Fig. 2 in Review of the fritillary species systematically close to Melitaea lutko Evans, 1932 (Lepidoptera: Nymphalidae) with analysis of their geographic distribution and interrelations with host plants

Fig. 2. Adults of Melitaea lutko Evans, 1932 and M. timandra binaluduca subsp. nov. A–H. UPS. I–P. UNS. A, I. M. lutko, ♂, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. B, J. M. timandra binaluduca subsp. nov., ♂, Iran, S Khorasan Prov., 35 km N Birjant t. C, K. M. timandra binaluduca subsp. nov., ♂, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. D, L. M. timandra binaluduca subsp. nov., ♂, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. E, M. M. lutko, ♀, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F, N. M. timandra binaluduca subsp. nov., ♂, Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. G, O. M. timandra binaluduca subsp. nov., ♀, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H, P. M. timandra binaluduca subsp. nov., ♀, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.Published as part of Kolesnichenko, Kirill A. & Kotlobay, Anatoly A., 2022, Review of the fritillary species systematically close to Melitaea lutko Evans, 1932 (Lepidoptera: Nymphalidae) with analysis of their geographic distribution and interrelations with host plants, pp. 1-60 in European Journal of Taxonomy 830 (1) on page 7, DOI: 10.5852/ejt.2022.830.1865, http://zenodo.org/record/683984

PLATE 6. VI in Early Stages and Notes on the Biology of Melitaea gina Higgins, 1941 (Lepidoptera: Nymphalidae) from Northern Zagros Mts (Iran) with the Special Remarks on Taxonomy

PLATE 6. VI instar caterpillar and head of IV–VI instar caterpillars of Melitaea gina Higgins, 1941 (a—view from above; b—lateral view): 1—VI instar caterpillar; 2—head of IV instar caterpillars; 3—head of V instar caterpillar; 4—head of VI instar caterpillar.Published as part of <i>Kolesnichenko, Kirill A. & Kotlobay, Anatoly A., 2023, Early Stages and Notes on the Biology of Melitaea gina Higgins, 1941 (Lepidoptera: Nymphalidae) from Northern Zagros Mts (Iran) with the Special Remarks on Taxonomy, pp. 223-238 in Zootaxa 5369 (2)</i> on page 231, DOI: 10.11646/zootaxa.5369.2.3, <a href="http://zenodo.org/record/10150324">http://zenodo.org/record/10150324</a&gt

PLATE 9 in Early Stages and Notes on the Biology of Melitaea gina Higgins, 1941 (Lepidoptera: Nymphalidae) from Northern Zagros Mts (Iran) with the Special Remarks on Taxonomy

PLATE 9. Holotype of M. gina safavides Gross & Ebert, 1975. 1—Holotype UPS, 2—holotype UNS, 3—labels.Published as part of <i>Kolesnichenko, Kirill A. & Kotlobay, Anatoly A., 2023, Early Stages and Notes on the Biology of Melitaea gina Higgins, 1941 (Lepidoptera: Nymphalidae) from Northern Zagros Mts (Iran) with the Special Remarks on Taxonomy, pp. 223-238 in Zootaxa 5369 (2)</i> on page 235, DOI: 10.11646/zootaxa.5369.2.3, <a href="http://zenodo.org/record/10150324">http://zenodo.org/record/10150324</a&gt

Early Stages and Notes on the Biology of Melitaea gina Higgins, 1941 (Lepidoptera: Nymphalidae) from Northern Zagros Mts (Iran) with the Special Remarks on Taxonomy

Kolesnichenko, Kirill A., Kotlobay, Anatoly A. (2023): Early Stages and Notes on the Biology of Melitaea gina Higgins, 1941 (Lepidoptera: Nymphalidae) from Northern Zagros Mts (Iran) with the Special Remarks on Taxonomy. Zootaxa 5369 (2): 223-238, DOI: 10.11646/zootaxa.5369.2.3, URL: https://www.mapress.com/zt/article/download/zootaxa.5369.2.3/5223

Fig. 19. Male genitalia and harpe. A–C in Review of the fritillary species systematically close to Melitaea lutko Evans, 1932 (Lepidoptera: Nymphalidae) with analysis of their geographic distribution and interrelations with host plants

Fig. 19. Male genitalia and harpe. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. Turkmenistan, Sary-Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H. Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. I. Afghanistan, Band-e-Amir, Hazarajat.Published as part of Kolesnichenko, Kirill A. & Kotlobay, Anatoly A., 2022, Review of the fritillary species systematically close to Melitaea lutko Evans, 1932 (Lepidoptera: Nymphalidae) with analysis of their geographic distribution and interrelations with host plants, pp. 1-60 in European Journal of Taxonomy 830 (1) on page 29, DOI: 10.5852/ejt.2022.830.1865, http://zenodo.org/record/683984

PLATE 5. II–V in Early Stages and Notes on the Biology of Melitaea gina Higgins, 1941 (Lepidoptera: Nymphalidae) from Northern Zagros Mts (Iran) with the Special Remarks on Taxonomy

PLATE 5. II–V instar caterpillars of Melitaea gina Higgins, 1941 (a—view from above; b—latera lview): 1—II instar caterpillar; 2—III instar caterpillar; 3—IV instar caterpillar; 4—V instar caterpillar.Published as part of <i>Kolesnichenko, Kirill A. & Kotlobay, Anatoly A., 2023, Early Stages and Notes on the Biology of Melitaea gina Higgins, 1941 (Lepidoptera: Nymphalidae) from Northern Zagros Mts (Iran) with the Special Remarks on Taxonomy, pp. 223-238 in Zootaxa 5369 (2)</i> on page 230, DOI: 10.11646/zootaxa.5369.2.3, <a href="http://zenodo.org/record/10150324">http://zenodo.org/record/10150324</a&gt

PLATE 2 in Early Stages and Notes on the Biology of Melitaea gina Higgins, 1941 (Lepidoptera: Nymphalidae) from Northern Zagros Mts (Iran) with the Special Remarks on Taxonomy

PLATE 2. Imago of M. gina Higgins, 1941. 1–6—imago of the late (summer) generation; 7–12—imago of the first (spring) generation. 1—male UPS, Iran, Northern Zagros Mts, Hamadan Province, Nahavand County, Shaban Rural District, 2 km south of Tazehnab-e Olya village, 2100 m., 20.07.2022; 2—female UPS, the same locality; 3—female UPS, ex pupa; 4—male UNS, the same locality; 5—female UNS, the same locality; 6—female UNS, ex pupa; 7—male UPS, Iran, Yasuj area, Khafr v, vicinity, 2200–2500 m., 10.05.2011; 8—male UPS, Iran, Yasuj area, Vezg pass, 2100 m., 15.05.2013; 9—female UPS, Iran, North Fars, 10 km NW Ardakan, 2500 m., 12.05.2008; 10—male UNS, Iran, Yasuj area, Khafr v, vicinity, 2200–2500 m., 10.05.2011; 11—male UNS, Iran, Yasuj area, Vezg pass, 2100 m., 15.05.2013; 12—female UNS, Iran, North Fars, 10 km NW Ardakan, 2500 m., 12.05.2008.Published as part of <i>Kolesnichenko, Kirill A. & Kotlobay, Anatoly A., 2023, Early Stages and Notes on the Biology of Melitaea gina Higgins, 1941 (Lepidoptera: Nymphalidae) from Northern Zagros Mts (Iran) with the Special Remarks on Taxonomy, pp. 223-238 in Zootaxa 5369 (2)</i> on page 226, DOI: 10.11646/zootaxa.5369.2.3, <a href="http://zenodo.org/record/10150324">http://zenodo.org/record/10150324</a&gt

Fig. 18. Male genitalia and harpe. A–C in Review of the fritillary species systematically close to Melitaea lutko Evans, 1932 (Lepidoptera: Nymphalidae) with analysis of their geographic distribution and interrelations with host plants

Fig. 18. Male genitalia and harpe. A–C. Melitaea shahvarica sp. nov. D–E. M. lutko Evans, 1932. F–I. M. mimetica Higgins, 1940. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D–E. Pakistan, Chitral, Chaghbini, CGNP [Chitral Gol National Park], alt. 2700 m. F–G. Afghanistan, Bamian Prov., Punjub Distr., 10 km. NE Varas v., alt. 2400 m. H–I. Afghanistan, Bamian Prov., Panjub Distr., 10 km. NE Varas vil., alt. 2400 m.Published as part of <i>Kolesnichenko, Kirill A. & Kotlobay, Anatoly A., 2022, Review of the fritillary species systematically close to Melitaea lutko Evans, 1932 (Lepidoptera: Nymphalidae) with analysis of their geographic distribution and interrelations with host plants, pp. 1-60 in European Journal of Taxonomy 830 (1)</i> on page 28, DOI: 10.5852/ejt.2022.830.1865, <a href="http://zenodo.org/record/6839847">http://zenodo.org/record/6839847</a&gt

PLATE 7 in Early Stages and Notes on the Biology of Melitaea gina Higgins, 1941 (Lepidoptera: Nymphalidae) from Northern Zagros Mts (Iran) with the Special Remarks on Taxonomy

PLATE 7. Pupa of Melitaea gina Higgins, 1941: 1—front view; 2—lateral view; 3—back view; 4—cremaster lateral view; 5—cremaster view from above.Published as part of <i>Kolesnichenko, Kirill A. & Kotlobay, Anatoly A., 2023, Early Stages and Notes on the Biology of Melitaea gina Higgins, 1941 (Lepidoptera: Nymphalidae) from Northern Zagros Mts (Iran) with the Special Remarks on Taxonomy, pp. 223-238 in Zootaxa 5369 (2)</i> on page 233, DOI: 10.11646/zootaxa.5369.2.3, <a href="http://zenodo.org/record/10150324">http://zenodo.org/record/10150324</a&gt

Review of the fritillary species systematically close to Melitaea lutko Evans, 1932 (Lepidoptera: Nymphalidae) with analysis of their geographic distribution and interrelations with host plants

This review considers a fascinating, from a zoogeographical viewpoint, group of closely related species: Melitaea lutko Evans, 1932, M. timandra Coutsis & van Oorschot, 2014, M. mimetica Higgins, 1940 stat. rev. and M. shahvarica sp. nov. It is a taxonomical and geographical review of these species, and data on the biology of M. shahvarica sp. nov. and nominate subspecies of M. timandra are discussed. A new species, M. shahvarica sp. nov. from Shahvar Mt. (Iran), and a new subspecies, M. timandra binaludica subsp. nov. from Kuh-e-Binalud Mts (Iran), are described. The specific structure of the group given in previous publications is critically evaluated. Hypotheses about a possible phylogenesis of the study group are provided