mordellochroa  species  of the Western Palaearcticregion (Coleoptera, Mordel-lidae). Entomologische Berichten, 37: 21-26. Borowiec, L. 1996. Mordellidae. Miastkowate (In-secta: Coleoptera). Fauna Polski, Tom 18, 189 pp. PolskaAkademia Nauk, Muzeum I Instytut Zoologii, Warszawa. Bright, D. E. 1986. A catalog of the Coleoptera of America North of  Mexico, Family: Mordellidae. USDA, ARS,  Agriculture Handbook 529-125, 22 pp. ElIlery, M. C.  1876. Essai monographique sur les Mordellides de L'Europe et des Contrees Lim-itrophes. L'Abeille, 14: 1-128. ErlIlisch, K.  1941.  Tribus  Mordellistenini  (Col. Mordell.) (8. Beitrag zur Kenntnis del' Mordel-liden). Mitteilungen del' Munchener Entomol-ogischen Gesellschaft, 31: 710-726. ErlIlisch, K. 1950. Die Gattungen del' Mordelliden del' Welt (16. Beitrag zur Kenntnis del' Mordel-liden). Entomologische Blatter, 46: 34-92. ErlIlisch, K. 1968. Contributions a  la connaissance de la faune entomologique de la C6te-D'Ivoire (J. Decelle, 1961-1964). 19. Coleoptera Mordel-lidae.  (55.  Beitrag zur Kenntnis del' Mordel-liden).  Musee  Royal  de  L-Afrique  Centrale, Tervuren, Belgique Annales, Serie in-8 , Sci-ences Zoologiques, 165: 257-287. Fabricius, J. C. 1798. Supplementum entomologi-ae systematicae, Hafniae, 572 pp. Franciscolo, M. 1957. A monograph of the South African genera and species ofMordellidae. Part 1. Morphology, subfamily Ctenidiinae and tribe Stenaliini. In South African Animal Life, Re-sults of the Lunds  University Expedition in 1950-1951,4: 207-291. Franciscolo, M.  1965. A monograph of the South African genera and species of  Mordellidae. Part 2.  Tribe Mordellini. In South African Animal Life, Results of the Lunds University Expedi-tion in 1950-1951, 11: 244-458. Franciscolo, M. 1967. A monograph of the South African genera and species ofMordellidae. Part 3. Tribe Mordellistenini. In South African An-imal Life, Results of the Lunds University Ex-pedition in 1950-1951, 13: 67-203. Franciscolo, M.  1982.  Mordellidae  (Coleoptera: Heteromera) from Rennell and Bellona Islands (Solomons). In T. Wolff (ed.), The Natural His-tory ofRennell Island, British Solomon Islands, University of Copenhagen, 8:  49-72. HatayalIla, T. 1985. Mordellidae: Tribe Mordellis-tenini. In Y.  Kurosawa et al.  (eds.),  The  Co-33 leoptera of Japan in Color. Hoikusha, Osaka, 387-397. Horak, J. 1993. Mordellidae, Scraptiidae. In J. Jel nek (ed.), Check-list of Czechoslovak Insects IV (Coleoptera). Seznam  eskoslovensk ch brouk . Folia Heyrovskyana, Suppl. I,  111-112, 115. JacklIlan, J. A. 1991. Notes on the nomenclature of Mordellidae  of America  north of Mexico. Coleopterists Bulletin, 45(4): 323-330. Kaszab, Z.  1979.  Felemas labfejizes bogarak II. Heteromera II. Mordellidae. Magyarorszag AI-latvilaga, Fauna Hungaria, Akademiai Kiado, Budapest, 9(2): 1-100. Leconte, J. L. 1862. Synopsis of the Mordellidae of the United States. Proceedings of the Natural Sciences of Philadelphia, 14: 43-51. Leconte, J. L. 1878. Additional descriptions of  new species, pp. 373-434. In E. A. Schwarz (ed.), The Coleoptera of  Florida. Proceedings of  the Amer-ican Philosophical Society, 17: 373-472. Liljeblad,  E. 1945. Monograph of  the family Mordel-lidae (Coleoptera) of North America, north of Mexico.  Miscellaneous Publications, Museum of Zoology, University of Michigan, No. 62, 229 pp. Lu, W., J. A. JacklIlan and P. W. Johnson. 1997. Male genitalia and phylogenetic relationships in North American Mordellidae (Coleoptera). Annals of the Entomological Society of Ameri-ca, 90(6): 742-767. MelsheilIler, F. E. 1845. Descriptions of new spe-cies  of Coleoptera of the United States. Pro-ceedings of the Academy of Natural Sciences of Philadelphia, 2: 213-223, 302-318. Ray, E. 1946. Studies on North American Mordel-lidae, III (Coleoptera). Pan-Pacific Entomolo-gist, 22(3): 90-99. Ray, E. 1947. Studies on North American Mordel-lidae,  V  (Coleoptera).  Pan-Pacific  Entomolo-gist, 23(3): 121-131. Say, T. 1824. Descriptions of coleopterous insects collected in the late expedition to  the Rocky Mountains,  performed  by  order  of Mr.  Cal-houn,  Secretary  of War,  under command  of Major Long. Journal of  the Academy of Natural Sciences of Philadelphia,  3(2):  238-282,  298-331, 403-464; 4(1): 83-99. Say, T. 1835. Descriptions of North American co-leopterous insects and observations on some already described. Boston Journal of Natural History, 1(2): 151-203. 34  Volmue 15, No.1, March, 2001, INSECTA MUNDI Schilsky,  J.  1908.  XII.  Beitrag  zur  deutschen Kaferfauna. Deutsche Entomologische Zeitschr-schrift, 604-608. Shiyake, S. 1995. A taxonomic study on the genus Tolidopalpus, with description of a new species (Coleoptera: Mordellidae). Bulletin of the Osa-ka Museum of Natural History, 49:  11-18. Shiyake, S.  1997. Two new species of the genus Tolidopalpus from the Philippines (Coleoptera, Mordellidae). Entomological Review of Japan, 52(1); 51-54. Takakuwa, M. 1985. Mordellidae: Tribe Mordelli-ni. In Y. Kurosawa et al. (eds.), The Coleoptera of Japan in Color. Hoikusha, Osaka: 376-387. 

A catalog of the Coleoptera of

A monograph of the South African genera and species of Mordellidae. Part 2. Tribe Mordellini.

A monograph of the South African genera and species ofMordellidae. Part 1. Morphology, subfamily Ctenidiinae and tribe Stenaliini.

A monograph of the South African genera and species ofMordellidae. Part 3. Tribe Mordellistenini.

A taxonomic study on the genus Tolidopalpus, with description of a new species (Coleoptera: Mordellidae).

Additional descriptions of new species,

Contributions a la connaissance de la faune entomologique de la C6te-D'Ivoire

De Nederlandse soorten van de keverfamilie Mordellidae.

Descriptions of coleopterous insects collected in the late expedition to the Rocky Mountains, performed by order of Mr. Calhoun, Secretary of War, under command of Major Long.

Descriptions of new species of Coleoptera of the United States.

Descriptions of North American coleopterous insects and observations on some already described.

Die Gattungen del' Mordelliden del' Welt (16. Beitrag zur Kenntnis del'

Essai monographique sur les Mordellides de L'Europe et des Contrees Limitrophes.

Felemas labfejizes bogarak II. Heteromera II.

Male genitalia and phylogenetic relationships in

Monograph of the family Mordellidae (Coleoptera) of North America,

Mordellidae (Coleoptera: Heteromera) from Rennell and Bellona Islands

Mordellidae: Tribe Mordellini. In

Mordellidae: Tribe Mordellistenini. In

Mordellidae. Miastkowate (Insecta: Coleoptera). Fauna Polski,

Notes on the nomenclature of Mordellidae of America north of Mexico.

Supplementum entomologiae systematicae,

Synopsis of the Mordellidae of the United States.

Tribus Mordellistenini (Col. Mordell.) (8. Beitrag zur Kenntnis del' Mordelliden). Mitteilungen del'

Two new species of the genus Tolidopalpus from the Philippines (Coleoptera,

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March 2001 
Nomenclatural changes for selected Mordellidae (Coleoptera) in 
North America 
John A. Jackman 
Texas A&M University, College Station, TX 
Wenhua Lu 
Jamestown, RI 
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Jackman, John A. and Lu, Wenhua, "Nomenclatural changes for selected Mordellidae (Coleoptera) in 
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INSECTA MUNDI, Vol. 15, No. 1, March, 2001 31 
Nomenclatural changes for selected Mordellidae 
(Coleoptera) in North America 
John A. Jackman 
Department of Entomology 
Texas A&M University 
College Station, Texas, 77843-2475, USA 
and 
Wenhua Lu 
The Conservation Agency 
6 Swinburne Street 
Jamestown, RI 02835, USA 
A b s t r a c t .  New nomenclatural changes are presented for selected North American mordellid beetles 
(Coleoptera: Mordellidae). T h e  following five genera are newly recorded from the  United States: Moi-dellaria 
Ermisch 1950, Falsornordellistei~a Ermisch 1941, Mordelliiza Schilsky 1908, Mordellochroa Emery 1876, and 
Pseudotolida Ermisch 1950. T h e  following species are transferred from Tornoxia and represent new 
combinations: Mordellaria borealis (LeConte 1862), Mordellaria fascifera (LeConte 1878), Mordellaria 
la,tipalpis (Ray 1946), Mordellaria serval (Say  1835), and Mordellaria z~izdulata (Melsheimer 1845). T h e  
following species are transferred from Mordellisteiza and represent new combinations: Falsornordellisteiza 
hebraica (LeConte 1862) and Falsoinordellisteiza pubesceizs (Fabricius 1798); Mordelliira blatchleyi (Liljeblad 
1945), Mordelliira pilosella (Ray 1947), Mordellirza pustulata (Melsheimer 1845), Mordelliira ustulata 
(LeConte 1862), and Mordelliira wiclzharni (Liljeblad 1945); Mordellochroa scapularis (Say  1824); Psez~dotolida 
arida (LeConte 1862), Pseudotolida lzrrausi (Liljeblad 1945), and Pseudotolida lutea (Melsheimer 1845). 
Mordelliira ustzslata (LeConte 1862) represents a return t o  species rank after being treated as a subspecies 
o f  Mordellisteira aizdreae LeConte 1862. There are a total o f  17 genera o f  Mordellidae i n  America north o f  
Mexico. 
Introduction 
The latest revision of the  Mordellidae i n  North 
America was published by Liljeblad (1945). The 
latest catalog of North American mordellids was 
completed by Bright (1986) and  was subsequently 
updated by Jackman (1991) resulting in  12 genera 
recognized i n  North America north of Mexico. Jack- 
m a n  (1991) suggested tha t  some of the  North Amer- 
ican species have not been correctly assigned to 
genera a s  European and Oriental authorities un- 
derstand them. Consequently, we continue to in- 
vestigate the generic placement of North American 
mordellids. 
Although various authors have added to the  
work, Franciscolo (1957) prepared the most com- 
prehensive study for the world genera. I n  this 
paper we propose nomenclatural changes tha t  a re  
justified based on our current  understanding of the 
genera. The characters used i n  the key by Fran- 
ciscolo (1957, 1965: 1967) provide the  basis for the 
nomenclatural changes. We transfer five species 
from Tolnoxia to Mordellaria. From Mordellistel~a, 
we transfer two species to Falsomordelliste7~a, five 
species to Mordelli7~a, one species to Mordello- 
chroa, and three to Pseudotolida. Our work in- 
creases the number of genera i n  America north of 
Mexico from 12 to 17 total. Further  study will likely 
justify additional nomenclatural reassignments. 
Mordellaria Errnisch 1950 
Mordellaria contains about 14 species i n  Afri- 
ca, Japan,  Chile, Taiwan, and  now North America. 
No worldwide key to species is available. Takakuwa 
(1985) provided a key to species i n  J apan  and 
Taiwan. The keys to species by Liljeblad (1945) and 
Ray (1946) will separate species in  North America 
even though these spocies a re  treated in  Tomoxia. 
Mordellaria has  the scutellum triangular or 
more or less rounded a t  the apex. Male genitalia of 
the genus (e.g., Lu e t  al. 1997) are very different 
from those of Tolnoxia redefined by Franciscolo 
(1982). These characters separate it from Tomoxia, 
which has  the scutellum quadrilateral or trapezoi- 
dal. Based on these characters, five North Ameri- 
3 2 Volume 15, No. 1, March, 2001, INSECTA MUNDI 
can species are here transferred from Torlzoxia: 
Mordellaria borealis (LeConte 1862); Mordellaria 
fascifera (LeConte 1878); Mordellaria latipalpis 
(Ray 1946); Mordellaria serual (Say 1835); and 
Mordellaria urtdulata (Melsheimer 1845), [NEW 
COMBINATIONS]. 
Falsomordellistena Ermisch 1941 
Fa.lson~ol-dellistelta contains about 25 species 
and is found primarily in Japan,  Taiwan, China, 
and now North America. Hatayama (1985) provid- 
ed a key to species i n  Japan and Taiwan. 
Falsontordellisterta has the penultimate tar- 
someres of the fore and middle legs emarginate, 
more or less bilobed, and usually dilated; the apical 
palpomere of the maxilla is securiform or scalene in  
both sexes; the eyes are small, oval shaped, and 
sometimes slightly emarginate behind the anten- 
na; eye facets are fine. Based on these characters, 
two North American species are here transferred 
from Mordelliste~ta: Falso~~zordell iste~~a hebraica 
(LeConte 1862); and Falso~~zordel l i s te~~apubesce~~s  
(Fabricius 1798), [NEW COMBINATIONS]. Both 
of the North American species have the elytra 
patterned. 
Mordellina Schilsky 1908 
This genus contains about 45 species found in  
Europe, Africa, China, Japan, Taiwan, and now 
North America. Ermisch (1968) and Francisco10 
(1967) provided keys to African species. Hatayama 
(1985) provided a key to species in  Japan and 
Taiwan. Batten (1976) provided a key to the species 
in  the Netherlands. 
Mordellirta has the penultimate tarsomeres of 
the fore and middle legs transversely truncate a t  
the apex, or very moderately emarginate, neither 
dilated nor bilobed; the apical palpomere of the 
maxilla is securiform, scalene, or cultriform, with 
sharp or sometimes rounded angles in both sexes; 
the eyes are large, shape unusually broad, emar- 
ginate behind the antenna; eye facets are coarse, 
and never smaller than 0.022 mm in diameter. 
Based on these characters, five North American 
species are here transferred from Mordellisterta: 
Mordellirza blatchleyi (Liljeblad 1945); Mordellirta 
pilosella (Ray 1947); Mordelli~zapustulata (Melshe- 
imer 1845); Mordelli7~a ustulata (LeConte 1862); 
and Mordellina wickharni (Liljeblad 1945), [NEW 
COMBINATIONS]. This restores species rank to 
M. ustulata, which has been treated as a subspecies 
of Mordelliste7ta andreae LeConte 1862 (Liljeblad 
1945). 
Mordellochroa Emery 1876 
Mordellochroa has a t  least six species world- 
wide and is found in  Japan,  Mongolia, Poland, 
Hungary, and probably Russia. Batten (1977) pro- 
vided a key to the Palearctic species, Kaszab (1979) 
and Horhk (1993) provided keys for the species in 
Hungary, and Borowiec (1996) provided a key to 
species in Poland. 
Mordellochroa has  the penultimate tarsomeres 
of the fore and middle legs transversely truncate a t  
the apex, or very moderately emarginate, neither 
dilated nor bilobed; the apical palpomere of the 
maxilla is malleiform i n  males and securiform or 
scalene in  the females. Based on these characters, 
one species is here transferred from Mordellisterta: 
Mordellochroa scapz~laris (Say 1824), [NEW COM- 
BINATION]. This species has blacli elytra with 
humeral spots reddish to yellow and the lateral 
ridges on the hind tibia quite short. 
Pseudotolida Ermisch 1950 
Pseudotolida has fewer than 10 species in  Ja-  
pan, China, Taiwan, Guatemala, and Brazil. Fran- 
ciscolo (1982) provided a key to some species. Shiy- 
ake (1995, 1997) compared the genus to other 
related genera. 
Pseudotolida has the penultimate tarsomeres 
of the fore and middle legs deeply emarginate, more 
or less bilobed and usually strongly dilated; the 
apical palpomere of the maxilla is malleiform in 
males and securiform in  females. Based on these 
characters, three North American species are here 
transferred from Mordellisterta: Pseudotolida ari- 
d a  (LeConte 1862); Pseudotolida Kl~ausi (Liljeblad 
1945); and Pseudotolida lutea (Melsheimer 1845), 
[NEW COMBINATIONS]. 
Acknowledgments 
We thank J a n  Horhk for pointing out that  Mordelis- 
tens scap~~lal- is  (Say) belongs in  Mordellochroa. 
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Ermisch, K. 1941. Tribus Mordellistenini (Col. 
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ogischen Gesellschaft, 31: 710-726. 
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(J. Decelle, 1961-1964). 19. Coleoptera Mordel- 
lidae. (55. Beitrag zur Kenntnis der Mordel- 
liden). Musee Royal de L-Afrique Centrale, 
Tervuren, Belgique Annales, Serie in-8 , Sci- 
ences Zoologiques, 165: 257-287. 
Fabricius, J. C. 1798. Supplementum entomologi- 
ae systematicae, Hafniae, 572 pp. 
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African genera and species of Mordellidae. Part  
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Stenaliini. I~L South African Animal Life, Re- 
sults of the Lunds University Expedition in 
1950-1951, 4: 207-291. 
Franciscolo, M. 1965. A monograph of the South 
African genera and species of Mordellidae. Part 
2. Tribe Mordellini. I~L South African Animal 
Life, Results of the Lunds University Expedi- 
tion in 1950-1951, 11: 244-458. 
Franciscolo, M. 1967. A monograph of the South 
African genera and species of Mordellidae. Part  
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pedition in 1950-1951, 13: 67-203. 
Franeiscolo, M. 1982. Mordellidae (C~leop te~a :  
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PP. 
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Male genitalia and phylogenetic relationships 
in North American Moi-dellidae (C~leopte~a) .  
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ca, 90(6): 742-767. 
Melsheimer, F. E. 1845. Descriptions of new spe- 
cies of Coleoptera of the United States. Pro- 
ceedings of the Academy of Natural Sciences of 
Philadelphia, 2: 213-223, 302-318. 
Ray, E. 1946. Studies on North American Mordel- 
lidae, I11 (Coleoptera). Pan-Pacific Entomolo- 
gist. 22(3): 90-99. 
Ray, E. 1947. Studies on North American Mordel- 
lidae, V (Coleoptera). Pan-Pacific Entomolo- 
gist, 23(3): 121-131. 
Say, T. 1824. Descriptions of coleopterous insects 
collected in the late expedition to the Rocky 
Mountains, performed by order of Mr. Cal- 
houn, Secretai-y of War. under cominand of 
Major Long. Journal of the Academy of Natural 
Sciences of Philadelphia, 3(2): 238-282, 298- 
331, 403-464; 4(1): 83-99. 
Say, T. 1835. Descriptions of North American co- 
leoptero-cts insects and observations on some 
already described. Boston Journal of Natural 
History, l(2): 151-203. 
3 4 Volume 15, No. 1, March, 2001, INSECTA MUNDI 
Schilsky, J. 1908. XII. Beitrag zur deutschen Shiyake, S. 1997. Two new species of the genus 
Kaferfauna. Deutsche Entomologische Zeitschr- Tolidopalpz~s from the Philippines (Coleoptera, 
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Shiyake, S. 1995. A taxonomic study on the genus 52(1): 51-54. 
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(Coleoptera: Mordellidae). Bulletin of the Osa- ni. In Y. Kurosawa e t  al. (eds.), The Coleoptera 
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Nomenclatural changes for selected Mordellidae (Coleoptera) in North America

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