The evolution of coleoid cephalopods and their present biodiversity and ecology

The present status of phylogeny and classification in coleoid cephalopods and the effect of evolution on the present ecology and biodiversity in the group are examined. The basis of knowledge of cephalopod phylogenywas formulated by Naef in the early 1920s, and his ideas and the progress made in the intervening 75 years are investigated. In the process, the roles that transitions between pelagic and benthic habitats played in the evolution of cephalopods are noted, and the possibility is advanced that the most recent “oceanic anoxic event” may have established a time marker for the divergence of some oegopsid families. The major advances since Naef’s work are: 1. The unusual nature of Vampyroteuthis has been recognized; 2. The sister-group relationship between the Neocoleoidea and the Belemnoidea has been established, but requires further confirmation;3. Monophyly has been confirmed for the Decapodiformes (new name), Octopodiformes and Octopoda by molecular and morphological methodologies; 4. The dates of origin of the Belemnoidea, Neocoleoidea, Sepioidea and fossil teuthoids have been extended to considerably earlier times. The major unsolved phylogenetic problems in need of immediate attention are the position of the Myopsida, relationships within the Sepioidea, the identification of the basal nodes within the Oegopsida, and the relationships of most “fossil teuthoids.

Structure of Titan's mid-range magnetic tail: Cassini magnetometer observations during the T9 flyby

We analyze the magnetic structure of Titan's mid-range magnetic tail (5-6 Titan radii downstream from the moon) during Cassini's T9 flyby. Cassini magnetometer (MAG) measurements reveal a well-defined, induced magnetic tail consisting of two lobes and a distinct central current sheet. MAG observations also indicate that Saturn's background magnetic field is close to the moon's orbital plane and that the magnetospheric flow has a significant component in the Saturn-Titan direction. The analysis of MAG data in a coordinate system based on the orientation of the background magnetic field and an estimation of the incoming flow direction suggests that Titan's magnetic tail is extremely asymmetric. An important source of these asymmetries is the connection of the inbound tail lobe and the outbound tail lobe to the dayside and nightside hemispheres of Titan, respectively. Another source could be the perturbations generated by changes in the upstream conditions

Cassini detection of Enceladus' cold water-group plume ionosphere

This study reports direct detection by the Cassini plasma spectrometer of freshly-produced water-group ions (O+, OH+, H2O+, H3O+) and heavier water dimer ions (HxO(2))(+) very close to Enceladus where the plasma begins to emerge from the plume. The data were obtained during two close ( 52 and 25 km) flybys of Enceladus in 2008 and are similar to ion data in cometary comas. The ions are observed in detectors looking in the Cassini ram direction exhibiting energies consistent with the Cassini speed, indicative of a nearly stagnant plasma flow in the plume. North of Enceladus the plasma slowing commences about 4 to 6 Enceladus radii away, while south of Enceladus signatures of the plasma interaction with the plume are detected 22 Enceladus radii away. Citation: Tokar, R. L., R. E. Johnson, M. F. Thomsen, R. J. Wilson, D. T. Young, F. J. Crary, A. J. Coates, G. H. Jones, and C. S. Paty ( 2009), Cassini detection of Enceladus' cold water-group plume ionosphere, Geophys. Res. Lett., 36, L13203, doi:10.1029/2009GL038923

Charged particle environment of Titan during the T9 flyby

The ion measurements of the Cassini Plasma Spectrometer are presented which were acquired on 26 December 2005, during the T9 flyby at Titan. The plasma flow and magnetic field directions in the distant plasma environment of the moon were distinctly different from the other flybys. The near-Titan environment, dominated by ions of Titan origin, had a split signature, each with different ion composition; the first region was dominated by dense, slow, and cold ions in the 16-19 and 28-40 amu mass range, the second region contained only ions with mass 1 and 2, much less dense and less slow. Magnetospheric ions penetrate marginally into region 1, whereas the region-2 ion population is mixed. A detailed analysis has led us to conclude that the first event was due to the crossing of the mantle of Titan, whereas the second one very likely was a wake crossing. The split indicates the non-convexity of the ion-dominated volume around Titan. Both ion distributions are analysed in detail

Discrete classification and electron energy spectra of Titan's varied magnetospheric environment

We analyse combined electron spectra across the dynamic range of both Cassini electron sensors in order to characterise the background plasma environment near Titan for 54 Cassini-Titan encounters as of May 2009. We characterise the encounters into four broad types: Plasma sheet, Lobe-like, Magnetosheath and Bimodal. Despite many encounters occurring close to the magnetopause only two encounters to date were predominantly in the magnetosheath (T32 and T42). Bimodal encounters contain two distinct electron populations, the low energy component of the bi-modal populations is apparently associated with local water group products. Additionally, a hot lobe-like environment is also occasionally observed and is suggestively linked to increased local pick-up. We find that 34 of 54 encounters analysed are associated with one of these groups while the remaining encounters exhibit a combination of these environments. We provide typical electron properties and spectra for each plasma regime and list the encounters appropriate to each. Citation: Rymer, A.M., H. T. Smith, A. Wellbrock, A.J. Coates, and D.T. Young (2009), Discrete classification and electron energy spectra of Titan's varied magnetospheric environment, Geophys. Res. Lett., 36, L15109, doi: 10.1029/2009GL039427

Plasma electrons above Saturn's main rings: CAPS observations

We present observations of thermal ( similar to 0.6 - 100eV) electrons observed near Saturn's main rings during Cassini's Saturn Orbit Insertion (SOI) on 1 July 2004. We find that the intensity of electrons is broadly anticorrelated with the ring optical depth at the magnetic footprint of the field line joining the spacecraft to the rings. We see enhancements corresponding to the Cassini division and Encke gap. We suggest that some of the electrons are generated by photoemission from ring particle surfaces on the illuminated side of the rings, the far side from the spacecraft. Structure in the energy spectrum over the Cassini division and A-ring may be related to photoelectron emission followed by acceleration, or, more likely, due to photoelectron production in the ring atmosphere or ionosphere

Discovery of heavy negative ions in Titan's ionosphere

Titan's ionosphere contains a rich positive ion population including organic molecules. Here, using CAPS electron spectrometer data from sixteen Titan encounters, we reveal the existence of negative ions. These ions, with densities up to similar to 100 cm similar to 3, are in mass groups of 10-30, 30-50, 50-80, 80-110, 110-200 and 200+ amu/charge. During one low encounter, negative ions with mass per charge as high as 10,000 amu/q are seen. Due to their unexpectedly high densities at similar to 950 km altitude, these negative ions must play a key role in the ion chemistry and they may be important in the formation of organic-rich aerosols (tholins) eventually falling to the surface

Cassini observations of the thermal plasma in the vicinity of Saturn's main rings and the F and G rings

The ion mass spectrometer on Cassini detected enhanced ion flux near Saturn's main rings that is consistent with the presence of atomic and molecular oxygen ions in the thermal plasma. The ring "atmosphere'' and "ionosphere'' are likely produced by UV photosputtering of the icy rings and subsequent photoionization of O-2. The identification of the O+ and O-2(+) ions is made using time-of-flight analysis and densities and temperatures are derived from the ion counting data. The ion temperatures over the main rings are a minimum near synchronous orbit and increase with radial distance from Saturn as expected from ion pick up in Saturn's magnetic field. The O-2(+) temperatures provide an estimate of the neutral O-2 temperature over the main rings. The ion mass spectrometer also detected significant O-2(+) outside of the main rings, near the F ring. It is concluded that between the F and G rings, the heavy ion population most likely consists of an admixture of O-2(+) and water group ions O+, OH+, and H2O+

Magnetic signatures of plasma-depleted flux tubes in the Saturnian inner magnetosphere

Initial Cassini observations have revealed evidence for interchanging magnetic flux tubes in the inner Saturnian magnetosphere. Some of the reported flux tubes differ remarkably by their magnetic signatures, having a depressed or enhanced magnetic pressure relative to their surroundings. The ones with stronger fields have been interpreted previously as either outward moving mass-loaded or inward moving plasma-depleted flux tubes based on magnetometer observations only. We use detailed multi-instrumental observations of small and large density depletions in the inner Saturnian magnetosphere from Cassini Rev. A orbit that enable us to discriminate amongst the two previous and opposite interpretations. Our analysis undoubtedly confirms the similar nature of both types of reported interchanging magnetic flux tubes, which are plasma-depleted, whatever their magnetic signatures are. Their different magnetic signature is clearly an effect associated with latitude. These Saturnian plasma-depleted flux tubes ultimately may play a similar role as the Jovian ones