Growth and yield models in Spain Historical overview, contemporary examples and perspectives

In this paper we present a review of forest models developed in Spain in recent years for both timber and non timber production and forest dynamics (regeneration, mortality). Models developed are whole stand, size (diameter) class and individual-tree. The models developed to date have been developed using data from permanent plots, experimental sites and the National Forest Inventory. In this paper we show the different sub-models developed so far and the friendly use software. Main perspectives of forest modeling in Spain are presented

Mixing effects on growth efficiency in mixed pine forests

Increased interest in mixed forests is due to evidence of them being more resource-use efficient and stable forest systems. However, intrinsic and extrinsic factors moderate interspecific species interactions generating different effects in productivity. Here, we explore a method to detect mixing effects in a specific mixture com_x0002_bination (Pinus sylvestris L. and Pinus pinaster Ait.), comparing the growth of mixed stands with that of mono_x0002_cultures. Combined tree and stand-level analyses also helped determine which mixing effects are most important for forest functioning and how changes at one level influence patterns at another level. Data from the Spanish National Forest Inventory were used to compare growth efficiency in mixed and pure stands; we relied on relative stand density indices to determine species-specific site occupancy. This same concept was used to evaluate competition status and inter/intra-specific competition effects as modifiers of potential growth at the tree-level. We observed that growth efficiency in both species increased with the proportion of the complementary species in the stand. At the tree-level, intraspecific competition was higher than interspe_x0002_cific competition in Scots pine tree growth, showing that it had benefited from the mixture. In contrast, mari_x0002_time pine did not show a competitive response to the interspecific interaction, indicating that tree growth was more strongly influenced by the competition structure (size-symmetric and size-asymmetric) than by the spe_x0002_cies of the competitors. Our results highlight the importance of combining stand-level analysis with that of tree-specific competition relationships when studying mixed-species forests

Adapting a model for even-aged Pinus pinea L. stands to complex multi-aged structures

Stone pine (Pinus pinea L.) stands have been usually managed as even-aged stands. Main objectives in management combine two main commercial productions, timber and pine nuts with other social aspects soil protection, recreational use and biodiversity conservation. Multifunctional management, together with the occurrence of successive events affecting regeneration have oriented managers to propose a management schedule based on the establishment and preservation of a low-stocking multi-aged complex structure on favourable locations. Despite the recent effort on modelling growth and yield on even-aged stands of stone pine, no studies focusing on modelling dynamics for uneven-aged stands have yet been developed up to present. In this study, a proposal is presented for adapting and calibrating an existing tree-level model, originally developed for even-aged stands of stone pine (model PINEA2), to multi-aged complex stands. Data from four multi-aged trials and 61 plots from the National Forest Inventory were used to adapt the whole set of functions included in the original model. In our study, four different methods have been proposed to adapt the original equations (1) direct validation and re-parameterization; (2) size class modelling; (3) refit of functions after removing typical even-aged covariates; and (4) multilevel calibration. Adaptation is based on assuming that a multi-aged stand of stone pine can be seen as the sum of independent, smaller, even-aged groups. The low densities of the stands, the early liberation of the most vigorous trees in all size classes and the major importance of root-level competition for water in Mediterranean forests are the main factors explaining these particular dynamics. Results show the suitability of the proposed method, attaining unbiased estimates with a degree of accuracy similar to that achieved in applying the original model to even-aged stands. The adapted model (PINEA_IRR) constitutes a flexible tool for the management and maintenance of stone pine stands, covering a wide range of within stand structural complexity, including forests in transition. © 2008 Elsevier B.V. All rights reserved

Tree ring wood density of Scots pine and European beech lower in mixed-species stands compared with monocultures

Mixed species stands are on the advance in Central Europe and many recently published studies have reported that they can overyield monocultures in terms of volume growth. However, as forest research has in the past been focused on monocultures, knowledge of how mixed-species stands and monocultures compare in terms of wood quality remains limited. Based on five triplets of fully stocked monocultures and mixed stands of Scots pine (Pinus sylvestris L.) and European beech (Fagus sylvatica L.), we analysed whether tree species mixing modifies wood quality and, more precisely, tree ring wood density. From a total of 322 trees we sampled increment cores for the analyses of tree ring width and tree ring wood density using a LIGNOSTATION™. We found that tree ring width of Scots pine was, on average, 14% wider in mixed compared with pure stands. Tree ring width of European beech did not differ between pure and mixed stands. Tree ring wood density was lower in mixed stands compared to pure stands for both Scots pine (−12%) and European beech (−8%). Tree ring wood density and tree ring width were negatively correlated in the case of Scots pine and positively correlated for European beech. When considering tree size and Stand density index, it was found that only tree ring width and mean tree ring wood density of European beech were influenced by stand density. Tree size had a significant effect only on tree ring wood density of European beech. The overall result of larger tree rings of Scots pine in mixed stands and a lower tree ring wood density of both species in mixed stands compared to pure stands was not influenced by stand density or tree size. Based on the measured values of tree ring wood density we conducted estimates of how mixed stands performed in terms of biomass. We found stem biomass to be 8% lower in mixed stands compared to pure stands. Reasons for the revealed differences in tree ring wood density and consequences for, among others, overyielding, carbon storage, and wood quality are discussed