Theoretical, methodological, and breeding results of the DIAS program in increasing grain production using the example of creation of a soft spring wheat variety - Kazakhstanskaya rannespelaya (origin, characteristics, economic value)

The selection of parental pairs is carried out considering their genealogical trees and the dynamics of the lim factors of the environment, in the creation area, with further hybridization, and subsequent multiple re-sowing of the population. The Kazakhstanskaya rannespelaya variety was created by selection from the F5 hybrid combination of Novosibirskaya 67 × Omskaya 9, the Interdepartmental program DIAS. The breeding record of Novosibirskaya 67 consists of 9 and Omskaya 9 – of 37 varieties, of which 19 are of winter origin. The inheritance of ear productivity, in both varieties, followed the type of complementary epistasis, with incomplete dominance and overdominance, with a large proportion, of recessive genes. In conditions of the Northern Trans-Urals, an early-ripening, yielding line was selected in F5 - Lutescens 1227-8-79, which became the ancestor of the Kazakhstanskaya rannespelaya variety. The variety has successfully passed the State variety testing and is registered in the Qostanay, North Kazakhstan and Kokchetav regions of Kazakhstan, and in the Chelyabinsk region of the Russian Federation, according to the grain quality it is included in the list of strong varieties. In the north of Kazakhstan, Kazakhstanskaya rannespelaya is cultivated on large areas. This variety, as a genetic source, has a good variety-forming ability. With its participation, 6 varieties were created in the Research Institute of the Northern Trans-Urals. Of these, Tyumenskaya 29 and Grenada are registered in 9 and 10 zones of the Russian Federation. It is used in the breeding programs of the Republic of Kazakhstan

Developmental Reaction Norms: the interactions among allometry, ontogeny and plasticity

How micro- and macroevolutionary evolutionary processes produce phenotypic change is without question one of the most intriguing and perplexing issues facing evolutionary biologists. We believe that roadblocks to progress lie A) in the underestimation of the role of the environment, and in particular, that of the interaction of genotypes with environmental factors, and B) in the continuing lack of incorporation of development into the evolutionary synthesis. We propose the integration of genetic, environmental and developmental perspectives on the evolution of the phenotype in the form of the concept of the developmental reaction norm (DRN) The DRN represents the set of multivariate ontogenies that can be produced by a single genotype when it is exposed to environmental variation. It encompasses: 1) the processes that alter the phenotype throughout the ontogenetic trajectory, 2) the recognition that different aspects of the phenotype are (and must be) correlated and 3) the ability of a genotype to produce phenotypes in different environments. This perspective necessitates the explicit study of character expression during development, the evaluation of associations between pairs or groups of characters (e.g., multivariate allometries), and the exploration of reaction norms and phenotypic plasticity. We explicitly extend the concept of the DRN to encompass adjustments made in response to changes in the internal environment as well. Thus, ‘typical’ developmental sequences (e.g., cell fate determination) and plastic responses are simply manifestations of different scales of ‘environmental’ effects along a continuum. We present: (1) a brief conceptual review of three fundamental aspects of the generation and evolution of phenotypes: the changes in the trajectories describing growth and differentiation (ontogeny), the multivariate relationships among characters (allometry), and the effect of the environment (plasticity); (2) a discussion of how these components are merged in the concept of the developmental reaction norm; and (3) a reaction norm perspective of major determinants of phenotypes: epigenesis, selection and constraint