A physiological measure of shifting connections in the Rana pipiens retinotectal system

The retinotectal connections of developing Rana tadpoles and froglets have been studied using light-pipe techniques to directly assay the pattern of the projection from the retina to the tectum. The projection site of the retina surrounding the optic nerve head was determined at two different stages of development (late larval and metamorphic frog) on the same animal. Small electrolytic marker lesions were used to mark the tectal sites to which the optic nerve head projected at these two times. Comparison of the positions of the two lesions gives a direct measure of the shift in the projection during the interlesion time interval of one week. The results indicate a shift in the projection of 275 µm week–1 in late larval life. Previous work in Xenopus using the light-pipe techniques indicated a qualitatively similar shift during equivalent stages of development, but significantly smaller in magnitude. In the present study, topographic postsynaptic units could be recorded at all stages investigated, indicating functional synapses between the optic nerve fibres and the tectum. Thus, these studies offer evidence of a significant shift in the functional connection pattern of the amphibian retinotectal map during development, in agreement with the recent anatomical data from other laboratories on the Rana and goldfish visual system

Aerothermal tests of spherical dome protuberances on a flat plate at a Mach number of 6.5

Aerothermal tests were conducted in the Langley 8-Foot High-Temperature Tunnel at a Mach number of 6.5 on a series of spherical dome protuberances mounted on a flat-plate test apparatus. Detailed surface pressure and heating-rate distributions were obtained for various dome heights and diameters submerged in both laminar and turbulent boundary layers including a baseline geometric condition representing a thermally bowed metallic thermal protection system (TPS) tile. The present results indicated that the surface pressures on the domes were increased on the windward surface and reduced on the leeward surface as predicted by linearized small-perturbation theory, and the distributions were only moderately affected by boundary-layer variations. Surface heating rates for turbulent flow increased on the windward surface and decreased on the leeward surface similar to the pressure; but for laminar boundary layers, the heating rates remained high on the leeward surface, probably due to local transition. Transitional flow effects cause heat load augmentation to increase by 30 percent for the maximum dome height in a laminar boundary layer. However, the corresponding augmentation for a dome with a height of 0.1 in. and a diameter of 14 in. representative of a bowed TPS tile was 14 percent or less for either a laminar or turbulent boundary layer

Granular Flow in a Vertically Vibrating Hopper

The behavior of the flow of glass spheres in a vertically vibrating hopper is examined. A two-dimensional hopper is mounted on a shaker that provides sinusoidal, vertical vibrations. Both the frequency and amplitude of the vibrations are adjustable. Hopper discharge rates and flow patterns are measured as the acceleration amplitude of the vibrations is increased from 0 to 4g's. Comparisons are made with unvibrated hopper flows and with a two-dimensional discrete element simulation model

Effects of vertical vibration on hopper flows of granular material

This paper examines the flow of granular material through a wedge-shaped hopper subject to vertical, sinusoidal oscillations. Experiments and discrete element computer simulations were conducted to investigate particle trajectories within and mass discharge rates from the hopper. With the hopper exit closed, side wall convection cells are observed in both the experiments and simulations. The convection cells are oriented such that particles move up along the inclined walls of the hopper and down along the centerline. Results from the computer simulation indicate that the convection cells are a result of the dilation of the granular bed during free fall and interaction with hopper walls. Measurements of the mean mass discharge rate for various vibration parameters were also made in both the experiments and simulations. The ratio of the mass discharge rate for a vibrating hopper to the mass discharge rate for a non-vibrating hopper scales with the oscillation velocity amplitude and exhibits a maximum value just greater than one for oscillation velocity amplitudes less than 0.5. The ratio is less than one for larger velocity amplitudes. A simple model taking into account the change in the effective gravity acting on the granular material over an oscillation cycle is examined. A significant deficiency in the model is that is assumes no material discharges from the hopper during part of each oscillation cycle for acceleration amplitudes greater than gravitational acceleration. Data from the simulations indicate that although the discharge rate from the hopper varies throughout an oscillation cycle, it never equals zero. The simulation was also used to examine particle horizontal position and velocity profiles at the hopper exit. Lastly, preliminary observations of the effects of localized vibration on a granular material in a closed hopper are presented

Investigation of f/2 and f/4 Waves in Granular Beds Subject to Vertical, Sinusoidal Oscillations

When a deep bed of granular material is subject to vertical, sinusoidal oscillations, a number of phenomena appear including two regimes of standing surface waves that form at one-half and one-quarter of the oscillation forcing frequency. These waves are referred to as f/2 and f/4 waves where f is the oscillation frequency. This paper presents the results from experiments and computer simulations designed to study the wavelength and wave amplitude dependence of the surface waves on the vibration parameters, collision coefficient of restriction, and the particle bed depth

Local measurements of velocity fluctuations and diffusion coefficients for a granular material flow

Measurements were made of two components of the average and fluctuating velocities, and of the local self-diffusion coefficients in a flow of granular material. The experiments were performed in a 1 m-high vertical channel with roughened sidewalls and with polished glass plates at the front and the back to create a two-dimensional flow. The particles used were glass spheres with a nominal diameter of 3 mm. The flows were high density and were characterized by the presence of long-duration frictional contacts between particles. The velocity measurements indicated that the flows consisted of a central uniform regime and a shear regime close to the walls. The fluctuating velocities in the transverse direction increased in magnitude from the centre towards the walls. A similar variation was not observed for the streamwise fluctuations. The self-diffusion coefficients showed a significant dependence on the fluctuating velocities and the shear rate. The velocity fluctuations were highly anistropic with the streamwise components being 2 to 2.5 times the transverse components. The self-diffusion coefficients for the streamwise direction were an order-of-magnitude higher than those for the transverse direction. The surface roughness of the particles led to a decrease in the self-diffusion coefficients

Viscous coupling of shear-free turbulence across nearly flat fluid interfaces

The interactions between shear-free turbulence in two regions (denoted as + and − on either side of a nearly flat horizontal interface are shown here to be controlled by several mechanisms, which depend on the magnitudes of the ratios of the densities, ρ+/ρ−, and kinematic viscosities of the fluids, μ+/μ−, and the root mean square (r.m.s.) velocities of the turbulence, u0+/u0−, above and below the interface. This study focuses on gas–liquid interfaces so that ρ+/ρ− ≪ 1 and also on where turbulence is generated either above or below the interface so that u0+/u0− is either very large or very small. It is assumed that vertical buoyancy forces across the interface are much larger than internal forces so that the interface is nearly flat, and coupling between turbulence on either side of the interface is determined by viscous stresses. A formal linearized rapid-distortion analysis with viscous effects is developed by extending the previous study by Hunt & Graham (J. Fluid Mech., vol. 84, 1978, pp. 209–235) of shear-free turbulence near rigid plane boundaries. The physical processes accounted for in our model include both the blocking effect of the interface on normal components of the turbulence and the viscous coupling of the horizontal field across thin interfacial viscous boundary layers. The horizontal divergence in the perturbation velocity field in the viscous layer drives weak inviscid irrotational velocity fluctuations outside the viscous boundary layers in a mechanism analogous to Ekman pumping. The analysis shows the following. (i) The blocking effects are similar to those near rigid boundaries on each side of the interface, but through the action of the thin viscous layers above and below the interface, the horizontal and vertical velocity components differ from those near a rigid surface and are correlated or anti-correlated respectively. (ii) Because of the growth of the viscous layers on either side of the interface, the ratio uI/u0, where uI is the r.m.s. of the interfacial velocity fluctuations and u0 the r.m.s. of the homogeneous turbulence far from the interface, does not vary with time. If the turbulence is driven in the lower layer with ρ+/ρ− ≪ 1 and u0+/u0− ≪ 1, then uI/u0− ~ 1 when Re (=u0−L−/ν−) ≫ 1 and R = (ρ−/ρ+)(v−/v+)1/2 ≫ 1. If the turbulence is driven in the upper layer with ρ+/ρ− ≪ 1 and u0+/u0− ≫ 1, then uI/u0+ ~ 1/(1 + R). (iii) Nonlinear effects become significant over periods greater than Lagrangian time scales. When turbulence is generated in the lower layer, and the Reynolds number is high enough, motions in the upper viscous layer are turbulent. The horizontal vorticity tends to decrease, and the vertical vorticity of the eddies dominates their asymptotic structure. When turbulence is generated in the upper layer, and the Reynolds number is less than about 106–107, the fluctuations in the viscous layer do not become turbulent. Nonlinear processes at the interface increase the ratio uI/u0+ for sheared or shear-free turbulence in the gas above its linear value of uI/u0+ ~ 1/(1 + R) to (ρ+/ρ−)1/2 ~ 1/30 for air–water interfaces. This estimate agrees with the direct numerical simulation results from Lombardi, De Angelis & Bannerjee (Phys. Fluids, vol. 8, no. 6, 1996, pp. 1643–1665). Because the linear viscous–inertial coupling mechanism is still significant, the eddy motions on either side of the interface have a similar horizontal structure, although their vertical structure differs

Local form interference in biological motion perception

Peer reviewedPublisher PD

Revisiting the 1954 Suspension Experiments of R. A.Bagnold

In 1954 R. A. Bagnold published his seminal findings on the rheological properties of a liquid-solid suspension. Although this work has been cited extensively over the last fifty years, there has not been a critical review of the experiments. The purpose of this study is to examine the work and to suggest an alternative reason for the experimental findings. The concentric cylinder rheometer was designed to measure simultaneously the shear and normal forces for a wide range of solid concentrations, fluid viscosities and shear rates. As presented by Bagnold, the analysis and experiments demonstrated that the shear and normal forces depended linearly on the shear rate in the 'macroviscous' regime; as the grain-to-grain interactions increased in the 'grain-inertia' regime, the stresses depended on the square of the shear rate and were independent of the fluid viscosity. These results, however, appear to be dictated by the design of the experimental facility. In Bagnold's experiments, the height (h) of the rheometer was relatively short compared to the spacing (t) between the rotating outer and stationary inner cylinder (h/t=4.6). Since the top and bottom end plates rotated with the outer cylinder, the flow contained two axisymmetric counter-rotating cells in which flow moved outward along the end plates and inward through the central region of the annulus. At higher Reynolds numbers, these cells contributed significantly to the measured torque, as demonstrated by comparing Bagnold's pure-fluid measurements with studies on laminar-to-turbulent transitions that pre-date the 1954 study. By accounting for the torque along the end walls, Bagnold's shear stress measurements can be estimated by modelling the liquid-solid mixture as a Newtonian fluid with a corrected viscosity that depends on the solids concentration. An analysis of the normal stress measurements was problematic because the gross measurements were not reported and could not be obtained

How does star formation proceed in the circumnuclear starburst ring of NGC 6951?

Gas inflowing along stellar bars is often stalled at the location of circumnuclear rings, that form an effective reservoir for massive star formation and thus shape the central regions of galaxies. However, how exactly star formation is proceeding within these circumnuclear starburst rings is subject of debate. Two main scenarios for this process have been put forward: In the first the onset of star formation is regulated by the total amount of gas present in the ring with star forming starting once a mass threshold has reached in a `random' position within the ring like `popcorn'. In the second star formation preferentially takes place near the locations where the gas enters the ring. This scenario has been dubbed `pearls-on-a-string'. Here we combine new optical IFU data covering the full stellar bar with existing multi-wavelength data to study in detail the 580 pc radius circumnuclear starburst ring in the nearby spiral galaxy NGC 6951. Using HST archival data together with Sauron and Oasis IFU data, we derive the ages and stellar masses of star clusters as well as the total stellar content of the central region. Adding information on the molecular gas distribution, stellar and gaseous dynamics and extinction, we find that the circumnuclear ring in NGC 6951 is ~1-1.5 Gyr old and has been forming stars for most of that time. We see evidence for preferred sites of star formation within the ring, consistent with the `pearls-on-a-string' scenario, when focusing on the youngest stellar populations. Due to the ring's longevity this signature is washed out when older stellar populations are included in the analysis.Comment: accepted for publication in A&A, 15 page