Stingless Bees (Hymenoptera: Apidae: Meliponini) Attracted to Animal Carcasses in the Brazilian Dry Forest and Implications for Forensic Entomology

The association of stingless bees with pig carcasses exposed in a Brazilian Dry Forest area were examined. Modified Shannon traps were settled together to collect these insects during two seasons (dry and rainy). 564 bees were collected, belonging to three tribes and nine species. The majority of bees (75.5%) was collected during the dry season and Partamona seridoensis Pedro & Camargo, 2003 (32.8%) and Trigonisca sp. (20.9%) were the most abundant species. Five decomposition stages were recognized, being the bloated, active and advanced decay the most attractive to the bees. Considerations about seasonal foraging and use of bees in the forensic entomology scope are presented

Scopogonalia echinura Young 1977

Scopogonalia echinura Young, 1977 (Figure 5) Material examined One male and one female: ‘ Goiânia – GO\ 30/ X/2000 \ B. Silva col.’ (DZUP); one male, same data, except: ‘ 06/ XI/ 2000 ’ (DZUP); one male, same data, except: ‘ 20/ XI/ 2000 ’ (DZUP); one female, same data, except: ‘ 13/ XI/ 2000 ’ (DZUP); one male, same data, except: ‘ 27/ XII/ 2000 ’ (DZUP); one male, same data, except: ‘ 06/ XI/ 2001 ’ (DZUP); one male: ‘CPo. [Campo] GRANDE MT [nowadays MS, Mato Grosso do Sul]\ Brasil – XI 1952 \ M. Alvarenga’ (DZUP); one male: ‘ Tabuleiro, Res. Biol. [Reserva Biológica] Guaribas,\ Paraíba, Brasil \ Rede de arrasto\ 06.IX.2010\ Creão & Afonso [Leal]’ (DSEC); one male and one female: ‘ BRASIL: Paraíba \ Mamanguape, REBIO [Reserva Biológica]\ Guaribas. Area 1\ Sweep. 01. Mar. 2012 \ A. H. Leal leg’. (DSEC). Comment Scopogonalia echinura had been previously recorded from Santa Elena de Uairén in south Venezuela (Young 1977) and from Vichada department in east Colombia (Freytag and Sharkey 2002), being here newly recorded from central and northeast Brazil. The original description (Young 1977, fig. 439f) does not mention the pair of tiny dorsolateral dentiform processes at the apex of the aedeagus observed in our studied specimens (Figure 5).Published as part of Leal, Afonso Henrique, Creão-Duarte, Antonio José & Mejdalani, Gabriel, 2016, Taxonomic review of Scopogonalia Young, 1977 (Insecta: Hemiptera: Cicadellidae: Cicadellini) with description of six new species, pp. 1513-1542 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 50 (23) on page 1529, DOI: 10.1080/00222933.2016.1166530, http://zenodo.org/record/520652

Scopogonalia osteiphera Leal & Creao-Duarte 2016, sp. nov.

Scopogonalia osteiphera Leal & Creão-Duarte sp. nov. (Figure 11) Diagnosis Sharpshooters with predominant green colour, transversal bone shaped black macula on anterior third of pronotum, fore wings with some cells and longitudinal veins dark brown to black; aedeagus with an apical and a subapical dentiform process on ventral portion and dorsoapical portion expanded as hood-like structure; paraphyses with rami strongly curved dorsally, with acute apices, embracing aedeagus. Comment The presence of a transversal bone shaped black macula on the pronotum is a symplesiomorphy of S. osteiphera. Although better observed in S. osteiphera, the presence of a hood-like structure on the dorsoapical portion of aedeagus is also symplesiomorphic for this species. The ventral subapical process of the aedeagus is a homoplastic autapomorphy of S. osteiphera, and the apical one is a symplesiomorphy of a large clade in which this species is included (Leal 2014). Type locality. 26.39477° S, 065.30345° W; 788 m; Choromoro, Tucumán, Argentina. Length. Male holotype, 4.6 mm; female paratypes, 5.6–6.1 mm. External morphology Head (Figure 11A), in dorsal view, moderately produced anteriorly, median length of crown about 1/2 interocular width and 1/3 transocular width; anterior margin narrowly rounded in dorsal view; ocelli located behind anterior eye angles, each closer to adjacent anterior eye angle than to median line of crown, situated at pair of slight concavities. Pronotum (Figure 11A) with width approximately equal to transocular width; with pair of concavities near anterior margin; lateral margins slightly convergent anteriorly; posterior margin slightly concave; dorsopleural carinae complete and oblique. Fore wings (Figure 11A) opaque; membrane including all of apical cells, extending forward along costal margin as far as basal half of wing. Hind legs with femoral setal formula 2:1:1; length of first tarsomere greater than combined length of two more distal ones. Coloration Ground colour of crown, anterior third of pronotum and mesonotum greenish-yellow, remainder of dorsum dark green (Figure 11A). Crown with Y-shaped black macula on median portion, with pair of arches at apices of arms of Y-shaped macula (Figure 11A). Pronotum with five black maculae: yellow anterior portion with median transversal one with broadened ends reminding the shape of a bone, and two transversal oval ones on lateral portions; posterior green portion with pair of arched maculae on posterior margin (Figure 11A). Mesonotum with pair of black maculae on basal angles, partially concealed under pronotum, and V-shaped macula on median portion (Figure 11A). Fore wings with some cells dark brown to black and concolourous stripes on and beside veins (Figure 11A). Face with frons greenish-yellow with slight transversal stripes on muscle impressions; remainder of face pale yellow. Lateral lobe of pronotum with black area just below dorsopleural carina; mesosternum black; legs and ventral portion of abdomen pale yellow; tergum of abdomen black. Male genitalia Pygofer (Figure 11B), in lateral view, moderately produced posteriorly; dorsal margin approximately rectilinear, oblique; posteroventral margin broadly convex; apex acute; numerous macrosetae near apex and along margins; slender process arising from ventral margin and extending posterodorsally as far as apex of pygofer. Subgenital plates (Figure 11C) subtriangular, not extending as far posteriorly as apex of pygofer, gradually tapered towards apex; macrosetae uniseriate along outer margin. Styles (Figure 11D) slender, without preapical lobe, not extending as far posteriorly as apex of connective, broadly curved ventrally, reminiscent of arches, with strong angular projection. Connective (Figure 11D) Y-shaped. Aedeagus (Figure 11E), in lateral view, directed posterodorsally, with two dentiform processes on ventral portion: apical and subapical one; dorsoapical portion expanded as hood-like structure. Paraphyses extending below shaft of aedeagus at base; rami strongly curved dorsally, with acute apices, embracing aedeagus. Female genitalia Abdominal sternite VII (Figure 11F) longer than wide, gradually tapered posteriorly; posterior margin with concavity, slight round prominence inside it. Valvulae II (Figure 11G) expanded beyond basal curvature and tapered towards acute apex; ventral margin approximately rectilinear; dorsal margin approximately parallel to ventral margin; preapical prominence distinct; inclined trapezoid teeth on basal and median portions (Figure 11H), becoming triangular towards apex (Figure 11I); first ones with posterior flat area, which becomes gradually smaller towards apex, where it is absent (Figure11J); denticles on all teeth and on apical portion; ventral dentate apical portion greater than dorsal one (Figure11J). Etymology The species epithet, osteiphera (from Greek: osteos, bone + phero, to bear), refers to the bone shaped macula on pronotum. Type material Holotype: male, ‘ ARGENTINA: Tucuman Prov. \ Km 1346 rt 9\ S 26.39477° W 065.30345°\ 788m 20.i.2008 \ D.M. Takiya sweep’ (DZRJ). Paratypes: one female, same data as holotype (DZRJ); one female: ‘ ARGENTINA: Misiones \ Prov. Rt 12 4KmW\ Capiovi 23.i.2008 \ D. M. Takiya sweep’ (DZUP); one female: ‘ ARGENTINA: Salta Prov. \ Palomitas, Km 1535 rt. 9\ S 24.89399° W 064.99311°\ 750m 19.i.2008 \D.M. Takiya sweep’ (DZUP); one female: ‘ ARGENTINA: Salta Prov.\ Los Nogales, Km 1547 rt. 9\ S 24.80173° W 064.99311°\ 800m 19.i.2008 \ D.M. Takiya sweep’ (DZUP). Comparative notes The new species externally resembles S. interruptula, as mentioned above. It differs from this by the paired paraphyses, by the male pygofer (Figure 11B) with acute apex, by the aedeagus (Figure 11E) with two ventral dentiform processes, as in S. subolivacea, and a hood-like structure, as in S. penicula and S. altmanni.Published as part of Leal, Afonso Henrique, Creão-Duarte, Antonio José & Mejdalani, Gabriel, 2016, Taxonomic review of Scopogonalia Young, 1977 (Insecta: Hemiptera: Cicadellidae: Cicadellini) with description of six new species, pp. 1513-1542 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 50 (23) on pages 1538-1540, DOI: 10.1080/00222933.2016.1166530, http://zenodo.org/record/520652

Scopogonalia dolixoura Leal & Creao-Duarte 2016, sp. nov.

Scopogonalia dolixoura Leal & Creão-Duarte sp. nov. (Figure 9) Diagnosis Light brown sharpshooters with head, in dorsal view, slightly produced anteriorly; male pygofer long and with dorsal margin, in lateral view, very concave; aedeagus with three dentiform processes on apical portion: two dorsolateral and one ventroapical. Comment The slightly produced head and the very concave shape of the male pygofer are autapomorphies of S. dolixoura (Leal 2014). Type locality. Parque Estadual da Serra de Santa Bárbara, Pontes e Lacerda, Mato Grosso, Brazil. Length. Male holotype, 6.5 mm. External morphology Head (Figure 9A), in dorsal view, slightly produced anteriorly, median length of crown about 1/3 interocular width and 1/5 transocular width; anterior margin broadly rounded in dorsal view; ocelli located behind anterior eye angles, each closer to adjacent anterior eye angle than to median line of crown, situated at pair of slight concavities. Pronotum (Figure 9A) with width less than transocular width; lateral margins parallel; posterior margin slightly concave; dorsopleural carinae complete and oblique; disc slightly striate, without setae; mesonotum with scutellum not striate. Fore wings (Figure 9A) completely hyaline. Hind legs with femoral setal formula 2:1:1; length of first tarsomere approximately equal to combined length of two more distal ones. Coloration Crown, pronotum and mesonotum light brown, fore wings amber (Figure 9A). Crown with dark brown round maculae on and around ocelli (Figure 9A). Dark brown stripe on lateral body portion, beginning on prothorax right behind eye and extending to tergum of abdomen (Figure 9A). Fore wings with dark brown stripe between radius and cubitus, enforcing the one on abdomen, visible by transparency of wings (Figure 9A). Face and ventral body surface light brown. Male genitalia Pygofer (Figure 9B), in lateral view, strongly produced posteriorly, slender; dorsal margin strongly concave; ventral margin slightly concave; apex obliquely truncate; numerous macrosetae on narrow apical portion and scattered along margins; slender process arising from ventral margin and extending posterodorsally, not attaining apex of pygofer. Subgenital plates (Figure 9C) subtriangular, not extending as far posteriorly as apex of pygofer, gradually tapered towards narrow apical portion; macrosetae multiseriate near base, becoming uniseriate towards apex. Styles (Figure 9D) with preapical lobe, not extending as far posteriorly as apex of connective, twisted, apex obliquely truncate, with strong angular projection; preapical lobe and angular projection directed ventrally. Connective (Figure 9D) Y-shaped. Aedeagus (Figure 9E), in lateral view, directed posteriorly and curved dorsally; apical portion with three small dentiform processes: ventral one and two dorsolateral ones. Paraphyses (Figure 9E) paired, symmetrical, extending below shaft of aedeagus; rami curved dorsally, apices slightly curved towards one another. Female. Unknown. Etymology The species epithet, dolixoura (from Greek: dolixouros, with long tail), refers to the long and slender male pygofer. Type material Holotype: male, ‘Parque [Estadual da Serra] de Santa Bárbara \ Pontes [e] Lacerda-MT [Mato Grosso state]\ Santa Rita-03-07/VII/02’, ‘ 16°02’34”S \ 59°16’22”N [W]\ Cavichioli leg’ (DZUP). Comparative notes Scopogonalia dolixoura externally resembles S. agkistroides, S. altmanni, S. echinura and S. oglobini due to the completely hyaline fore wings and the dark brown colour of the lateral body portions. Its aedeagus (Figure 9E) is similar to that of S. echinura (Figure 5) because both bear three dentiform processes on the apical portion: a ventral one and two dorsolateral ones. However, the aedeagus of S. dolixoura is longer beyond the dorsal curvature than that of S. echinura and its dorsolateral processes are more distant from the apex. It differs from this and the other known species by the long and slender male pygofer with the dorsal margin very concave.Published as part of Leal, Afonso Henrique, Creão-Duarte, Antonio José & Mejdalani, Gabriel, 2016, Taxonomic review of Scopogonalia Young, 1977 (Insecta: Hemiptera: Cicadellidae: Cicadellini) with description of six new species, pp. 1513-1542 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 50 (23) on pages 1535-1536, DOI: 10.1080/00222933.2016.1166530, http://zenodo.org/record/520652

Scopogonalia interruptula

Scopogonalia interruptula (Osborn, 1926) (Figures 3, 4) Cicadella interruptula Osborn, 1926. Scopogonalia interruptula: Young 1977. Diagnosis Green sharpshooters with dark broad longitudinal stripes; aedeagus, in lateral view, directed posterodorsally, almost straight from curvature at base, with pair of dentiform processes on dorsal portion; paraphysis (i.e. with a single ramus) directed posteriorly. Comment The paraphysis (i.e. with a single ramus) is an autapomorphy of S. interruptula (Leal 2014). Type locality. ‘Chapada’, Brazil. Length. Male allotype, 5.5 mm; female holotype, 6.0 mm. External morphology Head (Figure 3) moderately produced anteriorly, median length of crown about 1/2 interocular width and 1/3 transocular width; anterior margin narrowly rounded in dorsal view; ocelli located behind anterior eye angles, each closer to adjacent anterior eye angle than to median line of crown, situated at pair of slight concavities. Pronotum (Figure 3) with width approximately equal to transocular width of crown; lateral margins convergent anteriorly; dorsopleural carinae complete and oblique; disc smooth. Fore wings (Figure 3) opaque; membrane including all of apical cells, extending anteriorly along costal margin as far as basal half of wing. Hind legs with femoral apical setal formula 2:1:1; length of first tarsomere greater than combined length of two more distal ones. Coloration Background colour of crown, anterior third of pronotum, and mesonotum light green; remainder of dorsum dark green (Figure 3B). Crown with dark brown transversal stripe anterior to ocelli, forked at ends, and dark brown transversal stripe along posterior margin with projections attaining ocelli; dark brown line on median line of crown linking both transversal stripes (Figure 3B). Anterior third of pronotum with dark brown transversal stripe forked at ends, with short anterior branches, posterior branches extending posteriorly as far as mesonotum and returning to pronotum (Figure 3B). Mesonotum (Figure 3B) with three dark brown maculae: two on basal angles continuous with stripes of pronotum and median one extending from base to scutellar suture. Fore wings (Figure 3B) with dark brown stripes on and beside longitudinal veins. Face and ventral body parts yellow; pair of dark brown longitudinal stripes on muscle impressions. Male genitalia Pygofer (Figure 4A), in lateral view, moderately produced posteriorly; dorsal margin concave; ventral margin approximately rectilinear; apex narrowly rounded; numerous macrosetae from apical half to apex, scattered basally; slender process arising from ventral margin, extending posteriorly as far as apex of pygofer. Subgenital plates (Figure 4B) subtriangular, not extending posteriorly as far as apex of pygofer, gradually tapered towards narrow apical portion; macrosetae multiseriate near base, becoming uniseriate towards apex. Styles (Figure 4C) slender, without preapical lobe, not extending posteriorly as far as apex of connective; apex obliquely truncate. Connective (Figure 4C) Y-shaped. Aedeagus (Figure 4D), in lateral view, directed posterodorsally, almost straight from curvature at base; apex acute, with pair of dentiform processes on dorsal portion. Paraphysis (Figure 4D) with ramus directed posteriorly, apex acute. Female. Unavailable for dissection. Material examined Female holotype: ‘ Chapada \ Brazil \ Acc. No. 2966’, ‘Nov.’, ‘TYPE’, ‘ Cicadella \ interruptula\ H. O.’ (CMNH). Male allotype: ‘ Chapada \ Brazil \ Acc. No. 2966’, ‘ALLOTYPE’, ‘ Scopogonalia \ interruptula\ (Osb.)\ Young 1971’ (CMNH). Comparative notes Scopogonalia interruptula resembles externally S. paula and S. osteiphera sp. nov. (Figure 11A) by the green colour and the broad dark brown stripes on and beside the longitudinal veins of the fore wings. However, it is different from these and any other species of the genus because of the single paraphysis (Figure 4D), instead of the paraphyses of the other ones.Published as part of Leal, Afonso Henrique, Creão-Duarte, Antonio José & Mejdalani, Gabriel, 2016, Taxonomic review of Scopogonalia Young, 1977 (Insecta: Hemiptera: Cicadellidae: Cicadellini) with description of six new species, pp. 1513-1542 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 50 (23) on pages 1527-1528, DOI: 10.1080/00222933.2016.1166530, http://zenodo.org/record/520652

Scopogonalia altmanni Cavichioli 1986

Scopogonalia altmanni Cavichioli, 1986 Material examined One male and one female: ‘ ARGENTINA: Salta Prov.,\ 10 km N Rosario de la Frontera \ Km 1436 rt. 9, 814m \ S 25º42.546 ʹ W 064º56.623 ʹ \ D. M. Takiya 20.i.2008 sweep’ (DZRJ). Male also with label: ‘ Scopogonalia \ altmani [sic]\ Cavichioli\ D M TAKIYA det. 2007’. Comment Scopogonalia altmanni had been previously recorded from Paraná state, south Brazil (Cavichioli 1986). With the new record, the distribution of the species is expanded towards northwest Argentina, being the first record of the species out of Brazil.Published as part of Leal, Afonso Henrique, Creão-Duarte, Antonio José & Mejdalani, Gabriel, 2016, Taxonomic review of Scopogonalia Young, 1977 (Insecta: Hemiptera: Cicadellidae: Cicadellini) with description of six new species, pp. 1513-1542 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 50 (23) on page 1529, DOI: 10.1080/00222933.2016.1166530, http://zenodo.org/record/520652

Scopogonalia amazonensis Leal & Creao-Duarte 2016, sp. nov.

Scopogonalia amazonensis Leal & Creão-Duarte sp. nov. Diagnosis Green sharpshooters, crown and anterior third of pronotum yellow; aedeagus with two small semi-fused dentiform processes on ventral portion: one subapical and one apical, and pair of wing-shaped processes on dorsoapical portion. Comment The presence of the subapical dentiform process is a homoplastic autapomorphy of S. amazonensis sp. nov. shared with S. subolivacea, S. osteiphera sp. nov. and S. oglobini, whereas the pair of wing-shaped processes of the aedeagus is a homoplastic autapomorphy of S. amazonensis sp. nov. shared with S. subolivacea (Leal 2014). Type locality. Itacoatiara, Amazonas state, Brazil. Length. Male holotype, 5.7 mm. Male paratypes, 5.0– 5.9 mm; female paratypes, 6.0 mm. External morphology Head (Figure 6A) moderately produced anteriorly, median length of crown approximately 1/2 interocular width and 1/3 transocular width; anterior margin narrowly rounded in dorsal view; ocelli located behind anterior eye angles, each closer to adjacent anterior eye angle than to median line of crown, situated at pair of slight concavities. Pronotum (Figure 6A) with width approximately equal to transocular width; lateral margins convergent anteriorly; dorsopleural carinae incomplete, not attaining eyes, oblique. Fore wings (Figure 6A) opaque, membrane including all of apical cells, extending anteriorly along costal margin as far as basal half of wing. Hind legs with length of first tarsomere greater than combined length of two more distal ones. Coloration Background colour of crown, anterior third of pronotum, and mesonotum greenish-yellow, remainder of dorsum green (Figure 6A). Crown with round green maculae on and around ocelli (Figure 6A). Ventral region of body yellow; tergum of abdomen dark brown to black. Male genitalia Pygofer (Figure 6B), in lateral view, moderately produced posteriorly; dorsal margin slightly concave; posterodorsal margin broadly convex; apex acute; process arising from ventral margin and extending posterodorsally as far as apex of pygofer. Subgenital plates (Figure 6C) not extending posteriorly as far as apex of pygofer, gradually tapered towards apex; macrosetae uniseriate along outer margins. Styles (Figure 6D) slender, without preapical lobe, strongly curved. Connective (Figure 6D) Y-shaped. Aedeagus (Figure 6E), in lateral view, directed posterodorsally, with two tiny semi-fused dentiform processes on ventral portion, larger subapical one and smaller apical one, and pair of wing-shaped processes on dorsoapical portion (Figure 6E–F). Paraphyses (Figure 6E) paired, extending below aedeagus, with rami curved dorsally, not attaining aedeagus. Female genitalia Abdominal sternite VII (Figure 6G) longer than wide, lateral margins parallel, tapering near apex; posterior margin with concavity. Valvulae II (Figure 6H) expanded beyond basal curvature and gradually tapered towards acute apex; dorsal margin approximately rectilinear and parallel to ventral margin; preapical prominence distinct; teeth of basal and median portions inclined trapezoid (Figure 11H), becoming triangular towards apex (Figure 11I); first ones with flat posterior area, which becomes gradually smaller towards apex, where it is absent; denticles on all teeth and on apical portion; ventral dentate apical portion greater than dorsal one (Figure 11J). Etymology The species epithet, amazonensis, refers to the Amazon region, where the specimens were found. Type material Holotype: male, ‘Brasil-AM [Amazonas state]/ Itacoatiara \ Madeireira MIL, 024510S \583911W, 29–30.xi.2005 \ Luminosa móvel, J. A.\ Rafael, R. J. Machado &\ A. Silva Filho leg.’ (INPA). Paratypes: three males (INPA) with the same data as holotype; one male and two females: ‘ Rio Branco-AC [Acre state]\ 12-Jan-2004 \ Albuquerque, E.S.’ (DZUP). Comment Apparently, Young (1977, fig. 435) based on specimens of S. amazonensis his interpretation of S. subolivacea from Chanchamayo, Peru. The paratypes from Acre state have the pronotum almost completely yellow, with the posterior part light green. The fore wings also present a less intense tonality of green than the specimens from Amazonas do. However, the comparison of the male genitalia of specimens from both states support the hypothesis that they belong to the same species. Comparative notes This species resembles S. subolivacea, but differs from the latter by the small and semifused dentiform processes of the ventral aedeagal portion (Figure 6E) and by the narrower wing-shaped lateral processes (Figure 6E–F). The rami of the paraphyses are also longer in the new species than in S. subolivacea (Figure 6E).Published as part of Leal, Afonso Henrique, Creão-Duarte, Antonio José & Mejdalani, Gabriel, 2016, Taxonomic review of Scopogonalia Young, 1977 (Insecta: Hemiptera: Cicadellidae: Cicadellini) with description of six new species, pp. 1513-1542 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 50 (23) on pages 1529-1531, DOI: 10.1080/00222933.2016.1166530, http://zenodo.org/record/520652

Lycoderes albinoi Creao-Duarte & Cabral 2017, sp. nov.

<i>Lycoderes albinoi</i> Creão-Duarte & Cabral sp. nov. <p>(Figs. 1–4)</p> <p> <b>Diagnosis</b>. Body dark brown, almost black, except for forewings, with large central hyaline area, and abdomen, yellow; anterior pronotal process vertical, elevated, with suprahumeral horns arising at apex, divergent in frontal view, spatulate, longer than wide, tricarinate; posterior process sickle-shaped, extending from base of anterior process, arched and well separated from scutellum anteriorly, distal half tectiform, tapering to acute apex, extended slightly beyond apex of clavus.</p> <p>Measurements (in millimeters): Male/female. Total length (head to apex of forewings) 5.85/6.06; pronotum length (projection of suprahumeral horns to apex of posterior process) 5.37/5.49; length of forewings 5.36/5,48; width between humeral angles 1.71/1.91; width of head across eyes 1.67/1.77; head height 0.88/0.94; distance between eyes 1.12/1.23; distance between ocelli 0.46/0.50.</p> <p> <b>Description</b>. <b>Holotype male</b>. <b>Coloration</b>. Head, pronotum, ventral surface of thorax and legs dark brown, except for central area of forewings, hyaline, and eyes, ocelli and abdomen, yellow; head, legs and thorax with whitish and waxy pubescence; hind tibiae light brown, and tarsi yellow. <b>Head</b> (Fig. 2). Approximately 1.5x wider than long, eyes hemispherical, ocelli small, above centro-ocular line; postclypeus trilobed in frontal view. <b>Thorax</b>. Pronotum (Fig. 1) densely punctate, punctations coarser on posterior process; short, decumbent setae throughout; median carina weaker in metopidium, percurrent and sharper along posterior process; suprahumeral horns Vshaped in anterior view, spatulate, longer than wide, tri-faceted, delimited by carinae on anterior and posterior margins and on ventral surface from apex to base of posterior process; in lateral view, metopidium slightly curved anteriorly, short, not exceeding suprahumeral horns in length; posterior process sickle-shaped, extending from base of anterior process, sharply curved and well separated from scutellum anteriorly, distal half tectiform, tapering towards acute apex, reaching beyond apex of clavus; forewings (Fig. 1) obliquely truncate at apex; one discoidal cell, five apical cells: fourth cell triangular, not petiolate, and fifth cell pentagonal; two crossveins: <i>r-m</i> fused to branch M1+2 and <i>m-cu</i> to branches M3+4. Legs with tibiae spatulate, metathoracic tibiae with cucullate setae in distal half of row II.</p> <p>Female (Figs. 3,4): similar to male, slightly larger and lighter in color; proximal half of posterior process less sharply curved.</p> <p> Material examined. <b>Holotype male</b>, “ Brasil, BA[HIA], Uma | Fazenda UNACAU | 7-24.X.1986 /D. S. Amo- | rim & C. Vasconcelos ”; “ Mata Atlântica Primária | Armadilha Luminosa ” (DSEC / UFPB). Paratype female with same label data as holotype.</p> <p> <b>Notes on type specimens</b>. Specimens in good state of preservation. The holotype lacks the right suprahumeral horn. The left suprahumeral horn is missing in the female paratype, as well as the tarsi of both metathoracic legs. There is slight variation in the position of the crossvein <i>r-m</i> in the type series, however, in both the holotype and the paratype it is located at the base of M1+2 after the M fork. The crossvein <i>r-m</i> is a little farther from the fork in the holotype’s left forewing, and right after the point of bifurcation of M in the right forewing of the female, which is anomalous in this regard. In all cases, the fourth apical cell in the forewings is distinctly triangular.</p> <p> <b>Remarks.</b> In comparison to other congeneric species, <i>Lycoderes albinoi</i> <b>sp. nov.</b> most closely resembles <i>Lycoderes ancora</i> (Germar) and <i>Lycoderes furcifer</i> Sakakibara. In females of <i>Lycoderes ancora</i>, the suprahumeral horns diverge next to the base of the posterior process, whereas in females of <i>Lycoderes. albinoi</i> <b>sp. nov.</b>, the anterior process is more elongate, so that the suprahumeral horns diverge farther from the base of the posterior process. In males of <i>Lycoderes ancora</i>, the base of the posterior process is wider, reaching the base of the suprahumeral horns, a condition that is not observed in males of <i>Lycoderes albinoi</i> <b>sp. nov.</b> due to the much narrower base of the posterior process. The base of the posterior process is also close to the base of the suprahumeral horns in males and females of <i>Lycoderes furcifer</i>, however, the tip of the posterior process does not reach the apex of the clavus, as it does in <i>Lycoderes albinoi</i> <b>sp. nov.</b> In males of <i>Lycoderes furcifer</i>, the suprahumeral horns are tapered acutely towards the distal tips, and slightly curved ventrally. Contrastingly, the suprahumeral horns are gradually broadened toward the distal half in males of <i>Lycoderes albinoi</i> <b>sp. nov.</b>, forming a dorsal angle in frontal view, then tapering from the distal third to an acute tip.</p> <p> <b>Etymology</b>. This species is dedicated to Dr. Albino Morimasa Sakakibara, for his lifelong work training new systematists focused on auchenorrhynchan Hemiptera, and for his outstanding contributions to treehopper taxonomy.</p>Published as part of <i>Creão-Duarte, Antonio José, Cabral, Valberta Alves & Lourenço, Aline, 2017, A new species of Lycoderes Sakakibara (Hemiptera, Membracidae, Stegaspidinae) from Brazil in Zootaxa 4281 (1)</i>, DOI: 10.11646/zootaxa.4281.1.5, <a href="http://zenodo.org/record/815778">http://zenodo.org/record/815778</a&gt