The FRII Broad Line Seyfert 1 Galaxy: PKSJ 1037-2705

In this article, we demonstrate that PKSJ 1037-2705 has a weak accretion flow luminosity, well below the Seyfert1/QSO dividing line, weak broad emission lines (BELs) and moderately powerful FRII extended radio emission. It is one of the few documented examples of a broad-line object in which the time averaged jet kinetic luminosity, $\bar{Q}$, is larger than the total thermal luminosity (IR to X-ray) of the accretion flow, $L_{bol}$. The blazar nucleus dominates the optical and near ultraviolet emission and is a strong source of hard X-rays. The strong blazar emission indicates that the relativistic radio jet is presently active. The implication is that even weakly accreting AGN can create powerful jets. Kinetically dominated ($\bar{Q}>L_{bol}$) broad-line objects provide important constraints on the relationship between the accretion flow and the jet production mechanism.Comment: To appear in ApJ November 1, 2008, v687n1 issu

Optical Counterparts of Ultra-Luminous X-ray Sources identified from Archival Hubble Space Telescope/WFPC2

We present a systematic analysis of archival HST WFPC2 ``Association'' data sets that correlate with the Chandra positions of a set of 44 ultra-luminous X-ray sources (ULXs) of nearby galaxies. We have improved the Chandra-HST relative astrometry whenever possible. Disparate numbers of potential ULX counterparts are found, and in some cases none are found. The lack of or low number of counterparts in some cases may be due to insufficient depth in the WFPC2 images. Particularly in late-type galaxies, the HST image in the ULX region was often complex or crowded. We therefore address various scenarios for the nature of the ULX since it is not known which, if any, of the sources found are true counterparts. The optical luminosities of the sources are typically in the range 10^4-6 L_sun. In several cases color information is available, with the colors roughly tending to be more red in early-type galaxies. This suggests that, in general, the (potential) counterparts found in early-type galaxies are likely to be older stellar populations, and are probably globular clusters. Several early-type galaxy counterparts have blue colors, which may be due to younger stellar populations in the host galaxies, however these could also be background sources. In spiral galaxies the sources may also be due to localized structure in the disks rather than bound stellar systems. Alternatively some of the counterparts in late-type galaxies may be isolated supergiant stars. The observed X-ray/optical flux ratio is diluted by the optical emission of the cluster in cases where the system is an X-ray binary in a cluster, particularly in the case of a low-mass X-ray binaries in old cluster. (abridged)Comment: 35 pages with 9 figures formatted with emulateapj. Only subset of figures 1 and 2 are shown, for full version see http://xassist.pha.jhu.edu/ptak/hst_ulx_pape

The Anatomy of Asilisaurus kongwe, a Dinosauriform from the Lifua Member of the Manda Beds (~Middle Triassic) of Africa

The diagnosis of Dinosauria and interrelationships of the earliest dinosaurs relies on careful documentation of the anatomy of their closest relatives. These close relatives, or dinosaur “precursors,” are typically only documented by a handful of fossils from across Pangea and nearly all specimens are typically missing important regions (e.g., forelimbs, pelves, skulls) that appear to be important to help resolving the relationships of dinosaurs. Here, we fully describe the known skeletal elements of Asilisaurus kongwe, a dinosauriform from the Middle Triassic Manda Beds of the Ruhuhu Basin of Tanzania. The taxon is known from many disarticulated and partially articulated remains and, most importantly, from a spectacularly preserved associated skeleton of an individual containing much of the skull, pectoral and pelvic girdles, forelimb and hindlimb, and parts of the vertebral column including much of the tail. The unprecedented detail of the anatomy indicates that Asilisaurus kongwe had a unique skull that was short and had both a premaxillary and dentary edentulous margin, but retained a number of character states plesiomorphic for Archosauria, including a crocodylian-like ankle configuration and a rather short foot with well-developed metatarsals I and V. Additionally, character states present across the skeleton of Asilisaurus kongwe suggest it is more closely related to Silesaurus opolensis than to dinosaurs; thus suggesting high homoplasy and parallel trends within Silesauridae and within lineages of early dinosaurs. The anatomy of Asilisaurus kongwe and detailed description of early members of clades found outside Dinosauria are clearly needed to untangle the seemingly complex character evolution of the skeleton within avemetatarsalians.Fil: Nesbitt, Sterling J.. Virginia Polytechnic Institute; Estados UnidosFil: Langer, Max C.. Universidade de Sao Paulo; BrasilFil: Ezcurra, Martin Daniel. Consejo Nacional de Investigaciones Científicas y Técnicas. Oficina de Coordinación Administrativa Parque Centenario. Museo Argentino de Ciencias Naturales "Bernardino Rivadavia"; Argentin

Does the intermediate-mass black hole in LEDA 87300 (RGG 118) follow the near-quadratic Mbh-Mspheroid relation?

The mass scaling relation between supermassive black holes and their host spheroids has previously been described by a quadratic or steeper relation at low masses (105 < Mbh/Mo ≲ 107). How this extends into the realm of intermediate-mass black holes (102 < Mbh/Mo < 105) is not yet clear, although for the barred Sm galaxy LEDA 87300, Baldassare et al. recently reported a nominal virial mass of Mbh = 5 104 Mo residing in a "spheroid" of stellar mass equal to 6.3 108 Mo. We point out, for the first time, that LEDA 87300 therefore appears to reside on the near-quadratic Mbh-Msph,∗ relation. However, Baldassare et al. modeled the bulge and bar as the single spheroidal component of this galaxy. Here we perform a 3-component bulge+bar+disk decomposition and find a bulge luminosity which is 7.7 times fainter than the published "bulge" luminosity. After correcting for dust, we find that Mbulge = 0.9 108 Mo and Mbulge/Mdisk = 0.04 - which is now in accord with ratios typically found in Scd-Sm galaxies. We go on to discuss slight revisions to the stellar velocity dispersion (40 11 km s-1) and black hole mass () and show that LEDA 87300 remains consistent with the Mbh-σ relation, and also the near-quadratic Mbh-Msph,∗ relation when using the reduced bulge mass. LEDA 87300 therefore offers the first support for the rapid but regulated (near-quadratic) growth of black holes, relative to their host bulge/spheroid, extending into the domain of intermediate-mass black holes

Impact of Sauropod Dinosaurs on Lagoonal Substrates in the Broome Sandstone (Lower Cretaceous), Western Australia

Existing knowledge of the tracks left by sauropod dinosaurs (loosely ‘brontosaurs’) is essentially two-dimensional, derived mainly from footprints exposed on bedding planes, but examples in the Broome Sandstone (Early Cretaceous) of Western Australia provide a complementary three-dimensional picture showing the extent to which walking sauropods could deform the ground beneath their feet. The patterns of deformation created by sauropods traversing thinly-stratified lagoonal deposits of the Broome Sandstone are unprecedented in their extent and structural complexity. The stacks of transmitted reliefs (underprints or ghost prints) beneath individual footfalls are nested into a hierarchy of deeper and more inclusive basins and troughs which eventually attain the size of minor tectonic features. Ultimately the sauropod track-makers deformed the substrate to such an extent that they remodelled the topography of the landscape they inhabited. Such patterns of substrate deformation are revealed by investigating fragmentary and eroded footprints, not by the conventional search for pristine footprints on intact bedding planes. For that reason it is not known whether similar patterns of substrate deformation might occur at sauropod track-sites elsewhere in the world

Estimating Mass Properties of Dinosaurs Using Laser Imaging and 3D Computer Modelling

Body mass reconstructions of extinct vertebrates are most robust when complete to near-complete skeletons allow the reconstruction of either physical or digital models. Digital models are most efficient in terms of time and cost, and provide the facility to infinitely modify model properties non-destructively, such that sensitivity analyses can be conducted to quantify the effect of the many unknown parameters involved in reconstructions of extinct animals. In this study we use laser scanning (LiDAR) and computer modelling methods to create a range of 3D mass models of five specimens of non-avian dinosaur; two near-complete specimens of Tyrannosaurus rex, the most complete specimens of Acrocanthosaurus atokensis and Strutiomimum sedens, and a near-complete skeleton of a sub-adult Edmontosaurus annectens. LiDAR scanning allows a full mounted skeleton to be imaged resulting in a detailed 3D model in which each bone retains its spatial position and articulation. This provides a high resolution skeletal framework around which the body cavity and internal organs such as lungs and air sacs can be reconstructed. This has allowed calculation of body segment masses, centres of mass and moments or inertia for each animal. However, any soft tissue reconstruction of an extinct taxon inevitably represents a best estimate model with an unknown level of accuracy. We have therefore conducted an extensive sensitivity analysis in which the volumes of body segments and respiratory organs were varied in an attempt to constrain the likely maximum plausible range of mass parameters for each animal. Our results provide wide ranges in actual mass and inertial values, emphasizing the high level of uncertainty inevitable in such reconstructions. However, our sensitivity analysis consistently places the centre of mass well below and in front of hip joint in each animal, regardless of the chosen combination of body and respiratory structure volumes. These results emphasize that future biomechanical assessments of extinct taxa should be preceded by a detailed investigation of the plausible range of mass properties, in which sensitivity analyses are used to identify a suite of possible values to be tested as inputs in analytical models

New Information on the Cranial Anatomy of Acrocanthosaurus atokensis and Its Implications for the Phylogeny of Allosauroidea (Dinosauria: Theropoda)

Allosauroidea has a contentious taxonomic and systematic history. Within this group of theropod dinosaurs, considerable debate has surrounded the phylogenetic position of the large-bodied allosauroid Acrocanthosaurus atokensis from the Lower Cretaceous Antlers Formation of North America. Several prior analyses recover Acrocanthosaurus atokensis as sister taxon to the smaller-bodied Allosaurus fragilis known from North America and Europe, and others nest Acrocanthosaurus atokensis within Carcharodontosauridae, a large-bodied group of allosauroids that attained a cosmopolitan distribution during the Early Cretaceous.Re-evaluation of a well-preserved skull of Acrocanthosaurus atokensis (NCSM 14345) provides new information regarding the palatal complex and inner surfaces of the skull and mandible. Previously inaccessible internal views and articular surfaces of nearly every element of the skull are described. Twenty-four new morphological characters are identified as variable in Allosauroidea, combined with 153 previously published characters, and evaluated for eighteen terminal taxa. Systematic analysis of this dataset recovers a single most parsimonious topology placing Acrocanthosaurus atokensis as a member of Allosauroidea, in agreement with several recent analyses that nest the taxon well within Carcharodontosauridae.A revised diagnosis of Acrocanthosaurus atokensis finds that the species is distinguished by four primary characters, including: presence of a knob on the lateral surangular shelf; enlarged posterior surangular foramen; supraoccipital protruding as a double-boss posterior to the nuchal crest; and pneumatic recess within the medial surface of the quadrate. Furthermore, the recovered phylogeny more closely agrees with the stratigraphic record than hypotheses that place Acrocanthosaurus atokensis as more closely related to Allosaurus fragilis. Fitch optimization of body size is also more consistent with the placement of Acrocanthosaurus atokensis within a clade of larger carcharodontosaurid taxa than with smaller-bodied taxa near the base of Allosauroidea. This placement of Acrocanthosaurus atokensis supports previous hypotheses of a global carcharodontosaurid radiation during the Early Cretaceous

Guidelines for the use and interpretation of assays for monitoring autophagy (4th edition)1.

In 2008, we published the first set of guidelines for standardizing research in autophagy. Since then, this topic has received increasing attention, and many scientists have entered the field. Our knowledge base and relevant new technologies have also been expanding. Thus, it is important to formulate on a regular basis updated guidelines for monitoring autophagy in different organisms. Despite numerous reviews, there continues to be confusion regarding acceptable methods to evaluate autophagy, especially in multicellular eukaryotes. Here, we present a set of guidelines for investigators to select and interpret methods to examine autophagy and related processes, and for reviewers to provide realistic and reasonable critiques of reports that are focused on these processes. These guidelines are not meant to be a dogmatic set of rules, because the appropriateness of any assay largely depends on the question being asked and the system being used. Moreover, no individual assay is perfect for every situation, calling for the use of multiple techniques to properly monitor autophagy in each experimental setting. Finally, several core components of the autophagy machinery have been implicated in distinct autophagic processes (canonical and noncanonical autophagy), implying that genetic approaches to block autophagy should rely on targeting two or more autophagy-related genes that ideally participate in distinct steps of the pathway. Along similar lines, because multiple proteins involved in autophagy also regulate other cellular pathways including apoptosis, not all of them can be used as a specific marker for bona fide autophagic responses. Here, we critically discuss current methods of assessing autophagy and the information they can, or cannot, provide. Our ultimate goal is to encourage intellectual and technical innovation in the field