The first checklist of the tribe Olethreutini (Lepidoptera: Tortricidae: Olethreutinae) of Thailand

Muadsub, Sopita, Pinkaew, Nantasak (2023): The first checklist of the tribe Olethreutini (Lepidoptera: Tortricidae: Olethreutinae) of Thailand. Zootaxa 5349 (1): 1-65, DOI: 10.11646/zootaxa.5349.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5349.1.

Hiroshiinoueana curiosissima Pinkaew & Muadsub 2021, comb. nov.

Hiroshiinoueana curiosissima (Razowski, 2009) comb. nov. (Figs. 1–4, 9–14, 18–19) Atsusina curiosissima Razowski, 2009. Polish Journal of Entomology 78: 15–32. Specimens examined: 4♂, 3♀. THAILAND. Uthai Thani Prov., Huai Kha Khaeng W.S., 15 o 33ʹ11ʺN, 99 o 17ʹ45ʺE, alt. 250 m; 12 Oct 2020; np13607 (♂, genitalia slide NP3970); np13537 (♀); 17 Dec 2020; np13670 (♀, genitalia slide NP3971); np13671 (♀, genitalia slide NP3972); np13676 (♂), 15 o 36ʹ32ʺN, 99 o 18ʹ58ʺE, alt. 195 m; 14 Oct 2020; np13536 (♂, genitalia slide NP3968); np13606 (♂, genitalia slide NP3969); Pinkaew et al. leg.; KKIC. Diagnosis. Hiroshiinoueana curiosissima differs from all other species in the genus by the distinct unequal valvae, conspicuous small and short of right cucullus, absence of a tuft of long bristles on each valva, and presence of two clusters of moderately long curve spines below the left cucullus apex. Redescription. Head (Figs. 2, 4): Lower frons and upper frons light brown, vertex light brown mixed with brown, mostly scales iridescent when angled to light; labial palpi porrect, first segment yellowish white, second segment dilated to widened apex, yellowish white at base, slightly darker to light brown at apex, with three dark brown spots dorsobasally, medially, and apicoventrally respectively, third segment light brown, paler at apex; antenna brown, reaching middle of forewing. Thorax: Pronotal collar and tegulae brown, with greyish white tips, some scales iridescent when angled to light, mesonotum brown with yellowish tips, median with transverse band of light brown scales with white tips, posterior crest dark brown with yellowish tips. Forewing broad subrectangular, wingspan 12.1–12.6 mm in males (n = 4) (Fig. 1), 12.0–13.2 mm in females (n = 3) (Fig. 3), costa evenly curve; termen round; costal strigulae well developed, pinkish separated by blackish streaks; ground color silvery greyish brown, basal 1/3 with indistinct, irregular patch, brown to dark brown, broken by ground color and edged with pinkish brown scales, medially with large, indistinct irregular transverse patch, extending slightly oblique from costa to dorsum, brown mixed with dark brown, broken by ground color and edged with pinkish brown scales, inner margin extending from near middle of costa slightly oblique to middle of dorsum, outer margin extending from apical 1/4 of costa slightly oblique to tornus, near middle of patch with a small whitish grey X-shaped mark, with distinct oblique dark brown strip, edged with pinkish brown scales, extending oblique from apical 1/4 of costa to middle of termen, wing apex with small dark brown spot; under side greyish brown with white spots on costa. Hindwing (Figs. 1, 3) subrectangular, brown slightly paler to wing base; underside greyish brown. Abdomen: Tergum II with two small pits on anterolateral corners (Fig. 18), tergum VIII with enlarged posterior margin, rounded lobe (Fig. 19). Male genitalia (Figs.9–12).Tegumen subtriangular, rather narrow; uncus tubular lobe, rather short, slightly dilated apex, sparsely setose basally, apex with dense setae (Fig. 10); socii and gnathos indistinct; vinculum rather narrow; juxta subtriangular; caulis moderately long, wide cone-shaped, anellus surrounding apical 2/3 of phallus; phallus capsule-shaped, moderately long, widened medially, vesica with dense cornuti sockets (Fig. 12); valvae asymmetry; left valva larger than right, left valva elongate ovate, with distinct longitudinal fold from base to near apex, beneath with a large concave, dorsal margin fold downward over ventral margin, basal half with a large suboval-shaped patch of dense short spines mixed with moderately long spines, pointing inward, covered mostly area of sacculus except ventrobasally, inner margin of lobe round protruding inward, sacculus densely setose basally, cucullus elongate dome-shaped, rounded apex, moderately dense setae, with a group of eight moderately long, strong and curve spines beneath apex, beneath with a smaller group of two rather short, slightly curve spines (Fig. 11); right valva quite smaller than left, subtriangular, basal 2/3 covered with subcircular lobe, inner margin combined with basal excavation edge protruding inwardly, distinctly folded apicolaterally, lobe covered with dense of short to moderately long stout spines, mostly pointing inward, sacculus with moderately dense setae ventrobasally, cucullus smaller than sacculus, elongate subtriangular lobe, stout apex, moderately dense setae. Female genitalia (Figs. 13–14). Papillae anales densely setose; tergum VIII smooth, with sparse setae on subtriangular extension; sternum VII weakly sclerotized, with dense scale sockets, especially along very slightly concave posterior margin; sterigma strongly sclerotized, forming large long and curve tube with smooth surface, anterior half behind sternum VII with rather parallel margins, posterior half protruding upward over posterior margin of sternum VII, narrowed to posterior end (Fig. 13); colliculum sclerotized, long about half length of ductus bursae; ductus bursae rather narrow posterior half, anterior half widened to corpus bursae; ductus seminalis arising from anterior 1/3 of ductus bursae; corpus bursae ovate, granulate throughout, signum (Fig. 14) a scobinate patch, with distinct two large, tubular teeth pointing ventrally and two larger, rounded teeth pointing laterally. Distribution. Vietnam and Thailand. New record to Thailand. Remarks. Hiroshiinoueana curiosissima (Razowski, 2009) is superficailly similar to Gatesclarkeana idia Diakonoff, 1973 and Ukamenia thailandica Pinkaew & Muadsub, 2020 especially wing pattern and coloration of forewing. It can be distinguished by rather rounded termen and whitish grey X-shaped mark in forewing, hindwing without modified anal margin, and hind tibia without tuft of hair pencils. In contrast, G. idia and U. thailandica has straight forewing termen, middle of forewing with a rounded white spot in G. idia but absent in U. thailandica, anal marginal fold in hindwing and distinct hair pencils in hind tibia are also present. Although forewing pattern of H. curiosissima looks similar to G. idia and U. thailandica but it can be distinguished from the two latter by a large spiny lobe at base of valva (shared character of the genus Hiroshiinoueana) but it is small in G. idia and not present in U. thailandica. Furthermore, H. curiosissima lack spinules on the outer surface of sterigma in female genitalia but it is present in G. idia.Published as part of Pinkaew, Nantasak & Muadsub, Sopita, 2021, First record of the genus Hiroshiinoueana Kawabe, 1978 (Lepidoptera: Tortricidae: Olethreutinae), with a new record and a new species from Thailand, pp. 442-450 in Zootaxa 5023 (3) on pages 443-444, DOI: 10.11646/zootaxa.5023.3.8, http://zenodo.org/record/522633

Hiroshiinoueana partita Pinkaew & Muadsub 2021, sp. nov.

Hiroshiinoueana partita sp. nov. urn:lsid:zoobank.org:pub: 181D382E-3C33-4953-8690-1791E210A198 (Figs. 5–8, 15–17, 20–21) Type specimen. Holotype: ♂, THAILAND. Nakhon Nayok Prov., Khao Yai N.P., 14 o 23ʹ56ʺN, 101 o 22ʹ16ʺE, alt. 786 m; 10 Sep 2010; Pinkaew et al. leg.; np4798 (genitalia slide NP1514); KKIC. Paratype: 1♂, THAILAND. Nakhon Ratchasima Prov., Khao Yai N.P., 14 o 25ʹ55ʺN, 101 o 26ʹ37ʺE, alt. 791 m, 11 Nov 2009; Pinkaew et al. leg.; np3190 (♂, genitalia slide NP1258); KKIC. Diagnosis. Hiroshiinoueana partita sp. nov. is similar to H. wuzhishanica. It can be distinguished from the latter species by upperside of hind wing is light brown, and dark greyish brown hair pencil on the inner side of hind tibia, whereas H. wuzhishanica has pale greyish brown on the upperside of hindwing, and black hair pencils on the inner side of hind tibia. In male genitalia, H. partita sp. nov. have subtriangular pointing downward uncus, spines on the large lobe of sacculus are separated into two groups, neck is distinct, and smaller size of a tuft of long bristles is located ventromedially of sacculus, whereas H. wuzhishanica has small semicircular pointing upward uncus, spines on the large lobe of sacculus are not separated, neck is indistinct in left valva, and larger size of a tuft of long bristles is located beyond ventromedially of sacculus. Description. Head (Fig. 6): Lower frons light brown, upper frons and vertex brown mixed with light brown; labial palpi porrect, first segment light brown, second segment long, dilated to apex, basal 2/3 brown, apical 1/3 light brown, third segment light brown; antenna light brown, reaching middle of forewing. Pronotal collar and tegulae brown with some metallic silvery scales when angled to light; mesonotum light brown mixed with brown, posterior crest mostly loss; hind tibia light brown, basal half of inner side with a tuft of brown hair pencils, originated beyond base of tibia (Fig. 8). Forewing broad rectangular, wingsapn 12.7–13.3 mm in male (Fig. 5) (n = 2); costal margin evenly curve; termen rather round; costal strigulae indistinct, ground color brown with diffuse metallic silvery spots throughout; underside light greyish brown. Hindwing light brown, anal margin with long marginal fold (Fig. 7); underside light greyish brown with thickened, folded anal margin. Abdomen: Tergum II with two small pits on anterolateral corners (Fig. 20), tergum VIII with enlarged posterior margin, slightly concaved medially (Fig. 21). Male genitalia (Figs. 15–17). Tegumen widened basal half, narrower rounded dome-shaped apical half, dorsolateral corners folded, densely setose sockets; uncus small, subtriangular hairy lobe pointing ventrally (Fig. 16); socii moderately long plate-like, densely setose; gnathos moderately sclerotized, narrowed band, arising from midlength of tegumen, extending inwardly then abruptly bent upward; vinculum moderately wide; juxta subtrapezoid; caulis long, apical 2/3 enlarged; anellus wide, sclerotized, surrounding phallus, dorsal reach to apical 3/4 of phallus length; phallus rather short, slightly curve horn-shaped, pointed apex, cornuti absent; valvae symmetry, longitudinal folded, basal excavation with moderately dense, short setae on margin reaching sacculus basally; sacculus with a tuft of long bristles ventromedially, with a large oblique spiny lobe medially, suboval-shaped, protruding inwardly narrowed to phallus, dorsal margin folded, slightly widened to base of lobe and adjacent to base of cucullus, margined with row of dense, short and strong spines pointing oblique ventrally, inner margin combined with basal excavation edge, concaved apicomedially, outer margin round, pendant over ventromedially of sacculus, with aciculate patch along margin lateromedially, with large concave beneath lobe, wrinkled surface, lobe mostly covered with a large, oval-shaped patch of short and strong spines, distinctly split apically from row of spines on dorsal edge, pointing obliquely upward to phallus (Fig. 17); neck distinct, moderately dense setae ventrally; cucullus smaller than sacculus, widened crescent-shaped, densely setose ventral half to apex. Distribution. Thailand Etymology. The specific epithet (partita = separate) refers to separated group of spines on basal large lobe of valvae. Remarks. Wing pattern of Hiroshiinoueana partita sp. nov. in both type specimens are rather rubbed especially ground color and fascia but mottle metallic blue spots throughout the forewing are remained.Published as part of Pinkaew, Nantasak & Muadsub, Sopita, 2021, First record of the genus Hiroshiinoueana Kawabe, 1978 (Lepidoptera: Tortricidae: Olethreutinae), with a new record and a new species from Thailand, pp. 442-450 in Zootaxa 5023 (3) on pages 445-447, DOI: 10.11646/zootaxa.5023.3.8, http://zenodo.org/record/522633

FIGURES 1–2 in Description of Antaeola rhinosa sp. nov. (Lepidoptera: Tortricidae: Olethreutinae) from Thailand

FIGURES 1–2. Wing pattern of Antaeola rhinosa sp. nov. 1. Male (holotype) 2. Female (paratype, np3339). Scale bars = 2 mm.Published as part of Muadsub, Sopita & Pinkaew, Nantasak, 2023, Description of Antaeola rhinosa sp. nov. (Lepidoptera: Tortricidae: Olethreutinae) from Thailand, pp. 133-140 in Zootaxa 5323 (1) on page 134, DOI: 10.11646/zootaxa.5323.1.8, http://zenodo.org/record/820407

Sirindhornia bifida Pinkaew and Muadsub 2014, n. sp.

<i>Sirindhornia bifida</i> Pinkaew and Muadsub, n. sp. <p>Figures 1G–H, 3F, 4D</p> <p> <b>Diagnosis.</b> <i>Sirindhornia bifida</i> is the only species without a central orange mark in the hindwing, and together with <i>Sirindhornia</i> sp. (discussed below), can also be distinguished from all other known species of <i>Srinidhornia</i> by relatively large and confluent black spots in basal half of the forewing, diagonally connecting the base and the orange distal part, by the wide and strongly interconnected black lines in the distal half, and by the black line from middle of the costa straight to below the notch then downcurved parallel to termen. The entirely white labial palpi, an evenly curved costa, and much paler hindwings separate <i>S. bifida</i> from <i>Sirindhornia</i> sp. In the male genitalia the very narrow valva and a distally free, finger-like sacculus and a small narrowly ovate phallus are unique for the genus.</p> <p> <b>Description</b>. Head: As described for genus (Fig. 1G–H), but third segment of labial palpus entirely white.</p> <p> Thorax: As described for genus, but tegulae with central ovate black spot and black scales at base. Forewing length 3.8 mm in male (n = 1) (Fig. 3F); basal white area relatively short, 1/3 length to middle of termen, with relatively large and confluent black spots, some diagonally connecting base and orange distal wing part; distal 2/3 with orange ground color, its inner margin near costa preceded by two conspicuous black spots, medially somewhat projecting inward and from middle of wing edged by wide blackish band vertically to dorsum; strigulae and marks along costa as for genus but black line from middle of costa long, straight then ending in vertical, narrow band along middle of termen; notch with a small white mark; remainder of orange wing portion with wide, sinuate, strongly interconnected black bands and few black spots. Fringe as for genus. Underside as for genus. Hindwing with orange band along termen from apex to M 3, without orange patch near center. Fringe as for genus. Underside without patch.</p> <p>Abdomen: Male genitalia (Fig. 4D) [damaged] with socii bearing dense flattened modified spines on outer surface and 3 huge thorns on process projecting from outer margin, and with sclerotized gnathos arms along base; phallus small, narrowly ovate, more slender at base; valva long, narrow, spatulate, with distally free sacculus forming finger-like, slightly curved, setose process, outer valva surface with patch of long, modified scales in basal half, costal hook short with rounded tip.</p> <p>Holotype: ♂, Thailand, Trat Province, Trat Agroforestry R. St., 12°23'43"N 102°40'32"E, ca 30 m, 21–23 Apr 2013, N. Pinkaew, np 5007, ♂ genitalia slide NP1786. Deposited in KKIC.</p> <p> <b>Etymology.</b> The specific epithet <i>bifida</i> means ‘divided into two parts’ in Latin, referring to the sacculus which is distally separated from the valva.</p> <p> <b>Distribution.</b> Thailand.</p> <p> <b>Remarks.</b> This single male was collected in summer in the agroforestry ecosystem of the Trat Agroforestry Research and Training Station. Despite the highly modified genitalia, we name this species even though its genitalia are somewhat damaged.</p>Published as part of <i>Muadsub, Sopita & Pinkaew, Nantasak, 2014, Sirindhornia Pinkaew and Muadsub (Lepidoptera: Tortricidae), a new enarmoniine genus from Thailand, pp. 53-63 in Zootaxa 3869 (1)</i> on pages 61-62, DOI: 10.11646/zootaxa.3869.1.5, <a href="http://zenodo.org/record/4930835">http://zenodo.org/record/4930835</a&gt

Sirindhornia curvicosta Pinkaew and Muadsub 2014, n. sp.

<i>Sirindhornia curvicosta</i> Pinkaew and Muadsub, n. sp. <p>Figures 1E–F, 3E, 4C</p> <p> <b>Diagnosis.</b> <i>Sirindhornia curvicosta</i> is most similar to <i>S. chaipattana</i>, both with a broad nearly white distal margin to the white and black basal half of the forewing and a black line from before middle of costa downcurved and running towards dorsum, but <i>S. curvicosta</i> can be recognized by the distally strongly curved costa resulting in a narrow apex. In the male genitalia <i>S. curvicosta</i> has a more narrowly triangular uncus and the dorsal anellus process medially parallel-sided rather than distinctly narrowed as in <i>S. chaipattana</i>.</p> <p> <b>Description</b>. Head: Damaged, without scales; labial palpus as for genus (Fig. 1E–F), third segment of labial palpus with black scales in vertical groove on anterior face.</p> <p> Thorax: As for genus, tegulae white with thin black line along inner margin then recurved to center, and short black line on outer margin near base. Forewing length 5.2 mm in male (n = 1) (fig. 3E); costa more curved at wing apex, apex narrow; basal 2/5 white, with numerous, slender, connected black lines concentrated towards base and distally white with few black dots; distal 3/5 with reddish orange ground color, its inner margin near costa preceded by black spots and followed by a parallel, angled silvery band, slightly widening from costa to dorsum; strigulae and marks along costa as for genus, with one oblique, narrow black line from middle of costa downcurved; triangular white mark at notch; remainder of reddish orange wing portion with rather narrow, sinuate, slightly confluent silvery blackish bands and few black spots. Fringe white at notch. Underside as for genus. Hindwing very dark, with narrow orange patch along termen from apex to M 2 and a narrow orange streak in center. Underside with two longitudinal, greyish orange patches at middle of wing, divided by darker streak along fold between M 1 and M 2.</p> <p>Abdomen: Male genitalia (Fig. 4C) with tegumen moderately sclerotized; uncus flat, narrow, subtriangular, with dense setae dorsally, laterally connected with long, narrow, curved and raised subtriangular socii, densely setose, with two huge thorns on distal process, with long, only weakly curved, sclerotized gnathos arms; vinculum weakly sclerotized, wide; juxta small, caulis rather short; dorsal anellus process only slightly narrower medially, phallus long, slightly curved medially, bulbous in basal half and slight tapering to apex; valva slightly curved, elliptical with narrowly rounded apex; sacculus only small basal thickening, with dense setae along margin; modified scales on outer valva surface scattered.</p> <p>Holotype: ♂, Thailand, Chanthaburi Prov., Ang-Ed Com. for., ca 33 m, 12°36'04"N 102°19'50"E, 22–23 Dec 2011, N. Pinkaew. np 6233, ♂ genitalia slide NP 2071. Deposited in KKIC.</p> <p> <b>Etymology.</b> The specific epithet <i>curvicosta</i> refers to the strongly curved costa of the forewing just before the apex.</p> <p> <b>Distribution.</b> Thailand.</p> <p> <b>Remarks.</b> The single specimen was collected in the dry season in an area of a 50-year old rubber tree plantation which was reforested to secondary forest.</p>Published as part of <i>Muadsub, Sopita & Pinkaew, Nantasak, 2014, Sirindhornia Pinkaew and Muadsub (Lepidoptera: Tortricidae), a new enarmoniine genus from Thailand, pp. 53-63 in Zootaxa 3869 (1)</i> on page 61, DOI: 10.11646/zootaxa.3869.1.5, <a href="http://zenodo.org/record/4930835">http://zenodo.org/record/4930835</a&gt

FIGURES 11–12 in Description of Antaeola rhinosa sp. nov. (Lepidoptera: Tortricidae: Olethreutinae) from Thailand

FIGURES 11–12. Extended sterigma of Antaeola rhinosa sp. nov. 11. Typical lamella postvaginalis (paratype, NP4162) 12. Atypical lamella postvaginalis (paratype, NP4161).Published as part of <i>Muadsub, Sopita & Pinkaew, Nantasak, 2023, Description of Antaeola rhinosa sp. nov. (Lepidoptera: Tortricidae: Olethreutinae) from Thailand, pp. 133-140 in Zootaxa 5323 (1)</i> on page 139, DOI: 10.11646/zootaxa.5323.1.8, <a href="http://zenodo.org/record/8204077">http://zenodo.org/record/8204077</a&gt

FIGURES 8–9 in Description of Antaeola rhinosa sp. nov. (Lepidoptera: Tortricidae: Olethreutinae) from Thailand

FIGURES 8–9. Extended male genitalia parts of Antaeola rhinosa sp. nov. (holotype) 8. Uncus, socii, and gnathos 9. Ventroapical corner of sacculus.Published as part of Muadsub, Sopita & Pinkaew, Nantasak, 2023, Description of Antaeola rhinosa sp. nov. (Lepidoptera: Tortricidae: Olethreutinae) from Thailand, pp. 133-140 in Zootaxa 5323 (1) on page 137, DOI: 10.11646/zootaxa.5323.1.8, http://zenodo.org/record/820407

FIGURE 7 in Description of Antaeola rhinosa sp. nov. (Lepidoptera: Tortricidae: Olethreutinae) from Thailand

FIGURE 7. Morphological features of male genitalia of Antaeola rhinosa sp. nov. (holotype).Published as part of <i>Muadsub, Sopita & Pinkaew, Nantasak, 2023, Description of Antaeola rhinosa sp. nov. (Lepidoptera: Tortricidae: Olethreutinae) from Thailand, pp. 133-140 in Zootaxa 5323 (1)</i> on page 136, DOI: 10.11646/zootaxa.5323.1.8, <a href="http://zenodo.org/record/8204077">http://zenodo.org/record/8204077</a&gt

Antaeola rhinosa Muadsub & Pinkaew 2023, sp. nov.

Antaeola rhinosa sp. nov. (Figs. 1–12) Type material. Holotype: ♁. THAILAND: Nakhon Nayok Prov., Khao Yai N.P., 14°25ˊ55˝N 101°24ˊ05˝E, alt. 700 m, 4–7 Jan. 2017, N. Pinkaew et al. leg., np9807 (genitalia slide NP2935). Deposited in KKIC. Paratypes: 19♁, 6♀. THAILAND: Chaiyaphum Prov., Phu Khieo W.S., 16°22ˊ56˝N 101°33ˊ51˝E, alt. 875 m, 13 Mar. 2018, np11772 (♀, genitalia slide NP4162). Nakhon Nayok Prov., Khao Yai N.P., same collection data as holotype, 13 Sep. 2014, np8830 (♁); same collection data as holotype, 11 Jan. 2016, np7951 (♁); same collection data as holotype, 14 Jan. 2016, np8027 (♁, genitalia slide NP3519); same collection data as holotype, 5 May 2016, np8421 (♁); same collection data as holotype, 8 Sep. 2016, np9277 (♁); same collection data as holotype, 27 Nov. 2016, np9397 (♁); same data as holotype, np9805 (♁); 14°21ˊ49˝N 101°24ˊ37˝E, alt. 1,140 m, 20 Jan. 2010, np3291 (♁, genitalia slide NP1314), np3292 (♀, genitalia slide NP1312), np3339 (♀, genitalia slide NP4161); 14°21ˊ56˝N 101°24ˊ24˝E, alt. 1,140 m, 6 Sep. 2016, np9168 (♁, genitalia slide NP3521); 14°14ˊ13˝N 101°23ˊ37˝E, alt. 400 m, 5 Sep. 2016 (Day), np9077 (♀, genitalia slide NP3520); 14°26ˊ18˝N 101°22ˊ24˝E, alt. 740 m, 26 Nov. 2016, np9340 (♁); 14°23ˊ56˝N 101°22ˊ16˝E, alt. 786 m, 12 Sep. 2010, np4812 (♁). Sa Kaeo Prov., Pang Sida N.P., 14°02ˊ25˝N 102°15ˊ56˝E, alt. 310 m, 1 Mar. 2017, np9579 (♁); same collection data as preceding, 18 Oct. 2017, np11138 (♁), np11151 (♁, genitalia slide NP4160), np11178 (♁); 14°07ˊ37˝N 102°15ˊ30˝E, alt. 610 m, 17 Oct. 2017, np11017 (♁). Chanthaburi Prov., Khao Soi Dao N.P., 13°05ˊ49˝N 102°10ˊ21˝E, alt. 606 m, 18 May 2008, np2751 (♁, genitalia slide NP4159). Nakhon Si Thammarat Prov., Khao Nan N.P., 08°55ˊ25˝N 99°39ˊ49˝E, alt. 131 m, 9 Dec. 2007, np2528 (♁, genitalia slide NP1143); 08°48ˊ07˝N 99°34ˊ57˝E, alt. 152 m, 16 Feb. 2007, np2091 (♀, genitalia slide NP1021); same collection data as preceding, 15 Apr. 2007, np2252 (♀, genitalia slide NP932). Narathiwat Prov., Hala-Bala W.S., 05°47ˊ49˝N 101°50ˊ03˝E, alt. 60 m, 23‒29 Jan. 2012, np6611 (♁, genitalia slide NP2860). All were collected by N. Pinkaew et al. and deposited in KKIC. Diagnosis. The wing pattern of Antaeola rhinosa sp. nov. is superficially similar to that of A. antaea (Meyrick, 1912) but differs by the absence of a longitudinal creamy band along the subcostal area that is present in A. antaea. The male genitalia of A. rhinosa are distinguished from those of A. antaea by having a long, strongly curved uncus, with a narrowed apex bearing a distinct, narrow, horn-like process dorsoapically, whereas the uncus of A. antaea has a wide, spoon-shaped apex without a horn-like process. The female genitalia of A. rhinosa are distinguished by a pair of extended large oval spinulose sterigma lobes, whereas A. antaea has an aciculate heart-shaped sterigma. Description. Head (Figs. 3–4): Lower frons greyish white, upper frons light brown; vertex scales brown mixed with dark brown, with mostly light brown apices; antenna brown; labial palpus porrect, first segment light brown, white dorsally, second segment curved upward, widened apically, light brown mixed with brown with a yellowish white spot dorsobasally and a transverse brown band in apical 1/3 (paler in female), apical segment rather short and slender, obtuse, light brown mixed with brown. Thorax: Pronotal collar light brown mixed with brown; mesonotum and tegulae dark brown mixed with brown, posterior crest dark brown with light brown apices; male hind tibia (Fig. 6) with a tuft of long, creamy scales (i.e., hair pencil) extending from base to terminal end. Forewing subrectangular, wingspan 12.8–16.0 mm in males (n = 20) (Fig. 1), 16.1–17.1 mm in females (n = 6) (Fig. 2); costal margin evenly curved, termen slightly sinuate with two white marks, first a narrow transverse stripe between wing apex and M 2, second a spot between M 3 and CuA 1; costal strigulae well developed, light brown to greyish white oblique streaks separated by small blackish streaks, strigulae 1–2 paired, 3–4 double paired, 5–7 paired, 8 and 9 single; male wing patten mostly darker than female; with brown ground color mixed with diffused longitudinal dark brown narrow stripes, especially between distinct dark brown wing veins, with a distinct longitudinal, light brown, narrow stripe below vein Cu, extending from wing base to end of closed cell, tornal area light brown with dark brown narrow stripes between veins M 3 to CuA 2, basal 1/2 of dorsum with a large, longitudinal, dark brown patch continuing as a row of dark brown spots to near tornus; fringe scales greyish white except dark brown from M 2 to between M 3 and CuA 1; underside brown except greyish white between 1A+2A and dorsum, with yellowish white spots along costa, termen with two transverse, narrow, whitish lines from wing apex to between M 1 and M 2 and from M 3 to CuA 1. Hindwing rather broad, subrectangular, brown, slightly paler toward wing base, anal margin sclerotized, folded (Fig. 5); underside brown. Abdomen: Male genitalia (Figs. 7–9) with subrectangular tegumen, moderately sclerotized and wide, with moderately dense scale sockets laterally; uncus long and slender, basal 1/3 widened, subtriangular, with sclerotized lateral edges, remaining part narrow, medial 1/3 strongly hooked; apical 1/3 slightly tapered to obtuse apex, densely setose except at apex, with a small subtriangular lobe apicoventrally and a rather short, narrow, horn-shaped process, pointed apically, slightly curved upward; socius fused to sclerotized shoulder of tegumen, with moderately dense setae; gnathos large and sclerotized, originating at 1/2 of tegumen in length, pointing upward, lateral arms moderately wide with patch of small teeth laterally, apical 1/3 fused, subrectangular, with a narrow, median dentate patch, with wide, truncated apex and shallow excavation medially in some specimens (Fig. 8); vinculum moderately wide; juxta subtriangular; caulis rather short; anellus surrounding basal 1/5 of phallus; phallus moderately long, basal half subcylindrical, apical half tapered to apex, cornuti absent; valva long and slender, dorsobasally strongly curved; sacculus rather small, about 1/3 length of cucullus, with a patch of moderately dense setae basally, ventroapical corner round, irregularly serrate (Fig. 9), valva with strongly curved neck, medially with a short horizontal ridge, densely covered with short setae; cucullus long, slender, slightly narrowed medially, rounded apically, densely setose, mixed with dense spines along outer margin, ventrobasal angle round with an oblique patch of dense, moderately long spines reaching costal margin. Female genitalia (Figs. 10–12) with papillae anales densely setose, tergum VIII large and sclerotized, with subtriangular lateral extension densely microtrichiate, sternum VII moderately sclerotized, wide, deeply concave posteromedially, posterior half with dense microtrichia; ostium bursae small, elliptical; sterigma (Figs. 11–12) with a small semicircular ring-shaped lamella antevaginalis, with distinctly, large, sclerotized, spinulose lamella postvaginalis, a pair of separated ridges dorsally, ventrally fused with two large, protruding, divergent, oval spinulose lobes; colliculum narrow, 1/5 length of ductus bursae; ductus bursae long and slender; ductus seminalis arising near middle of ductus bursae; corpus bursae large, ovate, as long as ductus bursae, two small thorn-like signa from expanded subcircular sclerotized bases. Distribution. Thailand (Chaiyaphum, Chanthaburi, Nakhon Nayok, Nakhon Si Thammarat, Narathiwat, and Sa Kaeo Provinces). This new species can be found in a wide range of elevation between 60– 1,140 m. Etymology. The specific epithet rhinosa comes from rhinos (Greek) and refers to the horn-shaped process at the uncus tip that reminiscent of a rhinoceros’s horn.Published as part of Muadsub, Sopita & Pinkaew, Nantasak, 2023, Description of Antaeola rhinosa sp. nov. (Lepidoptera: Tortricidae: Olethreutinae) from Thailand, pp. 133-140 in Zootaxa 5323 (1) on pages 134-139, DOI: 10.11646/zootaxa.5323.1.8, http://zenodo.org/record/820407