The Sage-Grouse Habitat Mortgage: Effective Conifer Management in Space and Time

AbstractManagement of conservation-reliant species can be complicated by the need to manage ecosystem processes that operate at extended temporal horizons. One such process is the role of fire in regulating abundance of expanding conifers that disrupt sage-grouse habitat in the northern Great Basin of the United States. Removing conifers by cutting has a beneficial effect on sage-grouse habitat. However, effects may last only a few decades because conifer seedlings are not controlled and the seed bank is fully stocked. Fire treatment may be preferred because conifer control lasts longer than for mechanical treatments. The amount of conservation needed to control conifers at large temporal and spatial scales can be quantified by multiplying land area by the time needed for conifer abundance to progress to critical thresholds (i.e., “conservation volume”). The contribution of different treatments in arresting conifer succession can be calculated by dividing conservation volume by the duration of treatment effect. We estimate that fire has approximately twice the treatment life of cutting at time horizons approaching 100 yr, but, has high up-front conservation costs due to temporary loss of sagebrush. Cutting has less up-front conservation costs because sagebrush is unaffected, but it is more expensive over longer management time horizons because of decreased durability. Managing conifers within sage-grouse habitat is difficult because of the necessity to maintain the majority of the landscape in sagebrush habitat and because the threshold for negative conifer effects occurs fairly early in the successional process. The time needed for recovery of sagebrush creates limits to fire use in managing sage-grouse habitat. Utilizing a combination of fire and cutting treatments is most financially and ecologically sustainable over long time horizons involved in managing conifer-prone sage-grouse habitat

Restoring North America’s Sagebrush Steppe Ecosystem Using Seed Enhancement Technologies

Rangelands occupy over a third of global land area, and in many cases are in less than optimum condition as a result of past land use, catastrophic wildfire and other disturbance, invasive species, or climate change. Often the only means of restoring these lands involves seeding desirable species, yet there are few cost effective seeding technologies, especially for the more arid rangeland types. The inability to consistently establish desired plants from seed may indicate that the seeding technologies being used are not successful in addressing the primary sources of mortality in the progression from seed to established plant. Seed enhancement technologies allow for the physical manipulation and application of materials to the seed that can enhance germination, emergence, and/or early seedling growth. In this article we examine some of the major limiting factors impairing seedling establishment in North America’s native sagebrush steppe ecosystem, and demonstrate how seed enhancement technologies can be employed to overcome these restoration barriers. We discuss specific technologies for: (1) increasing soil water availability; (2) enhancing seedling emergence in crusting soil; (3) controlling the timing of seed germination; (4) improving plantability and emergence of small seeded species; (5) enhancing seed coverage of broadcasted seeds; and (6) improving selectivity of pre-emergent herbicide. Concepts and technologies in this paper for restoring the sagebrush steppe ecosystem may apply generally to semi-arid and arid rangelands around the globe

Saving the sagebrush sea: An ecosystem conservation plan for big sagebrush plant communities

Vegetation change and anthropogenic development are altering ecosystems and decreasing biodiversity. Successful management of ecosystems threatened by multiple stressors requires development of ecosystem conservation plans rather than single species plans. We selected the big sagebrush (Artemisia tridentata Nutt.) ecosystem to demonstrate this approach. The area occupied by the sagebrush ecosystem is declining and becoming increasingly fragmented at an alarming rate because of conifer encroachment, exotic annual grass invasion, and anthropogenic development. This is causing rangewide declines and localized extirpations of sagebrush associated fauna and flora. To develop an ecosystem conservation plan, a synthesis of existing knowledge is needed to prioritize and direct management and research. Based on the synthesis, we concluded that efforts to restore higher elevation conifer-encroached, sagebrush communities were frequently successful, while restoration of exotic annual grass-invaded, lower elevation, sagebrush communities often failed. Overcoming exotic annual grass invasion is challenging and needs additional research to improve the probability of restoration and identify areas where success would be more probable. Management of fire regimes will be paramount to conserving sagebrush communities, as infrequent fires facilitate conifer encroachment and too frequent fires promote exotic annual grasses. Anthropogenic development needs to be mitigated and reduced to protect sagebrush communities and this probably includes more conservation easements and other incentives to landowners to not develop their properties. Threats to the sustainability of sagebrush ecosystem are daunting, but a coordinated ecosystem conservation plan that focuses on applying successful practices and research to overcome limitations to conservation is most likely to yield success

Abundances of Coplanted Native Bunchgrasses and Crested Wheatgrass after 13 Years

Crested wheatgrass (Agropyron cristatum [L] Gaertm) has been seeded on more than 5 million hectares in western North America because it establishes more readily than native bunchgrasses. Currently, there is substantial interest in reestablishing native species in sagebrush steppe, but efforts to reintroduce native grasses into crested wheatgrass stands have been largely unsuccessful, and little is known about the long-term dynamics of crested wheatgrass/native species mixes. We examined the abundance of crested wheatgrass and seven native sagebrush steppe bunchgrasses planted concurrently at equal low densities in nongrazed and unburned plots. Thirteen years post establishment, crested wheatgrass was the dominant bunchgrass, with a 10-fold increase in density. Idaho fescue (Festuca idahoensis Elmer), Thurber’s needlegrass (Achnatherum thurberianum (Piper) Barkworth), basin wildrye (Leymus cinereus [Scribn. & Merr.] A. Löve), and Sandberg bluegrass (Poa secunda J. Presl) maintained their low planting density, whereas bluebunch wheatgrass (Pseudoroegneria spicata [Pursh] A. Löve), needle-and-thread (Hesperostipa comata [Trin. & Rupr.] Barkworth), and squirreltail (Elymus elymoides [Raf.] Swezey) densities declined. Our results suggest that densities of native bunchgrasses planted with crested wheatgrass are unlikely to increase and that some species may only persist at low levels. The high recruitment of crested wheatgrass suggests that coplanting of some native bunchgrasses may be a viable way of avoiding crested wheatgrass monocultures when this species is necessary for rehabilitation or restoration.Keywords: Agropyron cristatum, Revegetation, Sagebrush steppe, Restoratio

Of Grouse and Golden Eggs: Can Ecosystems Be Managed Within a Species-Based Regulatory Framework?

Declining greater sage-grouse populations are causing concern for the future of this species across the western United States. Major ecosystem issues, including exotic annual grass invasion and conifer encroachment, threaten vast acreages of sagebrush rangeland and are primary threats to sage-grouse. We discuss types of problems facing sage-grouse habitat and argue that complex ecosystem problems may be difficult to address under the Endangered Species Act as currently applied. Some problems, such as anthropogenic development, can be effectively regulated to produce a desired outcome. Other problems that are complex and involve disruption of ecosystem processes cannot be effectively regulated and require ongoing commitment to adaptive management. We believe that historical inertia of the regulatory paradigm is sufficient to skew management toward regulatory mechanisms, even though complex ecosystem problems impact large portions of the sage-grouse range. To overcome this situation, we suggest that the regulatory approach embodied in the Endangered Species Act be expanded to include promoting management trajectories needed to address complex ecosystem problems. This process should begin with state-and-transition models as the basis for a conceptual framework that outlines potential plant communities, their value as sage-grouse habitat, and their ecological status. Desired management trajectories are defined by maintenance of an ecologically resilient state that is of value as sage-grouse habitat, or movement from a less desired to a more desired state. Addressing complex ecosystem problems will involve shifting conservation roles. Under the regulatory approach, programmatic scales define regulatory policies, and local scales focus on implementing those policies. With complex ecosystem problems, programmatic scales empower local conservationists to make decisions necessary to adaptively manage problems. Putting ecosystem management on par with traditional regulatory actions honors obligations to provide regulatory protections while maintaining the capacity of the ecosystem to produce habitat and greatly expands the diversity of stakeholders willing to participate in sage-grouse conservation.Keywords: Sage-grouse, Endangered Species Act, State-and-transition, Sagebrus

Global change effects on plant communities are magnified by time and the number of global change factors imposed

Global change drivers (GCDs) are expected to alter community structure and consequently, the services that ecosystems provide. Yet, few experimental investigations have examined effects of GCDs on plant community structure across multiple ecosystem types, and those that do exist present conflicting patterns. In an unprecedented global synthesis of over 100 experiments that manipulated factors linked to GCDs, we show that herbaceous plant community responses depend on experimental manipulation length and number of factors manipulated. We found that plant communities are fairly resistant to experimentally manipulated GCDs in the short term (<10 y). In contrast, long-term (≥10 y) experiments show increasing community divergence of treatments from control conditions. Surprisingly, these community responses occurred with similar frequency across the GCD types manipulated in our database. However, community responses were more common when 3 or more GCDs were simultaneously manipulated, suggesting the emergence of additive or synergistic effects of multiple drivers, particularly over long time periods. In half of the cases, GCD manipulations caused a difference in community composition without a corresponding species richness difference, indicating that species reordering or replacement is an important mechanism of community responses to GCDs and should be given greater consideration when examining consequences of GCDs for the biodiversity–ecosystem function relationship. Human activities are currently driving unparalleled global changes worldwide. Our analyses provide the most comprehensive evidence to date that these human activities may have widespread impacts on plant community composition globally, which will increase in frequency over time and be greater in areas where communities face multiple GCDs simultaneously

Limitations to Postfire Seedling Establishment: The Role of Seeding Technology, Water Availability, and Invasive Plant Abundance

Seeding rangeland following wildfire is a central tool managers use to stabilize soils and inhibit the spread of invasive plants. Rates of successful seeding on arid rangeland, however, are low. The objective of this study was to determine the degree to which water availability, invasive plant abundance, and seeding technology influence postfire seedling establishment. Across four fire complexes, whole plots were either seeded using a rangeland drill, seeded by hand where seeds could be placed at an exact depth, or left as unseeded controls. Irrigation and weeding treatments were applied to subplots within whole plots in an incomplete factorial design. In three of the four fires, seeding method was the single factor limiting establishment with seedling density over sevenfold higher in the hand-seeded compared to the drill-seeded treatments. In contrast to our hypotheses, water and weeding had no positive effect on seedling establishment in any of the four fires; however, background weed density was relatively low. The native community recovered at all sites with minimal bunchgrass mortality. These results strongly suggest a need for a decision framework that evaluates postfire seeding needs relative to natural recovery. Based on these initial results, it appears modest improvements in seeding technology may yield substantial increases in seeding success. The Rangeland Ecology & Management archives are made available by the Society for Range Management and the University of Arizona Libraries. Contact [email protected] for further information.Migrated from OJS platform August 202

Regrowth and production of herbaceous riparian vegetation following defoliation

Stubble height regulations are frequently used to manage livestock grazing of herbaceous riparian vegetation. The objective of this study was to determine the impact of stubble height, time of clipping and soil water status on production and regrowth of herbaceous riparian vegetation. We used a randomized block design with 4 study sites on each of 3 small ( 2m width) streams in northern Harney County, Ore. In June and July of 2000-2003, 40 × 50 cm experimental plots were clipped to stubble heights of 5.1 (2 inch), 10.2 (4 inch), or 15.2 cm (6 inch), and paired control plots were left unclipped. Complete treatment sets were located adjacent to the stream and 4 m from the stream at each site. All plots were clipped to 1 cm in October and regrowth was calculated by comparing clipped and control plots. Water table depth was measured weekly using PVC wells. Results indicate that height regrowth was associated positively with stubble height (P < 0.01) and was less with July compared to June clipping (P = 0.02). Weight regrowth was also positively related to stubble height (P < 0.01) and decreased with July compared to June clipping (P = 0.04) whereas annual aboveground production increased with July clipping (P = 0.02). Annual production values for clipped plots were higher than for unclipped plots, indicating compensatory production in response to defoliation. Plots distant from the stream had less water availability, but regrowth and production were not strongly influenced by distance from active stream channel. Timing and intensity of defoliation were reliable predictors of regrowth and production performance. Most clipping height × time combinations produced end of season heights sufficient to meet current federal stubble height requirements (i.e., 10-15 cm). Our results provide insight on the timing and intensity of defoliation that will allow for adequate regrowth to meet different management objectives. However, other factors such as stream channel morphology, animal selectivity, and annual weather variation will need to be considered.The Journal of Range Management archives are made available by the Society for Range Management and the University of Arizona Libraries. Contact [email protected] for further information.Migrated from OJS platform August 202

Response of Bluebunch Wheatgrass and Medusahead to Defoliation

Our objective was to determine the short-term response of bluebunch wheatgrass and medusahead to defoliation of wheatgrass designed to stimulate regrowth through tillering. We hypothesized that defoliating bluebunch wheatgrass by 20% at the 3 to 3.5 leaf stage followed by a 50% defoliation at peak standing crop would increase its tillering and biomass production. Consequently, we expected a reduction of the density and biomass of medusahead over that of bluebunch wheatgrass defoliated 50% at peak standing crop. Treatments included four initial medusahead densities (200, 333, 444, 600 plants m-2) created by hand-pulling and three defoliation regimes factorially arranged (12 treatment combinations) in a randomized complete-block design and replicated four times at two sites. In 2006 and 2007, defoliation was accomplished by hand-clipping bluebunch wheatgrass 1) by 50% once at peak standing crop (late June); 2) by 20% at the 3 to 3.5 leaf stage, then again to 50% at peak standing crop (mid May, late June); or 3) plants were not clipped. Density was sampled in 2006 and 2007, and biomass was harvested only at Star Mountain (near Riverside, Oregon) in 2007 because Warm Springs (near Drewsey, Oregon) was burned by a wildfire before final 2007 data could be collected. In 2006, no treatments applied at either site detectably altered the number of tillers produced by bluebunch wheatgrass nor did they affect bluebunch wheatgrass density or biomass in 2007 at Star Mountain. Changes in medusahead density were not detected in 2006, but this annual invasive grass increased in density and biomass in 2007 at Star Mountain in plots receiving two defoliations. The relatively short growing period caused by summer drought and the relative intolerance of bluebunch wheatgrass to grazing make the twice-over grazing an unlikely practice for arid rangelands in the western United States. In fact, it could possibly increase the risk of annual grass invasion. The Rangeland Ecology & Management archives are made available by the Society for Range Management and the University of Arizona Libraries. Contact [email protected] for further information.Migrated from OJS platform August 202