Olenekian (Early Triassic) fossil assemblage from eastern Julian Alps (Slovenia)

New palaeontological and sedimentological data from the Lower Triassic strata of the eastern Julian Alps in Slovenia are presented., They are unusual for the Early Triassic of the Alps in representing a relatively deeper, unrestricted marine (mid-ramp) setting. There are two basic microfacies types in the section investigated (types A and B), which are organized as couplets with coarse-grained tempestitic deposits (microfacies A), overlain by laminated or bioturbated lime mudstones and/or marls (microfacies B), frequently containing ammonoids. This pattern is interpreted as storm deposition with occasional winnowing of bottom sediments and the formation of coarse-grained skeletal deposits (lags), followed by the slow settling of suspended particles, when the storm waned, in addition to background deposition. Dominantly lime mud deposition and the presence of ammonoids indicate deposition on a more distal, deeper ramp with an unrestricted connection to the open sea. Intense reworking of bottom skeletal-rich sediment and accumulation of storm lags suggest deposition above the storm wave base, possibly in a wide low-energy mid-ramp environment. Faunas from such settings have been reported relatively rarely from the Early Triassic of the Alps. The macrofauna contains ammonoids, bivalves and gastropods, whereas the microfauna is represented by foraminifer tests and conodont elements; rare fish remains also occur. In the foraminifer assemblages, species of Ammodiscus, Hoyenella, Glomospirella dominated, corresponding to the widespread “Glomospira-Glomospirella” foraminifer community, with some miliolids and nodosariids. The conodont fauna is characterized by Triassospathodus hungaricus (Kozur et Mostler), indicating an early Spathian (Olenekian) age. The fossil assemblage highlights the wide distribution of Early Triassic taxa in the Tethys and facilitates its worldwide correlation. Its relatively low diversity by comparison with shallow marine settings is interpreted as an evolutionary proximal-distal trend in the wake of the end-Permian mass extinction. Re-diversification first occurred in nearshore settings and expanded into deeper/distal marine environments through geological time

CONODONT BIOSTRATIGRAPHY AND LITHOSTRATIGRAPHY ACROSS THE PERMIAN-TRIASSIC BOUNDARY AT THE LUKAC SECTION IN WESTERN SLOVENIA

Detailed conodont biostratigraphy and lithostratigraphy of the Late Permian and Early Triassic beds were studied at the LukaC section in western Slovenia. The analyzed section is composed of the Bellerophon Formation ("evaporite-dolomite member") and the newly introduced Lukaè Formation ("transitional beds", "streaky limestone member" and "carbonate-clastic beds member"). The Permian-Triassic boundary interval is represented by "transitional beds" of carbonate facies deposited in shallow restricted marine conditions. The presence of H. parvus in sample L1 in the "transitional beds" marks the systemic boundary between Permian and Triassic. The studied interval is characterized by a diverse microfauna that contain conodonts, foraminifers, ostracods and gastropods. Six conodont zones have been recognized, in ascending order, the latest Changhsingian (uppermost Permian) praeparvus Zone, and the Griesbachian (lowermost Triassic) parvus, lobata, staeschei-isarcica, postparvus and anceps zones. This faunal succession represents the first known and the most complete conodont biozonation across the Permian-Triassic interval from the entire Dinaric region. The recognized conodont biozones can be correlated with the biozonation of the Southern Alps and of the GSSP Meishan D section.

THE LOWER TRIASSIC SHALLOW MARINE SUCCESSION IN GORSKI KOTAR REGION (EXTERNAL DINARIDES, CROATIA): LITHOFACIES AND CONODONT DATING

The paper aims to present Lower Triassic lithofacies definition and first conodont fauna of Gorski Kotar region, Croatia. The depositional environment is envisaged as shallow marine realm of a passive continental margin. Sedimentary complex differentiates in predominantly carbonate sedimentation that characterises the beginning of deposition with upward increasing trend of terrigeneous influx. Lithofacial units have been defined as oolitic bar facies, lagoonal facies, shoreface-offshore facies, ooid-sandy shoal facies, restricted bay facies and flat-pebble conglomerate facies.The following conodont taxa were collected: Ellisonia sp., Foliella gardenae, Hadrodontina sp., Hindeodus parvus, Hindeodus sp., Pachycladina obliqua, ?Parachirognathus sp., Platyvillosus costatus and Pl. hamadai. The oldest strata yield Hindeodus parvus marking lowermost Triassic. The biostratigraphical data enable recognition of the parvus-isarcicella zones, obliqua Zone and Platyvillosus Subzone. The finds of Hindeodus parvus, Platyvillosus costatus and Pl. hamadai represent their first records in the External Dinarides and enable correlation of the Early Triassic conodont faunas of the Western Tethyan realm

CONODONT DATING OF THE LOWER TRIASSIC SEDIMENTARY ROCKS IN THE EXTERNAL DINARIDES (CROATIA AND BOSNIA AND HERZEGOVINA)

Two Lower Triassic sedimentary successions have been dated by means of conodonts in the External Dinarides: Plavno section near Knin, Croatia and Bosansko Grahovo section in Bosnia and Herzegovina. Deposition in both sections shows similar characteristics, differentiated in three continuously deposited facies. The Siliciclastic facies was previously considered Seis beds and assigned to the lower Lower Triassic, the Mudstone facies, and the Siltstone-mudstone facies (occurring in the upper part of the succession) were formerly considered as Campil beds of the upper Lower Triassic. Vertical succession of Siliciclastic, Mudstone, and Siltstone-mudstone facies of both investigated sequences was interpreted as deepening of the environment envisaged as a transgressive trend in a shallow shelf environment. Facies successions at Plavno (690 m thick) and Bosansko Grahovo (229 m thick) differentiate for the presence of Dolostone facies in the lowest part of the Plavno succession. Conodont fauna of Dolostone facies at Plavno section is represented by isarcicellids, Isarcicella staeschei and I. isarcica (sample 3) that marks the Griesbachian isarcica Zone. The Siliciclastic facies of Plavno and Bosansko Grahovo sections is characterized by shallow-water euryhaline taxa attributed to the Smithian, part of the late Dinerian-Smithian obliqua Zone. This fauna is prevailed by Hadrodontina anceps and Pachycladina obliqua with co-occurrence of Smithian Parachirognathus ethingtoni and very rare presence of Foliella sp. or ?Furnishius sp. Discerned conodont taxa enable us to establish conodont zonation which gives new insight to the range of the so-called Siusi and Campil beds.

Suppressed competitive exclusion enabled the proliferation of Permian/Triassic boundary microbialites

During the earliest Triassic microbial mats flourished in the photic zones of marginal seas, generating widespread microbialites. It has been suggested that anoxic conditions in shallow marine environments, linked to the end‐Permian mass extinction, limited mat‐inhibiting metazoans allowing for this microbialite expansion. The presence of a diverse suite of proxies indicating oxygenated shallow sea‐water conditions (metazoan fossils, biomarkers and redox proxies) from microbialite successions have, however, challenged the inference of anoxic conditions. Here, the distribution and faunal composition of Griesbachian microbialites from China, Iran, Turkey, Armenia, Slovenia and Hungary are investigated to determine the factors that allowed microbialite‐forming microbial mats to flourish following the end‐Permian crisis. The results presented here show that Neotethyan microbial buildups record a unique faunal association due to the presence of keratose sponges, while the Palaeotethyan buildups have a higher proportion of molluscs and the foraminifera Earlandia. The distribution of the faunal components within the microbial fabrics suggests that, except for the keratose sponges and some microconchids, most of the metazoans were transported into the microbial framework via wave currents. The presence of both microbialites and metazoan associations were limited to oxygenated settings, suggesting that a factor other than anoxia resulted in a relaxation of ecological constraints following the mass extinction event. It is inferred that the end‐Permian mass extinction event decreased the diversity and abundance of metazoans to the point of significantly reducing competition, allowing photosynthesis‐based microbial mats to flourish in shallow water settings and resulting in the formation of widespread microbialites