Modulation of Negative Work Output from a Steering Muscle of the Blowfly Calliphora Vicina

Of the 17 muscles responsible for flight control in flies, only the first basalar muscle (b1) is known to fire an action potential each and every wing beat at a precise phase of the wing-beat period. The phase of action potentials in the b1 is shifted during turns, implicating the b1 in the control of aerodynamic yaw torque. We used the work loop technique to quantify the effects of phase modulation on the mechanical output of the b1 of the blowfly Calliphora vicina. During cyclic length oscillations at 10 and 50 Hz, the magnitude of positive work output by the b1 was similar to that measured previously from other insect muscles. However, when tested at wing-beat frequency (150 Hz), the net work performed in each cycle was negative. The twitch kinetics of the b1 suggest that negative work output reflects intrinsic specializations of the b1 muscle. Our results suggest that, in addition to a possible role as a passive elastic element, the phase-sensitivity of its mechanical properties may endow the b1 with the capacity to modulate wing-beat kinematics during turning maneuvers

Cosmological Parameters

The discussion of cosmological parameters used to be a source of embarrassment to cosmologists. Today, measurements of the cosmological parameters are leading the way into the era of precision cosmology. The CMB temperature is measured to four significant figures, T_0=2.7277+/-0.002 K; the Hubble constant is now determined with a reliable error estimate, H_0=(65+/-5) km sec^-1 Mpc^-1; the mass density of baryons is precisely determined by big-bang nucleosynthesis Omega_B = (0.019+/-0.001) h^-2; and the age of the Universe inferred from the ages of the oldest stars is 14+/-1.5 Gyr, which is consistent the expansion age. Further, we have the first full accounting of matter and energy in the Universe, complete with a self consistency check. Expressed as a fraction of the critical density it goes like this: neutrinos, between 0.3% and 15%; stars, between 0.3% and 0.6%; baryons (total), 5+/-0.5%; matter (total),40% +/- 10%; smooth, dark energy, 80% +/- 20%; totaling to the critical density (within the errors).Comment: 27 pages LaTeX with 8 eps figures. To be published in The Proceedings of Particle Physics and the Universe (Cosmo-98), edited by David O. Caldwell (AIP, Woodbury, NY

Toward a Transnational History of World War I

Michael S. Neiberg delivered the keynote address at the 19th Military History Colloquium, held at the University of Waterloo, Waterloo, ON, 1–3 May 2008. This is the text of his address