A complete checklist with new records and geographical distribution of the rove beetles (Coleoptera, Staphylinidae) of Brazil

This paper presents the first comprehensive list of 2,688 species of Staphylinidae (Coleoptera) recorded from Brazil. The list is based on the taxonomic and ecological literature, and new records from some insect collections, and includes locality references for each species. In addition, Brazilian localities and the country-level distribution outside of Brazil are provided for each species. Brazilian localities are organized by state, and include the bibliographic reference and page number where each locality was reported. All localities are geo-referenced, organized by state, and listed in an Appendix.Este trabalho apresenta a primeira lista completa das 2.688 espécies de Staphylinidae (Coleoptera) registradas para o Brasil. A lista inclui todas as localidades citadas para cada espécie e baseia-se na literatura taxonômica e ecológica disponíveis. Cada localidade inclui a referência bibliográfica e o número da página onde foram citadas. Também são apresentados registros inéditos obtidos de algumas coleções de insetos. Além das localidades brasileiras são citados todos os países com ocorrência conhecida para cada espécie. As localidades brasileiras, listadas no Apêndice, estão organizadas por estado e georreferenciadas

Revision of Oedichirus.

137 p. : ill. (some col.), maps ; 26 cm.The New World species of Oedichirus Erichson, 1839, are revised, redescribed, or newly described, illustrated, and included in keys for identification. The morphology of the ventral pterothoracic sclerites is examined and Matsuda's (1970) interpretation is largely adopted over that of Ferris (1940b). Newly discovered characters of the median gonocoxal plate and associated vulvar plate, including its microstructures, are discussed. Prior to the present paper New World species were known in Brazil and Costa Rica, but are herein newly reported for the Dominican Republic, Mexico, Ecuador, Peru, Bolivia, and Argentina; a specimen intercepted at a port in the United States was said to have come from Nicaragua. Eight species were previously described; 19 new ones are added herein, 12 from Brazil (O. apiculus, O. batillus, O. bicristatus, O. bullaglaber, O. bullahirtus, O. clavolateralis, O. clavulus, O. echinatus, O. exilis, O. glabrihamus, O. lunatus, and O. procerus), two from Mexico (O. isthmus and O. sinuosus), and one each from Argentina (O. misionesiensis), Bolivia (O. dilophus), Ecuador (O. distortus), Peru (O. hamatus), and Dominican Republic (O. dominicanensis). A neotype is designated for O. geniculatus and lectotypes are designated for O. brunneus Wendeler, O. pictipes Bierig, O. ohausi Wendeler, O. optatus Sharp, O. sparsipennis Bernhauer, and O. speculifrons Bernhauer. Some species are included in species groups

Classification of species groups, phylogeny, natural history, and catalogue (Coleoptera, Staphylinidae, Oxytelinae)

367 p. : ill., maps ; 26 cm.Includes bibliographical references (p. 335-355) and indexes."The purpose of this monograph is to review and reinterpret what is known about Bledius with the goal of prompting further study of the genus. Included are a new infrageneric classification for the world; discussions of the life cycle, habitat, distribution, enemies, chemical secretions, economic importance, immature stages, fossils, and phylogeny; and a cross-indexed annotated catalogue to the taxonomic names and associated literature. Formal taxonomic changes include the following: Bledius minniensis from Armenia is a new species, Neobledius is a new synonym of Bledius and Elbidus, and Microbledius and Psamathobledius are new synonyms of Bledius. New combinations include B. karachiensis transferred from Neobledius and actitus, litoreus, and playanus transferred from Microbledius. New names include albanicus and jutlandensis which replace the preoccupied nebulosus Koch and atlanticus Lohse, respectively. The following names are labeled as new status: B. castaneipennis Mannerheim is elevated from synonymy with opacus Block; atramentarius Rottenberg is elevated from synonymy with and replaces bos Fauvel which was described a year later; limicola Tottenham is elevated from synonymy with and replaces germanicus Wagner which is preoccupied by germanicus Gravenhorst; minor Mulsant and Rey is elevated from synonymy with and replaces devillei Bondroit which was an unnecessary replacement name. Two names are emended: cariniceps for carinlceps and ghesquierei for guesquierei. Six names, bubalus Gistel, castaneus Mulsant and Rey, chimerinus Gistel, germanicus Gravenhorst, gyllenhalii Laporte, and westerhauseri Gistel are used herein for the first time since their original publication. All eight currently recognized subgenera are treated as synonyms of Bledius. In their place a classification of 34 species groups is proposed. This classification is based on examination of 422 of the 439 species of Bledius. The species were assigned to species groups based on examination of type material or subsequently identified specimens. Nine species were assigned to species groups from characters in the original description -- each such assignment is clearly indicated. Eight species are listed as incertae sedis for one or more of the following reasons: material was unavailable for study, the description did not cite sufficient information for placement in a species group, the types were lost, or the species was unknown. Descriptions, illustrations, a list of included species and localities, and a distributional map are provided for each species group along with a key to the group. No effort was made to define or revise species. The monophyly of Bledius is established by the presence of a cluster of secretory pores in a prosternal pit or depression and the possession in females of a pair of elongate, undivided genital sclerites on segment IX. Both Bledius and its sister genus Eppelsheimius have a central row of spine-like setae on the hypopharynx; they share other characters as well. The relationships among the species groups are shown in a cladogram on which many characters are homoplasic. Bledius comprises two main lineages. One with 14 species groups and 88 species includes, among others, the large robust species with horns. Most species of this branch live in saline habitats. Two of the species groups are restricted to the New World. The sister lineage includes the remaining 20 species groups and more than 340 species. A terminal cluster of species groups along with species in three other groups live in saline habitats; the others, including the four most speciose groups, live in freshwater habitats. In this lineage 3 species groups are confined to the New World, 11 to the Old, and the remaining 6 are widespread in both hemispheres. The primitive habitat for Bledius was near salt water. One descendent remains there; the other adapted to the freshwater habitat. In this second lineage some highly speciose groups developed near freshwater and there was a reinvasion of the saline habitat. Bledius is a large genus of 439 species, some of which occur in huge populations. Species are found on all continents except Antarctica and on most continental islands but are absent from most oceanic islands. Although only a third of the species lives in saline habitats (inland as well as coastal), these species represent two-thirds of the species groups. Two-thirds of the species live in freshwater habitats, representing the remaining third of the species groups. Bledius may have lived as far back as the Cretaceous. Nine fossil species have been described as belonging in Bledius, however, two of these are no longer included. Of the remaining seven only Bledius glaciatus from the Pleistocene and B. primitiarum of the Oligocene are likely to be Bledius. Another probable member of Bledius is Staphylinus lesleyi Scudder also of the Oligocene; this species has not been moved to Bledius. Although the oldest probable fossils of Bledius are about 35 million years old, evidence suggests that the Oxytelinae existed about 150 million years ago during the Jurassic. Species of Bledius are most abundant in unvegetated or lightly vegetated, sunny, moist sand adjacent to rivers, lakes, and oceans. They preferentially select their habitat based on soil moisture, salinity, texture, and, indirectly at least, size of the sand grains, availability of food, and amount of shade. The pressure of predators may influence habitat limits but that may be partly due to the physiological weakening of a Bledius that is not living in its optimal habitat. Although as yet undemonstrated, food plant specificity may influence habitat selection by Bledius. Bledius lives in burrows which adults excavate with their mandibles and strengthen with their protibiae. The burrows are several millimeters to about 40 cm deep and about 1 to 5 mm in diameter. Larvae also construct and live in burrows. Eggs are laid in special egg chambers; some species cache food in the burrow. The burrow serves not only as living quarters for Bledius but helps protect them. Bledius can live at a relative humidity of 100 percent to no less than 93 percent. The humidity within the burrow is nearly 100 percent. Some seacoastal species live in the intertidal zone where they remain during high tide. Individuals that stay in the burrow during high tide live in an air bubble, which acts as a physical gill, and are immediately active after the tide recedes. Individuals exposed directly to seawater fly away or survive by becoming comatose and recover only slowly when removed from the water; most, however, die after a few hours of immersion. Usually the adults overwinter in deep burrows. Eggs are deposited during spring or early summer and develop in two to three weeks. The larval stage lasts for six to eight weeks and the five instars are each of undetermined length. Pupae exist for about a week. Egg to adult then is two to three months. During the summer adults live for three to six weeks. Females oviposit several times; the number of generations per year is unclear but may vary according to species and climate. Larval characters have been published for 19 species. Adults and larvae feed on algae and diatoms that live in the moisture surrounding each sand grain. The green algae there are said to be stunted filamentous forms. Some Bledius facultatively store algae. Some species are said to exhibit subsocial behavior by attending to their offspring. Evidence for this interpretation is inconclusive. Carabid beetles of Dyschirius are regarded as the principal predators of Bledius. However, little evidence is found to support a species-specific host-prey relationship. More likely, the relationship of Bledius and Dyschirius is based on geographical distribution and habitat preference. Over 50 species of Dyschirius are reported with over 80 species of Bledius. Other Carabidae reported as actual or probable predators of Bledius include species of Cillenum, Bembidion, Pogonus, Dicheirotrichus, Clivina, and Schizogenius. The ichneumonid wasp, Barycnemis blediator, is a parasitoid of Bledius spectabilis. Sandpipers and rainbow trout also eat Bledius. Fungi of the order Laboulbeniales commonly infest Bledius and, although usually regarded as benign, one report suggests otherwise. Species of Bledius secrete a characteristic, penetratingly fragrant chemical of five components. That this secretion functions as a defensive chemical is debatable. No experimentally supported hypotheses have explained the function of this compound. One species in Japan was reported to do economically significant damage to clay beds used to produce salt by evaporation of seawater. The literature for and taxonomic names used in Bledius are summarized in a catalogue. Fifteen generic-level names have been used in Bledius; 646 species-level names have been used, 434 of which are presently listed as valid and extant. All names that have been included in Bledius are listed in the catalogue. The species names are cross-indexed and the references annotated"--P. 5-6

Subtribe Cylindroxystina

83 p. : ill., maps ; 26 cm.Includes bibliographical references (p. 83)."The Neotropical tribe Cylindroxystini is reduced to a subtribe, Cylindroxystina, of the Paederini. Cylindroxystina is distinguished from all other subtribes of the Paederinae by the sickle-shaped, edentate mandibles, nipple-shaped fourth segment of the maxillary palpus, single pair of paratergites of abdominal segment III, and cylindrical abdomen. The paratergites of segments IV to VI are absent, the tergum and sternum of each segment are fused, and the segment is cylindrical. Segment VII also lacks paratergites, but the tergum and sternum remain separate. Two genera, Cylindroxystus and Neolindus, are included in the subtribe. For Cylindroxystus, 2 species, pluviosus and borinquense, both from the Caribbean, are transferred from Lobrathium and 11 new species are described as follows: crenus, flavus, luridus, and sinuosus from Ecuador; furvus and messus from Peru; alleni and redactus from Panama; lyrus from Venezuela; concavoperculum from Guyana; and cavus from Brazil. Cylindroxystus is distinguished from Neolindus by the third antennal segment that is cupulate or has a carina surrounding the apex. Twenty-seven new species are described in Neolindus as follows: agilis, brewsterae, densus, hamatus, lodhii, sinuatus, and unilobus from Brazil; cephalochymus from Peru; bidens, bullus, dichymus, lirellus, milleri, parallelus, procarinatus, prolatus, and retusus from Ecuador; pumicosus from Colombia; brachiatus, plectrus, and rudiculus from Venezuela; apiculus, basisinuatus, campbelli, and punctogularis from Panama; and cuneatus and incanalis from Costa Rica. Neolindus is separated from Cylindroxystus by the presence of one or two pairs of cephalic trichobothria which are lacking in Cylindroxystus. Until now Cylindroxystus was known only by one specimen from Costa Rica, and Neolindus by a few specimens from Brazil and Peru. The subtribe is redescribed and both genera and all the species are described and illustrated and their phylogenetic relationships discussed. Keys to the males of both genera are provided. Most of the present work is based on characteristics of the male. Females were unavailable for over half of the species, and it was not always possible to reliably associate the sexes. Species of both genera have been collected rarely and most seem to have been collected from leaf litter on the forest floor at some distance from streams"--P. 3

Oxytelinae

8 p. : ill. ; 24 cm.Includes 1 bibliographical reference (p. 8)

Aequatorialis, mandibularis, and semiferrugineus groups and two new genera (Coleoptera, Staphylinidae, Oxytelinae)

p. 113-[254] : ill., maps ; 27 cm.Includes bibliographical references (p. 252-253) and index."Bledius is redescribed, and a key to the species-groups of the New World is presented. The species of the aequatorialis, mandibularis, and semiferrugineus groups of Bledius Leach from the Nearctic and West Indian areas and two new genera, Microbledius and Psamathobledius, are described. Illustrations, keys, diagnoses, and distributional and habitat data are presented for each species. Three new species, Microbledius playanus, Microbledius actitus, and Microbledius litoreus are described and M. forcipatus (Le Conte), M. albidus (Sharp), M. albipennis (Bernhauer), M. bifasciatus (Sharp), M. miles (Bernhauer), M. minutissimus (Bernhauer), and M. weiseri (Bernhauer) are transferred from Bledius. Psamathobledius contains P. caribbeanus (Blackwelder), P. microcephalus (Fauvel) and P. punctatissimus (Le Conte) (B. esposus Blackwelder is a new synonymy); all are transferred from Bledius. Bledius aequatorialis Mutchler, B. beattyi Blackwelder, and B. ceratus Blackwelder are included in the aequatorialis group. Bledius mandibularis Erichson, B. pallipennis Say (B. gularis Le Conte is a new synonymy), B. fortis Le Conte, B. ferratus Le Conte, and B. jacobinus Le Conte (B. actus Herman is a new synonymy) are in the mandibularis group. Bledius semiferrugineus Le Conte (B. canaliculatus Notman is a new synonymy), B. foraminosus Casey (B. relictus Fall and B. deceptivus Fall are new synonymies), B. tallaci Fall, B. gravidus Casey (B. missionensis Hatch is a new synonymy), B. rotundicollis Le Conte, B. rubiginosus Erichson, B. fumatus Le Conte, B. philadelphicus Fall (B. dickersoni Notman and B. mixtus Notman are new synonymies), and B. analis Le Conte, B. nitidicollis Le Conte, B. assimilis Casey, and B. coulteri Hatch are included in the semiferrugineus group"--P. 115

Armatus, basalis, and melanocephalus groups (Coleoptera, Staphylinidae, Oxytelinae)

p. 73-172 : ill., maps ; 26 cm.Includes bibliographical references (p. 170-171) and index."In the present paper, the second of a three part series on Bledius, the armatus, basalis, and melanocephalus groups from the Nearctic region are discussed. In addition to a revised key to the species groups of the New World, illustrations, keys, diagnoses and distributional and habitat data are presented for each species. Bledius notialus, B. aquilonarius, and B. thinopus are newly described. Bledius cuspidatus Le Conte, B. agonus Casey, B. tenuis Casey, and B. nelsoni Hatch are new junior synonyms of B. flavipennis Le Conte. Bledius lectus Casey, B. piceus Fall, B. specularis Fall, B. regularis Fall, and B. condonensis Hatch are new junior synonyms of B. opacifrons Le Conte. Bledius nigriceps Notman is a new junior synonym of B. politus Erichson. Bledius gradatus Fall and B. boddyi Hatch are new junior synonyms of B. eximius Casey. Bledius arizonensis Fall is a new junior synonym of B. strenuus Casey. Bledius ignavus Casey and B. misellus Casey are new junior synonyms of B. basalis Le Conte. Bledius confinus Fall is a new junior synonym of B. melanocephalus (Say). Bledius melanocephalus (Say), the only species in the melanocephalus group, is recognized and discussed for the first time in more than 150 years. Bledius cordatus (Say), B. basalis Le Conte, B. turbulentus Casey, B. dimidiatus Le Conte, B. opaculus Le Conte, B. neglectus Casey, and B. thinopus, new species, comprise the basalis group and, but for B. dimidiatus, all live on the coast of eastern North America. The armatus group includes B. flavipennis Le Conte, B. nitidiceps Le Conte, B. opacifrons Le Conte, B. ineptus Casey, B. consimilis Fall, B. monstratus Casey, B. fenyesi Bernhauer and Schubert, B. politus Erichson, B. episcopalis Fall, B. eximius Casey, B. bellicus Blackwelder, B. strenuus Casey, B. aquilonarius, new species, and B. notialus, new species. All species of the armatus group except B. politus live principally west of the Mississippi River"--P. 75

Annularis and emarginatus groups (Coleoptera, Staphylinidae, Oxytelinae)

145 p. : ill., maps ; 26 cm.Includes bibliographical references (p. 141-143) and index."In the present report, the third in a series on Bledius, the annularis and emarginatus groups are discussed and a new species is described in the aequatorialis group. A key to the species, descriptions, diagnoses, illustrations, and distributional and habitat data for each species are presented. Bledius habrus, melanocolus, aurantius, jucundus, naius, nardus, omega, venus, viriosus, wudus, and zophus are newly described. Thirteen nominal species are recognized as synonyms. Bledius albidipennis Bernhauer and ornatus LeConte are new junior synonyms of albonotatus Mäklin; apicalis Fall is a new junior synonym of diagonalis LeConte; adjustus Casey and transitus Fall are new junior synonyms of gentilis Casey; kincaidi Hatch is a new junior synonym of parvicollis Casey; rusticus Fall is a new junior synonym of bicolor Casey; luteipennis LeConte, medialis Fall, oregonensis Hatch, and pleuralis LeConte are new junior synonyms of suturalis LeConte; and borealis Blatchley and bowronensis Hatch are new junior synonyms of turgidus Casey. The species in the United States included in the emarginatus group are cognatus LeConte, emarginatus (Say), and wudus Herman. They are found in the eastern and southeastern United States. The species are similar and not easily distinguished. Bledius albonotatus Mäklin, aurantius Herman, bicolor Casey, cedarensis Hatch, confusus LeConte, diagonalis LeConte, gentilis Casey, gracilis Casey, habrus Herman, jucundus Herman, laticollis LeConte, melanocolus Herman, monticola Casey, naius Herman, nardus Herman, newelli Hatch, omega Herman, parvicollis Casey, persimilis Fall, phytosinus LeConte, ruficornis LeConte, suturalis LeConte, tarandus Herman, tau LeConte, turgidus Casey, venus Herman, villosus Casey, viriosus Herman and zophus Herman are species from the United States and Canada included in the annularis group and which I can identify. Also included in the annularis group are nine species listed together in what I call the annularis complex: annularis LeConte, breretoni Hatch, honestus Casey, languidus Casey, mysticus Fall, nebulosus Casey, sinuatus LeConte, stabilis Casey, and washingtonensis Hatch. After years of study and collecting I have been unable to resolve the species of this complex which is widespread in Canada and the northern United States. I was unable to resolve three other species: fasciatus (Say), longipennis Mäklin, and verticalis Notman; the types of each have been lost but I discuss each of them. The annularis group species are widespread in the United States and Canada and most are found in shaded soil near fresh water. Only two, albonotatus and newelli, are associated with coastal, saline habitats. Several species, notably ruficornis, tarandus, and turgidus, have transcontinental distributions, and one, albonotatus, is found along the coast from western Alaska to Baja California. The addition of a new species, susae, to the aequatorialis group brings that group up to four. However, susae is the only one of the group known only from the United States which is found only on the mainland. It is also the sole species of Bledius in which the females have a pronotal horn; although pronotal horns are known in a number of New and Old World species, heretofore no females have been reported with such a modification. Since publication of Parts I and II of this monograph, I have examined over 6000 more specimens of 41 previously revised species. In Appendix II all the new records are included; for many species extensions of their previously known range are recorded. Lectotypes are designated for the species of Bledius described by LeConte, Casey, and Fall. For Part III, 28,408 specimens were examined"--P. 4

Nomenclatural changes, Staphylinidae.

83 p. ; 26 cm.Includes bibliographical references (p. 61-83).More than 775 nomenclatural changes are proposed and nomenclatural problems are discussed for the Staphylinidae. Two generic names are proposed as replacement names: Torobus for species formerly in Trigonopselaphus, and Ryvkinius for the preoccupied Mesoporus Ryvkin. Type species are fixed or problems with the type species are discussed for: Entomoculia, Eumegalopsidia (an unavailable name), Anthobium, Lesteva, Phyllodrepoidea, Aleioglyphesthus, Bolitogyrus, Cephalonthus, Diatrechus, Euremus, Indoquedius, Philothalpus, Philonthopsis, Pseudoremus, Leptophius, Stenus (Nestus), Coproporus, and Paratachinus. The type species for Entomoculia, Anthobium, Phyllodrepoidea, Leptophius, and Stenus (Nestus) are fixed under provisions of article 70.3 of the Code. Emendations are cited or one of multiple original spellings adopted in Bledius, Bryoporus, Carpelimus, Edaphus, Eleusis, Gauropterus, Indosorius, Lispinus, Lithocharodes, Medon, Osorius, Philonthus, Piestus, Pinophilinus, Platydracus, Priochirus, Prognathoides, Quedius, Stenus, and Trigonuras. New combinations are proposed in Arrhenopeplus (3), Dialycera (2), Eusphalerum (14), Nacaeus (19), Pseudoxyporus (4), Homalotrichus (1), Thinodromus (2), Gabrius (1), Platydracus (57), Torobus (9), Bryophacis (7), Ischnosoma (16), Lordithon (66), Ryvkinius (1), and Sepedophilus (244). Fifty-nine new synonyms are listed for Micropeplus (1), Anthophagus (2), Eusphalerum (1), Phloeonomus (1), Eleusis (1), Leptochirus (1), Neolosus (1), Osorius (5), Priochirus (1), Anotylus (1), Bledius (1), Carpelimus (1), Ochthephilus (1), Apoquedius (1), Diatrechus (1), Erichsonius (2), Gabrius (2), Gastrisus (1), Hesperus (2), Heterothops (2), Leptacinus (1), Notolinus (1), Phallolinus (1), Philonthus (6), Quedius (5), Staphylinus (2), Tasgius (1) Xantholinus (4), Stenus (7), Sepedophilus (1), and Tachinus (1). Polyphematiana E. Strand is a junior synonym of Trigonopselaphus Gemminger and Harold. Eleven names are resurrected from synonymy, one each in Phyllodrepoidea, Anotylus, Oxytelus, Loncovilius, Philonthus, Platydracus, Lordithon, Mycetoporus, and Tachinus and two in Stenus. Under provisions of article 23.9.1 of the Code, 28 junior synonyms are protected in Amphichroum (1), Carpelimus (1), Deleaster (1), Proteinus (1), Bisnius (1), Leptacinus (1), Megalinus (1), Neobisnius (1), Ocypus (1), Philonthus (3), Quedius (4), Tasgius (2), Stenus (5), Bolitobius (1), Lordithon (1), Mycetoporus (1), and Tachyporus (2). Provisions of the same article protect 5 junior homonyms in Eusphalerum (1), Xylodromus (1), Quedius (1), Tachinus (1), and Tachyporus (1). Under provisions of article 23.9.3 of the Code, 15 junior synonyms will be referred to the Commission for rulings under the plenary power. Pending the outcome of these cases, use of the junior name is maintained. The affected species are in Eusphalerum (1), Lesteva (1), Omalium (1), Phloeostiba (1), Xylodromus (1), Anotylus (1), Bledius (1), Carpelimus (1), Ocypus (1), Philonthus (1), Quedius (4), and Lordithon (1). Under provisions of article 23.9.5 of the Code, 64 junior primary homonyms will be referred to the Commission for rulings under the plenary power. Pending the outcome of these deliberations, use of the junior name is maintained. The species are in Eusphalerum (1), Mannerheimia (1), Omaliomimus (1), Omaliopsis (1), Omalium (4), Phyllodrepa (1), Pycnoglypta (1), Xylodromus (1), Anotylus (1), Bledius (1), Carpelimus (1), Oxyporus (1), Belonuchus (2), Bisnius (1), Cafius (2), Cheilocolpus (1), Diatrechus (1), Endeius (1), Gabrius (5), Hesperus (2), Leptacinus (1), Nordus (1), Paederomimus (1), Philonthus (10), Platydracus (3), Quedius (2), Staphylinus (6), Xantholinus (2), Xanthopygus (1), Xenopygus (2), Carphacis (1), Coproporus (1), Tachinomorphus (1), and Tachinus (3). Replacement names are proposed for 90 preoccupied names in 37 genera: Euaesthetus (1), Leptotyphlus (1), Micropeplus (1), Eusphalerum (2), Lesteva (1), Olophrum (1), Omalium (2), Clavilispinus (1), Eleusis (3), Holotrochus (3), Leptochirus (1), Lispinus (4), Osorius (3), Priochirus (2), Thoracochirus (1), Anotylus (2), Carpelimus (2), Platystethus (1), Atanygnathus (1), Bisnius (1), Diochus (1), Gabrius (3), Hesperus (2), Heterothops (1), Leptacinus (3), Philonthus (19), Quedius (4), Staphylinus (3), Tasgius (1), Dianous (1), Bolitobius (1), Carphacis (1), Coproporus (3), Lordithon (3), Sepedophilus (5), Tachinus (4), Tachyporus (1). Eight names are resurrected to replace preoccupied names in Anotylus (1), Philonthus (1), Quedius (1), Stenus (2), Lordithon (1), Tachinomorphus (1), and Tachinus (1). Twenty-six senior homonyms, in Hapalaraea (1), Omalium (2), Phyllodrepa (1), Bledius (1), Oxytelus (1), Bisnius (1), Quedius (1), Staphylinus (15), Carphacis (1), Sepedophilus (1), and Tachinus (1), are regarded to be "nomina dubia" and the junior names conserved by fiat, without sanction by the Code. Psephidonus is the older name, but application to the Commission will be made to reject that name in favor of the younger Geodromicus. Tachyporiniformes is an unavailable name

Rove-beetle subfamily Oxytelinae

p. 345-454 : ill. ; 27 cm.Includes bibliographical references (p. 429-433) and index