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Intensity of larval diapause in the bamboo borer, Omphisa fuscidentalis

Larvae of the bamboo borer, Omphisa fuscidentalis, enter larval diapause in September and pupate in the following June (Singtripop et al., 1999). We examined the changes in the responses of larvae to exogenous 20-hydroxyecdysone (20E) in order to estimate the progress of diapause development. In this respect, we adopted two terms, responsiveness and sensitivity of larvae to 20E. Responsiveness was estimated by the percentage of larvae that pupated, and sensitivity was evaluated by the duration from the day of 20E injection to pupation. The responsiveness of larvae declined gradually from September to November when larvae were least responsive to 20E, and then increased markedly from January to February. This indicates that the intensity of diapause increases from September to November and terminated gradually thereafter. Thus the sequence of events as the larval responses to 20E is characterized by a V-shaped curve. Sensitivity of larvae to 20E was at the same level from September to December, and increased remarkably from December to January. The abrupt increase in the sensitivity occurred one month earlier than the bottom of the V-shaped curve of larval responsiveness, suggesting that the increases in the responsiveness and sensitivity in the latter hall of diapause may be brought about by respective mechanisms

Larval growth and diapause in a tropical moth, Omphisa fuscidentalis hampson

The bamboo borer, Omphisa fuscidentalis, is a moth found in northern Thailand, Lao and Myanmar and its larvae feed on the inner pulp of bamboo shoots. In a tropical highland (about 500 m sea level) forest at 19°N near Chiang Mai, Thailand, the larvae feed on at least 5 bamboo species. Nucleotide sequence analysis of the region of mitochondrial cytochrome C oxidase subunit 1 gene amplified by the polymerase chain reaction (PCR) verified that larvae collected from different bamboos belong to the same species. Adults appeared in early August and laid clusters of eggs on the newly grown bamboo shoot. The newly hatched larvae bore a hole in the shoot, enter an internode of the shoot and feed on the inner pulp. After maturation in September, the larvae remain in an internodal cavity of bamboo for up to 9 months, from September to the following June. Number of larval instars was estimated by measuring the width of head capsules remained in internodes of bamboo shoots. The growth curve of the width fitted to Dyar\u27s law and the mature larvae were estimated to be 5th instar. Mature larvae were collected in the field each month and their body weight, head capsule width, protein and fat contents and hemolymph ecdysteroid titer were measured. Body weight continuously decreased during the 9 months whereas head capsule width remained constant. Fat content fluctuated during this period while protein level remained at a similar level until March, after which it significantly increased. During this period, hemolymph ecdysteroid concentrations remained low. Current results show that the bamboo borer larvae enter diapause at the end of feeding period of the fifth (last) larval instar and the larval diapause lasts until June

Clubiona octoginta Dankittipakul, sp. n.

Clubiona octoginta Dankittipakul sp. n. (Figs 1, 15–19, 41– 47) Type material: Holotype: ɗ, Pha Tam National Park [Khong Chiam District, Ubon Ratchathani Province, Northeastern Thailand], deciduous forest, 250 m, 15.– 20.xi. 2006, P. Dankittipakul (MHNG). Paratypes: 3 ɗ, 2 Ψ, same data as holotype (MHNG, TNHM). Etymology. The specific epithet is a Latin numeral meaning eighty. This new species is named in honor of His Majesty King Bhumibol Adulyadej the Great of Thailand who has served his country with indefatigable dedication. His Majesty is the longest-ruling monarch in Thailand's history and the longest-reigning ruler in the world. This new species is described to commemorate the celebrations on the most auspicious occasion of His Majesty the King’s 80 th Birthday Anniversary on December 2007. Diagnosis. Clubiona octoginta sp. n. is closely related to C. campylacantha sp. n. Although differences between them on the basis of genitalia are mostly trivial, they are not conspecific and represent distinct species. Clubiona octoginta sp. n. is easily separable in having different opisthosomal color pattern (Fig. 41 vs. Fig. 34). The conical embolus of C. octoginta sp. n. is more reduced and the triangular retrolateral tibial apophysis is apparently shorter and broader at base compared to that of C. campylacantha sp. n. Clubiona octoginta sp. n. is also more slender than C. campylacantha sp. n., which is relatively large and stoutly built species. The separate status of C. octoginta sp. n. is further justified by differences in female genitalia. In C. octoginta sp. n., the spermathecal head is provided with pear-shaped terminal apparatus (Fig. 19). In contrast, this structure represents a short, digitiform tubercle in the females of C. campylacantha sp. n. (Fig. 14). The female of C. octoginta sp. n. also resembles that of C. scandens known from Borneo in similar pattern of the vulva in which the spermathecal heads are indistinct although the terminally dilated trigonal apparatuses are conspicuously enlarged (Figs 19, 47; see also Deeleman-Reinhold 2001: figs 58–59). The most obvious difference is the shape of epigynal atrium, which in C. scandens can be recognized by the columnar depression with indistinct anterior atrial margin. Also, the terminal portions of spermathecal heads are much bigger in C. scandens than in C. octoginta sp. n., covering most anterior part of the vulva but hidden behind the spermathecal heads in this new species (Fig. 18). Description. Male (holotype): Total 5.4; prosoma 2.4 long, 1.7 wide; opisthosoma 3.0 long, 1.5 wide. Prosoma (Fig. 41). Ovoid, pars cephalica relatively broad, in profile slightly higher behind longitudinal fovea; integument smooth. Carapace yellow, anteriorly brown, a pair of Y-shaped purplish markings starting from behind PME and PLE almost reaching dark reddish fovea. Chelicerae dark brown, dorsally with dark pattern. Labium and endites pale brown. Sternum pale yellow. Eyes. Eyes with black rings and surrounded by purplish markings. Eye sizes and interdistances: AME 0.11, ALE 0.11, PME 0.11, PLE 0.11, AME–AME 0.11, AME–ALE 0.06, PME–PME 0.28, PME–PLE 0.18; MOQ: 0.28 long, 0.36 anterior width, 0.51 posterior width. Legs. Yellow, femora with dark brown ventral band; tibiae with dark distal and proximal annuli; anterior metatarsi with distal dark band; tarsi brown. Leg formula 4213. Leg measurements: I II III IV Femora 1.40 1.55 1.32 1.60 Patellae + tibiae 1.86 1.95 1.45 1.91 Metatarsi 0.85 0.95 1.20 1.64 Tarsi 0.53 0.55 0.50 0.51 Total 4.64 5.00 4.47 5.66 Opisthosoma (Fig. 41). Elongate-oval; dorsum yellow, with broken dark median band, reaching half opisthosoma length; a pair of broken lateral bands fused posteriorly; venter pale, without markings. Male palp (Figs 15 –16, 42– 44). Retrolateral tibial apophysis trigonal, distinctly broad at base, its apex sharply pointed. Cymbium longer than wide, slightly excavated dorso-apically. Bulb oval, with indistinct anterio-proapically membranous portion; sperm duct sinuate, forming a double loop. Embolus represented by short, triangular tubercle, broad at base, gradually tapering toward its apex, apico-prolaterally fused with curved conductor. Female (paratype): Total length 5.9; prosoma 2.2 long, 1.6 wide; opisthosoma 3.7 long, 2.2 wide. Resembling the males but slightly larger in size (Fig. 1); prosoma narrowed in cephalic area; dorsal shield of prosoma with indistinct color marking; dorsum of opisthosoma with pattern less defined. Eyes. Eyes with black rings and surrounded by purplish markings. Eye sizes and interdistances: AME 0.10, ALE 0.10, PME 0.11, PLE 0.10, AME–AME 0.06, AME–ALE 0.03, PME–PME 0.23, PME–PLE 0.11; MOQ: 0.28 long, 0.31 anterior width, 0.46 posterior width. Legs. Leg measurements: I II III IV Femora 1.30 1.40 1.23 1.54 Patellae + tibiae 1.82 1.95 1.46 1.90 Metatarsi 0.88 0.92 1.20 1.70 Tarsi 0.51 0.53 0.46 0.55 Total 4.51 4.77 4.35 5.69 Epigyne and vulva (Figs 17 –19, 45– 47). Anterior epigynal atrium trapezoid, margin rebordered, anterior atrial margin straight. Genital orifices situated on posteriorlateral part of the atrium. Spermathecal stalks short; spermathecal head with triangular terminal apparatus carrying numerous minute pores on its anterior portion; spermathecal bases elongated tubular, encircling small fertilization ducts. Membranous bursae rounded, situated posterior to the spermathecae. Natural history. The type specimens were collected from dry deciduous forests. They were all obtained from a Malaise trap in November. The national park is predominately covered with dry dipterocarp forests. Distribution. Known only from the type locality, Pha Tam National Park, northeastern Thailand (Fig. 69).Published as part of Dankittipakul, Pakawin & Singtripop, Tippawan, 2008, Five new species of the spider genus Clubiona Latreille (Araneae: Clubionidae) from Thailand, pp. 34-60 in Zootaxa 1747 on pages 39-41, DOI: 10.5281/zenodo.18167

The spitting spider family Scytodidae in Thailand, with descriptions of three new Dictis species (Araneae)

Volume: 117Start Page: 121End Page: 14

Systaria Simon 1897

Systaria Simon, 1897 Systaria Simon, 1897 a: 87. Lehtinen 1967: 237; Deeleman-Reinhold 2001: 202. Hebrithele Berland, 1938: 137 (synonymised by Deeleman-Reinhold 2001: 202). Type species by original designation: S. drassiformis Simon, 1897. Diagnosis. Representatives of Systaria can be distinguished from all other Asian Systariinae by a combination of characters: the dark and large AME; the widely separated anterior lateral spinnerets; the lengthened distal segment of posterior lateral spinnerets in females; the broad sperm duct running along outer margin of the tegulum; the lack of a median apophysis; the sclerotized bursae located anterior to smaller spermathecae. Members of Systaria can be distinguished from those of Xantharia by the absence of an oblique depression on gnathocoxae, the oval sternum being narrower anteriorly than in the middle, and the tarsi are covered with denser claw tufts. Members of Tamin can be distinguished from those of Systaria by having smaller AME, the sparse claw tuft and the widely separated gnaphosid-like anterior lateral spinnerets are provided with long spigots in males (AME being largest and anterior lateral spinnerets being distinctively shorter in males than in females in Systaria species). Description. Large to medium-sized clubionid spiders without distinctive coloration pattern. Prosoma oval, longer than wide, widest between coxae II and III, narrowed in front but always more than 2 / 3 its maximum width; thoracic groove well defined; fovea deep, longitudinal. AME and PME circular, ALE and PLE oval, AME dark, largest, other pearly white, subequal in size; PER slightly wider than AER, AER recurved, PER slightly procurved; AME separated by less than their radius, less than their diameter apart from ALE; PME separated by twice their diameter, slightly farther from PLE; ALE and PLE separated by almost their diameter. MOQ slightly longer than wide, posteriorly wider than anteriorly; clypeus height at AME, twice their diameter. Chelicerae long and slender, vertical, with slight lateral condyle; 3 promaginal and 2–3 retromarginal teeth; fang moderately long. Gnathocoxae longer than wide, slightly wider anteriorly, parallel or slightly excavated medially on lateral margins, without oblique depression, slightly divergent, with anterior median scopula. Labium slightly longer than wide, rebordered. Sternum oval, domed, slightly narrower anteriorly than in the middle, with triangular sclerotized extensions to or between coxae. Leg formula 4123, prograde; trochanters deeply notched; femora with strong spines; patellae short, without spine; anterior tibiae with conspicuous ventral spines; anterior metatarsi with weak ventral spines; metatarsi and tarsi with distoventral scopulae; tarsi with dense claw tuft. Two tarsal claws, dentate. Opisthosoma ovoid, tapering posteriorly, covered with dense bristles, without dorsal scutum, some species with brownish gray reticulations. Six spinnerets, anterior lateral spinnerets cylindrical, tapering apically, never contiguous, long, widely separated, usually up to one diameter apart, two-segmented, distal segment very short, with one major ampullate and several piriform gland spigots; posterior median spinnerets short, cylindrical in both sexes; posterior lateral spinnerets two-segmented, both segments long, distal segment carrying one large apical and several smaller spigots. Male palp with femur and patella unmodified; tibia with elevated distoventral hump, and complex retrolateral tibial apophysis. Cymbium with sharply pointed baso-retrolateral cymbial tubercle or oblique basal ridge; invaginated at base on retrolateral side, its surface partially membranous or lightly sclerotized, without pigmentation. Tegulum clearly separated from almost indistinct subtegulum, elongate-oval, protruding posteriorly, strongly excavated meso-prolaterally to accommodate embolus, with broad sperm duct running along its outer margin. Embolic base membranous, originating meso-prolaterally; embolus triangular, gradually tapering toward its apex. Median apophysis absent. Conductor membranous, situated apically. Epigyne lightly sclerotized, with shallow epigynal atrium and paired copulatory orifices. Atrium often provided with a pair of ridges, presumably to guide and support male copulatory organ towards insemination pore. Copulatory orifices leading to short insemination ducts which closely attached to posterior part of bursae. Bursae sclerotized, situated anteriorly, connected to spermathecae via mostly short ducts. Fertilization ducts variable in shape. After copulation, insemination duct often inflated, forming a spherical ball containing a plug of dark substance (Figs 14, 19, 25, 31); virgin females as in Figs 13, 37, 39, 40. Taxonomic remarks. The two following characters mentioned are considered synapomorphic and diagnostic for the genus. A sharply pointed tubercle situated at base on retrolateral side of the cymbium (= cymbial tubercle; Fig. 1, CT) is found in the males of S. bifida sp. nov., S. decidua sp. nov. and S. acuminata sp. nov. (Figs 9, 32). In other Systaria species this area is clearly elevated, forming an oblique basal ridge; its coloration is slightly darker than the rest of the cymbium. This modification can be easily observed from a retrolateral view of the male palp (Figs 10, 33). The females examined and treated in the present study all share a similar pattern of internal genitalia in which a pair of bursae situated anterior to posterior spermathecae are heavily sclerotized, often with the remaining of numerous gland ductules. Although there is no doubt that the Chinese species belong to Systaria, illustrations of their internal genitalia show a different pattern: the anterior bursae seem to be membranous. Zhang et al. (2009) documented that ‘Bursae situated anterior of copulatory orifices, large sac-shaped and semi -transparent ’. However, it appears to us that most females of Systaria possess sclerotized bursae (see also Deeleman-Reinhold 2001). It is of interest to note that in Systaria the entrance to the vulva (inseminations ducts) is often plugged, possibly to prevent further matings of the female (Deeleman-Reinhold, pers. comm.). The phenomenon of plugged copulatory organs in female spiders is known in a variety of families, but was not yet recognized and reported for Systariinae. Natural history. Most of the specimens of Systaria species treated in this study were directly obtained by digging their retreats inside the caves or nearby the cave entrances. Distribution. Southern China, Southeast Asia including New Guinea.Published as part of Dankittipakul, Pakawin & Singtripop, Tippawan, 2011, Seven new species of Systaria Simon, 1897 from Southeast Asia (Araneae, Clubionidae, Systariinae), pp. 16-32 in Zootaxa 2905 on pages 18-19, DOI: 10.5281/zenodo.20710

Systaria cervina Simon 1897

Systaria cervina (Simon, 1897) Figs 4–8 Syrisca cervina Simon, 1897 b: 500, description of female. Systaria cervina (Simon). Deeleman-Reinhold, 2001: 203, figs 236– 237, transfer from Syrisca. Type material. Holotype: Ƥ, PHILIPPINES: Rizal: Antipolo (MNHN, AR 13751), examined. [Antipolo is a city locating in the Province of Rizal, ca. 25 km east of Manila. At the back side of Antipolo Church is a large mystical cave known as Inday Nelly Mystical Cave. It is likely that the female syntype was collected from this cave.] Other material examined. PHILIPPINES: Quezon Province: 23, 1Ƥ, 2 juveniles (presumably female), Pagbilao, Cueva Balisen, 20 February 1975, leg. P. Strinati, (MHNG, Syst- 25449). Diagnosis. Males of S. cervina can be distinguished from those of S. elberti (Strand, 1913) by the sickleshaped RTA provided with a small dorsal incision in lateral view (Figs 5–6). Females can be distinguished from those of S. elberti by the anterior bursae abruptly narrowing, forming short, posterior stalks (Fig. 8) instead of being elongate-oval, and by the much narrower insemination ducts connected between anterior bursae and posterior spermathecae. The male palpal structure of S. cervina also closely resembles S. insulana (Rainbow, 1902) but can be recognized by the more elongated RTA which is greatly narrower at base, and by the smaller conductor. Description. Male (MHNG, Syst- 25449): Total length 7.38. Prosoma 3.32 long, 2.28 wide. Opisthosoma 4.06 long, 1.92 wide. Leg formula: 4123; I 10.32 (2.78, 1.36, 2.66, 2.38, 1.14); II 8.66 (2.56, 1.18, 2.38, 1.62, 0.92); III 7.78 (2.08, 0.82, 1.76, 2.18, 0.94); IV 11.56 (3.24, 1.02, 3.08, 3.16, 1.06). Spination. Femora I d 1 - 1 - 1, p 1 - 1 - 1, r 0-1 - 1; II d 1 - 1 - 1, p 1 - 1 - 1, r 0-1 -0; III d 1 - 1 - 1, p 1 - 1 -0, r 1 - 1 - 1; IV d 1 - 1 - 1, p 1 - 0-1, r 1 - 1 - 1. Tibiae: I v 2 - 2 - 1 - 1; II v 2 - 2 - 2; III d 1 -0-0, p 1 -0-0, r 2 - 2 - 2, v 2 - 2 - 2; IV d 0-1 -0, p 1 - 1-2, r 1 - 1 -0, v 2 - 2 - 2. Metatarsi: I v 1 -0-0; II v 1 - 1 - 1; III p 1 - 1 - 1, r 1 - 1-2, v 0-1 - 1; IV p 1 - 1 - 1, r 2 - 2 - 2, v 1 - 1 - 1. Eye size and interdistances: AME 0.12, ALE 0.08, PME 0.08, PLE 0.06; AME– AME 0.16, AME–ALE 0.14, PME–PME 0.22, PME–PLE 0.14, ALE–PLE 0.12; MOA 0.26 long, anterior width 0.28, posterior width 0.26. Palp (Figs 4–6). Ventral surface of tibia with elevated, distal swelling. RTA slender, sickle-shaped, with small dorsal incision (Figs 5–6). Cymbium fold partially membranous, membrane situated basally. Cymbial ridge dark reddish-brown, slightly elevated, running obliquely. Tegulum oval, excavated baso-prolaterally. Embolus triangular, membranous at base. Conductor membranous, flange-like, situated apically. Female (MHNG, Syst- 25449): Total length 10.52. Prosoma 4.80 long, 3.32 wide. Opisthosoma 5.72 long, 3.06 wide. Leg formula: 4123; I 8.88 (2.60, 1.30, 2.52, 1.62, 0.84); II 8.30 (2.54, 1.22, 2.22, 1.58, 0.74); III 7.94 (2.84, 0.82, 1.70, 1.86, 0.72); IV 12.42 (3.70, 1.24, 2.76, 3.22, 1.50). Spination. Femora: I d 1 - 1 - 1, p 1 - 1 -0, r 0-1 - 1; II d 1 - 1 - 1, p 1 - 1 -0, r 1 - 1 - 1; III d 1 - 1 - 1, p 0-1 - 1, r 1 - 1 - 1; IV d 1 - 1 - 1, p 1 - 1 - 1, r 0-1 -0. Tibiae: I v 2 - 1 - 1; II v 0-2 -0; III p 1 - 1 -0, r 1 - 0- 1, v 2 - 2 - 2; IV p 1 - 1 -0, r 1 - 1 -0, v 2 - 2 - 2. Metatarsi: II v 1 - 1 -0; III p 1 - 1 - 1, r 1 - 1-2, v 2 - 1 - 1; IV p 1 - 1-2, r 1 - 1-2, v 2 - 2 - 2. Eye size and interdistances: AME 0.16, ALE 0.12, PME 0.12, PLE 0.10; AME–AME 0.18, AME–ALE 0.16, PME– PME 0.24, PME–PLE 0.18, ALE–PLE 0.14; MOA 0.26 long, anterior width 0.38, posterior width 0.36. Genitalia (Figs 7–8). Epigynal plate triangular, weakly sclerotized. Copulatory orifices teardrop-shaped, situated medially. Anterior bursae sclerotized, rounded anteriorly, abruptly narrowing posteriorly. Single pair of spherical spermathecae situated posteriorly. Fertilization ducts lanceolated. Natural history. New specimens of S. cervina were collected from their retreats at the entrance of a huge limestone cave. Distribution. The Provinces of Rizal and Quezon, Luzon Island, the Philippines (Fig. 43).Published as part of Dankittipakul, Pakawin & Singtripop, Tippawan, 2011, Seven new species of Systaria Simon, 1897 from Southeast Asia (Araneae, Clubionidae, Systariinae), pp. 16-32 in Zootaxa 2905 on pages 20-21, DOI: 10.5281/zenodo.20710

Clubiona Latreille 1804

Clubiona Latreille, 1804 Type species: Clubiona pallidula (Clerck, 1757).Published as part of Dankittipakul, Pakawin & Singtripop, Tippawan, 2008, Five new species of the spider genus Clubiona Latreille (Araneae: Clubionidae) from Thailand, pp. 34-60 in Zootaxa 1747 on page 36, DOI: 10.5281/zenodo.18167

Systaria insolita Dankittipakul & Singtripop, 2011, sp. nov.

Systaria insolita sp. nov. Figs 26–31 Type material. Holotype: 3, THAILAND: Chaiyaphum Province: Tard Tone National Park, 300 m, 8 December 2006, leg. P. Dankittipakul (MHNG, Sys-04). Paratypes: THAILAND: Phitsanulok Province: 2 Ƥ, Nakhon Thai District, Phu Hin Rong Kla National Park, 800 m, rotten log, 20 November 2002, leg. P. Dankittipakul (MHNG, Sys-05). Diagnosis. Males of S. insolita sp. nov. can be recognized by a peculiar shape of the embolus which is rather short and stout (Fig. 26) instead of elongate and filiform as in other males of Systaria, and by the beak-shaped dorsal process on the RTA (Figs 27–29). Females can be recognized by a pair of hoods on epigynal atrium (Fig. 30), and by the peculiar shape of the anterior bursae (Fig. 31). Etymology. The specific epithet is a Latin adjective (insolitus,-a, -um), referring to an unusual form of the embolus. Description. Male (holotype): Total length 7.78. Prosoma 3.54 long, 2.42 wide. Opisthosoma 4.24 long, 2.02 wide. Leg formula: 4123; I 11.12 (2.90, 1.54, 2.80, 2.50, 1.38); II 9.38 (2.74, 1.28, 2.56, 1.80, 1.00); III 8.08 (2.22, 0.84, 1.86, 2.24, 0.92); IV 11.88 (3.36, 1.04, 3.16, 3.20, 1.12). Spination. Femora I d 1 - 1 - 1, p 1 - 1 - 1, r0- 0-1; II d 1 - 1 - 1, p 1 - 1 - 1, r 1 - 0-1; III d 1 - 1 - 1, p 0-1 - 1, r 1 - 0-1; IV d 1 - 1 - 1, p 0-1 - 1, r 1 - 1 - 1. Tibiae: I v 2 - 2 - 2; II v 2 - 2 - 1; III d 0-1 -0, p 1 - 1 - 1, r 2 - 2 - 2, v 2 - 2 - 2; IV d 0-1 -0, p 1 - 1 -0, r 1 - 1 -0, v 2 - 2 - 2. Metatarsi: I v 1 -0-0; II v 0-1 - 1; III p 1 - 1-2, r 1 - 1 - 1, v 1 - 0-1; IV p 1 - 1 - 1, r 2 - 2 - 2, v 1 - 1 - 1. Eye size and interdistances: AME 0.12, ALE 0.08, PME 0.08, PLE 0.06; AME–AME 0.16, AME–ALE 0.14, PME–PME 0.22, PME–PLE 0.16, ALE–PLE 0.12; MOA 0.28 long, anterior width 0.30, posterior width 0.28. Palp (Figs 26–29). Ventral surface of tibia with small, elevated hump (Fig. 27). RTA elongated; ventral process hook-shaped, directed anteriad (Figs 27, 29); dorsal process sharply pointed, directed posteriad (Figs 26, 29). Cymbial fold partially membranous. Cymbial ridge slightly elevated. Embolus triangular, its apex short and stout; embolic base membranous. Conductor spoon-shaped. Female (paratype, MHNG, Sys-05): Total length 10.58. Prosoma 4.82 long, 3.40 wide. Opisthosoma 5.76 long, 3.12 wide. Leg formula: 4123; I 9.48 (2.88, 1.38, 2.60, 1.70, 0.92); II 8.78 (2.66, 1.28, 2.32, 1.64, 0.88); III 8.32 (2.90, 0.88, 1.82, 1.94, 0.78); IV 12.70 (3.60, 1.32, 2.86, 3.32, 1.60). Spination. Femora: I d 1 - 1 - 1, p 1 - 1 -0, r 0-1 - 1; II d 1 - 1 - 1, p 0-1 - 1, r 0-1 - 2; III d 1 - 1 - 1, p 1 - 0-1, r 1-2; IV d 1 - 1 - 1, p 1 - 0-1, r0- 0-1. Tibiae: I v 2 - 2 -0; II v 0-2 -0; III p 1 - 1 -0, r 0- 1 - 1, v 2 - 2 - 2; IV p 1 - 1 -0, r 1 - 1 -0, v 2 - 2 - 2. Metatarsi: II v 1 - 1 -0; III p 1 - 1 - 1, r 1 - 1 - 1, v 2 - 0-2; IV p 1 - 1-2, r 1 - 1-2, v 2 - 2 - 2. Eye size and interdistances: AME 0.14, ALE 0.12, PME 0.10, PLE 0.10; AME–AME 0.16, AME–ALE 0.14, PME–PME 0.22, PME–PLE 0.16, ALE–PLE 0.12; MOA 0.26 long, anterior width 0.38, posterior width 0.36. Genitalia (Figs 30–31). Epigynal atrium deep, with pair of median hoods probably functioning as guiding ridges. Copulatory orifices situated on posterior margin of epigynal atrium. Anterior bursae sclerotized, elongated, widest posteriorly, distinctly narrowed anteriorly. Spermathecae elongate-oval. Fertilization ducts acuminated. Natural history. Types of S. insolita sp. nov. were collected from deciduous forests. The females built retreats beneath bark of a large rotten pine tree. Distribution. Northeastern Thailand (Fig. 43).Published as part of Dankittipakul, Pakawin & Singtripop, Tippawan, 2011, Seven new species of Systaria Simon, 1897 from Southeast Asia (Araneae, Clubionidae, Systariinae), pp. 16-32 in Zootaxa 2905 on pages 26-28, DOI: 10.5281/zenodo.20710