13 research outputs found

    Shearia teletskaya Mikhaljova 2000

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    <i>Shearia teletskaya</i> Mikhaljova 2000 <p> <i>Shearia teletskaya</i> Mikhaljova 2000: 167 –168, 167: figs 39–40.</p> <p> <i>Shearia teletskaya</i> — Mikhaljova & Golovatch 2001: 111; Mikhaljova 2004: 160 –161, 160: figs 375–376, 112: map 15.</p> <p> <i>Material examined</i> (all from Russia, Siberia, Republic of Altai, Ulagan District, Altai State Nature Reserve, collected by P.S. Nefediev and J.S. Nefedieva): 1 male, 4 juveniles (CN), SE of Teletskoe Lake, environs of cordon Chiri, upper reaches of Bayas River, sparse growth of <i>Pinus sibirica</i>, 1,900 m a.s.l., 14 August 2005; 1 male (CN), 1 male (IBSS), SE of Teletskoe Lake, environs of cordon Chiri, watershed of Kyga and Bayas rivers, 22 numbered sites at different altitudes (= Kyga Profile), site 16, subalpine (= subgoltsy) sparse growth of <i>Pinus sibirica</i>, 2,060 m a.s.l., 15 August 2005; 1 male, 8 juveniles (CN), SE of Teletskoe Lake, environs of cordon Chiri, watershed of Kyga and Bayas rivers, 22 numbered sites at different altitudes (= Kyga Profile), site 12, sparse growth of <i>Pinus sibirica</i> and <i>Abies sibirica</i>, 1,900 m a.s.l., 16 August 2005.</p> <p> <i>Distribution</i>: Russia: Siberia (Republic of Altai).</p>Published as part of <i>Mikhaljova, Elena V., Nefediev, Pavel S. & Nefedieva, Julia S., 2008, A new species and new records of millipedes of the family Diplomaragnidae (Diplopoda, Chordeumatida) from Altai, pp. 49-56 in Zootaxa 1931</i> on page 54, DOI: <a href="http://zenodo.org/record/184844">10.5281/zenodo.184844</a&gt

    Sibiriulus multinicus Mikhaljova 2001

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    Sibiriulus multinicus Mikhaljova, 2001 Figs 8–18 Sibiriulus multinicus Mikhaljova 2001 in Mikhaljova & Golovatch 2001: 106 –107, 106: figs 4– 11. Sibiriulus dentiger - Nefediev 2001: 84. Sibiriulus multinicus - Mikhaljova 2004: 70 –72, 71: figs 100 –113, 74: map 9; Mikhaljova & Nefediev 2003: 85, 85: figs 1–6. Material examined: 1 male, 1 juvenile, 1 fragment (TAI) from Russia, Siberia, Republic of Altai, Turochak District, Altaisky State Nature Reserve, near cordon Baigazan, 409 m a.s.l., Padus avium, collected 3 September 2005 by M.B. Sakhnevitch. Material re-examined: 1 male paratype (IBSS) from Russia, Siberia, Altai Mts, Katunsky Mt. Range, Katunsky Nature Reserve, near Multa, Sredneye Multinskoye Lake, ca. 30 air-km SE of Ust-Koksa, 1,600– 2,200 m a.s.l., collected 30 July– 5 August 1994 by S.I. Golovatch and A.B. Ryvkin; 1 male (IBSS) from Russia, Siberia, Altai Province, Smolenskoe District, near Smolenskoe village, bank of Poperechka River, Salix, collected 27 August 2000 by P.S. Nefediev. Distribution. Russia: Siberia (Republic of Altai, southeastern part of Altai Province). Remarks. The species was originally described from the Altai Mountains; Katunsky Nature Reserve is the type locality (Mikhaljova & Golovatch 2001). The species shows variation in gonopod configuration (Mikhaljova & Nefediev 2003). The fresh material from the Altaisky Reserve adds information concerning the variability of this species. The distal part of the gonopod mesomere is variable in shape. It is prolonged to a different degree, or without prolongation, with an apical fovea and a minute knob, with an apical fovea and without a knob but with an apical hook in specimens from the Altaisky State Nature Reserve (Figs 8–9), Aigulaksky Mt. Range, Republic of Altai (Figs 10–11) and the Smolenskoe District (Figs 12–14), in contrast to a relatively simple apex of the mesomere in a specimen from the Katunsky Nature Reserve (Figs 15, 16). A process of the gonopod opisthomere is long, styletiform; it is placed approximately in the middle portion of the inner side of the opisthomere in specimen from Katunsky Nature Reserve (Fig. 15), in contrast to the short process of the gonopod opisthomere which is placed apically or subapically in specimens from Altaisky State Nature Reserve (Fig. 17) and Smolenskoe District (Fig. 18). Configuration of the gonopod promere is practically invariable. Genus Sibiriulus contains three species known from Siberia.Published as part of Mikhaljova, Elena V., Nefediev, Pavel S. & Nefedieva, Julia S., 2007, New data on millipedes of the family Julidae (Diplopoda, Julida) from Altai, Siberia, pp. 57-63 in Zootaxa 1541 on pages 60-62, DOI: 10.5281/zenodo.17784

    The millipede genus Sibiriulus GuliÄŤka, 1963 revisited, with the description of a new species from the Altais, southwestern Siberia, Russia (Diplopoda: Julida Julidae)

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    Nefediev, Pavel S., Nefedieva, Julia S., Krugova, Tatiana M. (2020): The millipede genus Sibiriulus GuliÄŤka, 1963 revisited, with the description of a new species from the Altais, southwestern Siberia, Russia (Diplopoda: Julida Julidae). Zootaxa 4869 (3): 421-429, DOI: https://doi.org/10.11646/zootaxa.4869.3.

    Altajosoma deplanatum Stuxberg 1876

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    <i>Altajosoma deplanatum</i> (Stuxberg 1876) <p> <i>Craspedosoma deplanatum</i> Stuxberg 1876: 36, 39: figs.</p> <p> <i>Craspedosoma deplanatum</i> — Stuxberg 1876a: 317; Lokšina & Golovatch 1979: 383. <i>Altajosoma pinetorum</i> Guliċka 1972: 37 –38, 37: fig. 1.</p> <p> <i>Altajosoma pinetorum</i> — Lokšina & Golovatch 1979: 382; Shelley <i>et al</i>. 2000: 62.</p> <p> <i>Diplomaragna deplanata</i> — Shear 1990: 19 –21, 20: figs 29–33; Mikhaljova 1993: 22.</p> <p> <i>Diplomaragna pinetorum</i> — Shear 1990: 38; Mikhaljova 1993: 25.</p> <p> <i>Altajosoma deplanatum</i> — Mikhaljova 2000: 160: figs 16–19; 2004: 170–171, 171: figs 389–393, 162: map 21;; Mikhaljova & Golovatch 2001:108; Nefediev 2002: 30; 2002a: 35; Mikhaljova & Nefediev 2003: 86.</p> <p> <i>Material examined</i> (all from Russia, Siberia, Republic of Altai, collected by P.S. Nefediev and J.S. Nefedieva): 1 male, 6 juveniles (CN), Ulagan District, Altai State Nature Reserve, SE of Teletskoe Lake, environs of cordon Chiri, watershed of Kyga and Bayas rivers, 22 numbered sites at different altitudes (= Kyga Profile), site 14, subalpine sparse growth of <i>Pinus sibirica</i>, 1,935 m a.s.l., 14 August 2005; 1 male (CN), Ulagan District, Altai State Nature Reserve, SE of Teletskoe Lake, environs of cordon Chiri, watershed of Kyga and Bayas rivers, 22 numbered sites at different altitudes (= Kyga Profile), site 6, forest (<i>Pinus sylvestris, Abies sibirica, Populus tremula</i>), 860 m a.s.l., 18 August 2005; 1 male (CN) Ulagan District, 15 km NNW of Aktash, Taldukel’ Lake, forest (<i>Larix sibirica</i>), 17–18 July 2006; 1 male (CN), 1 male (IBSS), Ulagan District, 25 km NNE of Aktash, Ulagan Pass, sparse growth of <i>Pinus sibirica</i> and <i>Larix sibirica</i> with <i>Betula rotundifolia</i>, 2,080 m a.s.l., 31 July 2006; 1 male (CN), Ulagan District, 18 km NNW of Aktash, NW of Sorulukel’ Lake, top of the hill, forest (<i>Pinus sibirica, Larix sibirica</i>), 10 August 2007; 1 male (CN), Ulagan District, 18 km NNW of Aktash, NW of Sorulukel’ Lake, forest (<i>Pinus sibirica, Larix sibirica</i>), 13 August 2007; 2 males (CN), Turochak District, Altai State Nature Reserve, N of Teletskoe Lake, Yailyu, broad gully (<i>Betula pendula, Pinus sylvestris</i>), 19 August 2005.</p> <p> <i>Distribution</i>: Russia: Siberia (Kemerovo Area, between Mariinsk in Kemerovo Area and Achinsk in Krasnoyarsk Province, Republic of Khakassia, Novosibirsk and Tomsk areas, Republic of Altai).</p> <p> <i>Remarks</i>: At present, environs of Aktash are the southernmost records of this species.</p>Published as part of <i>Mikhaljova, Elena V., Nefediev, Pavel S. & Nefedieva, Julia S., 2008, A new species and new records of millipedes of the family Diplomaragnidae (Diplopoda, Chordeumatida) from Altai, pp. 49-56 in Zootaxa 1931</i> on pages 51-52, DOI: <a href="http://zenodo.org/record/184844">10.5281/zenodo.184844</a&gt

    Shearia calycina Mikhaljova 2000

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    <i>Shearia calycina</i> Mikhaljova 2000 <p> <i>Shearia calycina</i> Mikhaljova 2000: 166 –167, 167: figs 37–38.</p> <p> <i>Shearia calycina</i> — Mikhaljva & Golovatch 2001: 110; Mikhaljova 2004: 163 –164, 163: figs 379–380, 164: map 22.</p> <p> <i>Material examined</i> (all from Russia, Siberia, Republic of Altai, Kosh-Agach District, collected by P.S. Nefediev and J.S. Nefedieva): 1 male, 3 females, 7 juveniles (CN), 22 km SSE of Belyi Bom, near Achik, sparse growth of <i>Pinus sibirica</i> with <i>Larix sibirica, Betula rotundifolia, Lonicera altaica</i>, 2,000–2,200 m a.s.l., 22 July 2006; 1 male, 1 female, 2 juveniles (CN), 1 male (IBSS), 22 km SSE of Belyi Bom, near Achik, S riverside slope of Shavla River, subalpine meadow, under <i>Pentaphylloides fruticosa</i>, 22 July 2006.</p> <p> <i>Distribution</i>: Russia: Siberia (Republic of Altai).</p> <p> <i>Remarks</i>: Originally described from 2,800 m a.s.l., northern extremity of Yuzhno-Chuiskii Mt. Range, watershed of Sebystei and Kokuzek rivers, South Altai Mts (Mikhaljova 2000), this species has since never been rediscovered. Therefore, the above samples are the first records of this species at 2,000–2,200 m a.s.l. in the northwest extremity of Severo-Chuiskii Mt. Range.</p> <p> <b> <i>Shearia densecava</i> (Guli</b> ě <b>ka 1972)</b></p> <p> <i>Altajosoma densecavum</i> Guliċka 1972: 38, 37: fig 2.</p> <p> <i>Altajosoma densecavum</i> — Lokšina & Golovatch 1979: 382.</p> <p> <i>Diplomaragna densecava</i> — Shear 1990: 38 –39; Mikhaljova 1993: 22.</p> <p> <i>Shearia densecava</i> — Mikhaljova 2000: 165 –166, 166: figs 33–36; Mikhaljova & Golovatch 2001: 110; Mikhaljova & Nefediev 2003: 86.</p> <p> <i>Material examined</i> (all from Russia, Siberia, Republic of Altai): 1 male (CN), Shebalino District, Kumalyr, 85° 34' – 51° 10', <i>Larix sibirica</i> park-like forest, 1,050 m a.s.l., collected 14 August 2005 by A.S. Babenko; (all below from Ulagan District, collected by P.S. Nefediev and J.S. Nefedieva) 1 male (CN), 1 male (IBSS), 15 km NNW of Aktash, Taldukel’ Lake, forest (<i>Larix sibirica</i>), 17–18 July 2006; 2 males, 1 female, 10 juveniles (CN), near ford on the left bank of Kalbakaya River, 89° 05' – 50° 21', sparse growth of <i>Larix sibirica</i> with <i>Pinus sibirica</i> and <i>Betula rotundifolia</i> on the hill slope, 25 July 2006; 2 males (CN), 25 km NNE of Aktash, Ulagan Pass, sparse growth of <i>Pinus sibirica</i> and <i>Larix sibirica</i> with <i>Betula rotundifolia</i>, 2,080 m a.s.l., 31 July 2006; 2 males (CN), Aktash, top of Sardyma Mt., forest (<i>Pinus sibirica, Larix sibirica</i>), 5 August 2006; 3 males (CN), 18 km NNW of Aktash, NW of Sorulukel’ Lake, top of the hill, forest (<i>Pinus sibirica, Larix sibirica</i>), 10 August 2007; 1 male, 27 juveniles (CN), 18 km NNW of Aktash, SW of Sorulukel’ Lake, bottom of N slope of Aigulakskii Mt. Range, forest (<i>Pinus sibirica, Larix sibirica</i> with <i>Betula rotundifolia</i>), 11 August 2007; 1 male, 1 female (CN), 18 km NNW of Aktash, SW of Sorulukel’ Lake, forest (<i>Pinus sibirica, Larix sibirica</i>), under mushroom cap, 12 August 2007; 1 male (CN), 18 km NNW of Aktash, NW of Sorulukel’ Lake, forest (<i>Pinus sibirica, Larix sibirica</i>), 13 August 2007.</p> <p> <i>Distribution</i>: Russia: Siberia (Republic of Altai).</p> <p> <i>Remark</i> s: At present Kumalyr is the northernmost record of this species, and Kalbakaya River is the easternmost one.</p>Published as part of <i>Mikhaljova, Elena V., Nefediev, Pavel S. & Nefedieva, Julia S., 2008, A new species and new records of millipedes of the family Diplomaragnidae (Diplopoda, Chordeumatida) from Altai, pp. 49-56 in Zootaxa 1931</i> on pages 53-54, DOI: <a href="http://zenodo.org/record/184844">10.5281/zenodo.184844</a&gt

    Megaphyllum sjaelandicum Meinert 1868

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    Megaphyllum sjaelandicum (Meinert, 1868) Fig. 19 Material examined: 1 male, 1 female (IBSS) from Russia, Siberia, Republic of Altai, Shebalino District, Kumalyr, 85 ° 34 ' – 51 ° 10 ', Larix sibirica park-like forest, 1050 m a.s.l., collected 14 August 2005 by A.S. Babenko; 1 male, no. 50 (PSU) from Russia, Urals, Perm Province, Kishertsky District, "Predural'e" protected territory, Kamen' Mezhevoy, southern slope, steppe part, limestone, collected 31 May 1996 by N.S. Mazura; 2 males, 1 female, no. 102 (IBSS), 1 male, no. 102 (PSU) from Russia, Perm Province, Kachka village, new Betula forest, pitfall traps, collected 2–13 June 2001 by S.L. Esyunin. Remarks. This species is distributed in Central and Eastern Europe. Hitherto the easternmost known records are from the Ural Region (Komlev & Esyunin 1996). This is the first record of the species in Siberia. This species appears to be variable. The mesal side of the gonopod promere in a male from Altai has a small projection distally (Fig. 19). The gonopod promere is devoid of such a projection in males from European countries and the Ural Region. In addition, the gonopod flagella are somewhat longer, and the gonopod mesomere processes are somewhat more obtuse in a male from Altai in comparison with ones in males from the Urals. A still undescribed species referred to as Megaphyllum aff. sjaelandicum was recorded in the West (= Rudnyi) Altais, East Kazakhstan (Golovatch 1992). It seems plausible that it too belongs to M. sjaelandicum, taking into account the variability of latter species.Published as part of Mikhaljova, Elena V., Nefediev, Pavel S. & Nefedieva, Julia S., 2007, New data on millipedes of the family Julidae (Diplopoda, Julida) from Altai, Siberia, pp. 57-63 in Zootaxa 1541 on pages 62-63, DOI: 10.5281/zenodo.17784

    Altajosoma corniferum Mikhaljova, Nefediev & Nefedieva, 2008, sp.n.

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    <i>Altajosoma corniferum</i> sp.n. <p>Figs 1–3</p> <p> <i>Material examined</i>: <i>Holotype</i>: 1 male (IBSS), Russia, Republic of Altai, Ongudai District, Seminskii Pass, forest (<i>Pinus sibirica</i>), 1,800 m a.s.l., collected 15 August 2005 by A.S. Babenko.</p> <p> <i>Diagnosis</i>: The species differs from congeners mainly by the large, horn-shaped lateral sheath processes of the posterior gonopod colpocoxites.</p> <p> <i>Description</i>: Male. Length 17 mm, width 1.6 mm with paraterga. Coloration in alcohol light brown. Legs yellowish, with marbled brownish distal parts. Eyes black. Antennae broken.</p> <p>Body with 32 segments. Anterior part of head sparsely setose, vertigial suture hardly visible. Eye patches subtriangular, with 23–27 ocelli. Collum semicircular, its macrochaetae broken. Somite 2 narrower than both head (with genae) and somite 3. Paraterga beginning on somite 2, well developed on somites 4–27, reduced on somite 28, onward missing. Metazonital macrochaetae in a transverse row on somites 29–31, like an extended triangle on preceding somites. Macrochaetae pointed, but not very sharply so. Anterolateral macrochaetae shortest, caudolateral and medial ones subequal in length. Axial suture well developed.</p> <p>Legs long and slender. Leg pairs 1 and 2 typically reduced in size, with tarsal brushes as usual; each claw with long setiform outgrowth (=filament) ventrally at base, and two small additional claws dorsally. Other pregonopodal legs gradually enlarged toward gonopods. Leg pairs 3–7 with group of funnel-shaped tarsal papillae apically near claw. Claws 3 and 4 each with long setiform outgrowth ventrally and two small additional claws dorsally at base. Claws 5–7 each with a long setiform outgrowth at base ventrally only. Postgonopodal legs (including leg pairs 10 and 11) with funnel-shaped tarsal papillae apically near claw. Tarsal papillae gradually missing toward hind part of body. Claws of postgonopodal legs (including claws of leg pairs 10 and 11) each with a long setiform outgrowth ventrally and two small additional claws dorsally at base; long setiform outgrowth and additional claws gradually missing toward posteriormost legs (the additional claws missing former). Claws of posteriormost legs without both a long setiform outgrowth ventrally and two small additional claws dorsally. Legs pairs 10 and 11 with coxal glands. Coxa 11 (Fig. 1) with ventral process covered with sparse setae; coxal gland opening on an anterior wall of coxa.</p> <p>Anterior gonopod telopodites 2-segmented, subflagelliform (Fig. 2); first telopodite segment and distal part of coxosternum attached to the adjacent mesal portion of posterior gonopod by thin membranous film. Distal part of telopodite hidden inside narrow sheath groove. Posterior gonopod colpocoxites fused basally, curved caudad. Sheath groove stretching to anterior surface of colpocoxite. Lateral sheath process of colpocoxite (lp) large, as horn curved basally, with small mesal protuberance subbasally; mesal surface of lateral sheath process shagreen. Mesal sheath process of colpocoxite absent. Angiocoxite with a globule in posterior view. Posterior angiocoxal process (pp) large, curved forward; its apex pointed. Anterior angiocoxal process (ap) digitiform, with a blunt apex (Fig. 3). Posterior gonopod telopodites 2-segmented, setose, with a long femur.</p> <p>Female. Unknown.</p> <p> <i>Etymology</i>: The specific epithet refers to the horn-shaped lateral sheath processes of the posterior gonopod colpocoxites.</p>Published as part of <i>Mikhaljova, Elena V., Nefediev, Pavel S. & Nefedieva, Julia S., 2008, A new species and new records of millipedes of the family Diplomaragnidae (Diplopoda, Chordeumatida) from Altai, pp. 49-56 in Zootaxa 1931</i> on pages 49-51, DOI: <a href="http://zenodo.org/record/184844">10.5281/zenodo.184844</a&gt

    Altajosoma bakurovi subsp. bakurovi Shear 1990

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    <i>Altajosoma bakurovi bakurovi</i> (Shear 1990) <p> <i>Diplomaragna bakurovi</i> Shear 1990: 22, 23: figs 38–41.</p> <p> <i>Diplomaragna bakurovi</i> — Mikhaljova 1993: 18.</p> <p> <i>Altajosoma bakurovi</i> — Mikhaljova 2000: 161: fig. 20; 2004: 178–179, 178: figs 419–422, 116: map 16; Mikhaljova & Golovatch 2001:108; Mikhaljova & Nefediev 2003: 86; Nefediev 2002: 30.</p> <p> <i>Material examined</i> (all from Russia, Siberia, Republic of Altai, collected by P.S. Nefediev and J.S. Nefedieva): 1 male, 22 juveniles (CN), Turochak District, NW of Teletskoe Lake, 2 km E of Iogach, forest (<i>Pinus sibirica, Abies sibirica, Betula pendula</i> with fern), 434 m a.s.l., 14–17 July 2004; 1 male, 1 juvenile (CN), Turochak District, Altai State Nature Reserve, environs of Teletskoe Lake (central part), near cordon Kokshi, forest (<i>Pinus sylvestris, Betula pendula</i>), 18 August 2005; 2 males (CN), Turochak District, Altai State Nature Reserve, N of Teletskoe Lake, Yailyu, broad gully (<i>Betula pendula, Pinus sylvestris</i>), 19 August 2005; 1 male (CN), 1 male (IBSS) Ulagan District, Altai State Nature Reserve, SE of Teletskoe Lake, near cordon Chiri, mouth of Chiri River, forest (<i>Betula pendula</i>), 11–12 August 2005; 1 male (CN), Ulagan District, Altai State Nature Reserve, 5 km SE of cordon Chiri, Bayas River, forest (<i>Salix viminalis</i>), 12 August 2005.</p> <p> <i>Distribution</i>: Russia: Siberia (Novosibirsk, Kemerovo and Tomsk areas, Krasnoyarsk Province, Republic of Altai).</p>Published as part of <i>Mikhaljova, Elena V., Nefediev, Pavel S. & Nefedieva, Julia S., 2008, A new species and new records of millipedes of the family Diplomaragnidae (Diplopoda, Chordeumatida) from Altai, pp. 49-56 in Zootaxa 1931</i> on page 51, DOI: <a href="http://zenodo.org/record/184844">10.5281/zenodo.184844</a&gt

    Shearia oiskaya Mikhaljova 2000

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    <i>Shearia oiskaya</i> Mikhaljova 2000 <p> <i>Shearia oiskaya</i> Mikhaljova 2000: 165: figs 31–32.</p> <p> <i>Shearia oiskaya</i> — Mikhaljova & Golovatch 2001: 110; Mikhaljova 2004: 167 –169, 168: figs 387–388, 164: map 22.</p> <p> <i>Material examined</i>: 7 males, 2 females, 22 juveniles (CN), 1 male (IBSS) Russia, Republic of Altai, Ulagan District, Altai State Nature Reserve, 9 km S of Dzhulukul’ Lake, near Pak-Yyash Lakes, <i>Betula rotundifolia</i>, 2,200 m a.s.l., 27 July 2006, collected by P.S. Nefediev and J.S. Nefedieva.</p> <p> <i>Distribution</i>: Russia: Siberia (Krasnoyarsk Province, Republic of Altai).</p> <p> <i>Remarks</i>: The species has hitherto been known from West Sayan Mts., Krasnoyarsk Province, its <i>terra typica</i> (Mikhaljova 2000). Therefore, the above material is the first record of this species in the Republic of Altai.</p>Published as part of <i>Mikhaljova, Elena V., Nefediev, Pavel S. & Nefedieva, Julia S., 2008, A new species and new records of millipedes of the family Diplomaragnidae (Diplopoda, Chordeumatida) from Altai, pp. 49-56 in Zootaxa 1931</i> on page 54, DOI: <a href="http://zenodo.org/record/184844">10.5281/zenodo.184844</a&gt

    Genus Leptoiulus Verhoeff, 1894 new to the fauna of the Asian part of Russia, with description of a new species from the Altai and its comparison with the European Leptoiulus trilineatus (C. L. Koch, 1847) (Diplopoda, Julida, Julidae)

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    Mikhaljova, Elena V., Nefediev, Pavel S., Nefedieva, Julia S., Dyachkov, Yuri V. (2015): Genus Leptoiulus Verhoeff, 1894 new to the fauna of the Asian part of Russia, with description of a new species from the Altai and its comparison with the European Leptoiulus trilineatus (C. L. Koch, 1847) (Diplopoda, Julida, Julidae). Zootaxa 3974 (2): 267-276, DOI: 10.11646/zootaxa.3974.2.1
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