Andricus istvani Tavakoli & Melika & Sadeghi & Pénzes & Assareh & Atkinson & Bechtold & Mikó & Zargaran & Aligolizade & Barimani & Bihari & Pirozi & Fülöp & Somogyi & Challis & Preuss & Nicholls & Stone 2008, new species

Andricus istvani Melika, new species Figs 2–17 Type material. HOLOTYPE female: IRAN, Lorestan, Ghelaei, V.2005. Q. brantii. Leg. M. Tavakoli. Lor 177. PARATYPES: 1 female and 14 males with the same labels as the holotype. The holotype and 2 male paratypes are deposited in HNHM; 5 male paratypes in RIFR; 1 female and 7 male paratypes in the collection of SPL. Other material examined. 28 males and 12 galls labelled as " Iran, Kermanshah, Pave, V.2005. Q. brantii. Leg. M. Tavakoli ". Etymology. In honour of Dr. István Mikó, our friend and researcher of the Systematic Parasitoid Laboratory, Plant Protection and Soil Conservation Service of County Vas, Tanakajd, Hungary. Diagnosis. The only known Western Palaearctic sexual Andricus which in the forewing in both, females and males, lacks cilia on margins is A. crispator Tschek, which is the most closely related species. However, in A. crispator OOL 3.4–3.6 times as long as the length of the lateral ocellus; the transfacial distance 1.8–2.0 times as long as the height of eye; the pedicel 1.5 times as long as broad; F10 1.5 times as long as F9; R 1 in the forewing on a short distance running along the wing margin, the areolet triangular, distinct; the median mesoscutal line absent; the mesoscutellum posteriorly elongated into a blunt spine-like tip; the mesopleuron, except coriaceous speculum, transversely striate, while in A. istvani, new species, OOL only 2.3 times as long as the length of the lateral ocellus; the transfacial distance only 1.2 times as long as the height of eye; the pedicel 2.0 times as long as broad; F10 2.5 times as long as F9; R 1 in the forewing do not reaching wing margin, the areolet absent; the median mesoscutal line in a form of a short triangle; the mesoscutellum posteriorly rounded; the mesopleuron, including speculum, uniformly transversely delicately striate. Description. FEMALE (holotype). Body and antennae entirely and uniformly light brown; eyes dark grey; legs slightly lighter; wings with distinct brown veins. Head delicately coriaceous, with very few short white setae, 2.1 times as broad as long from above, slightly broader than high in front view and slightly broader than mesosoma. Gena delicately coriaceous, slightly broadened behind eye, visible in front view behind eye. Malar space coriaceous, with striae radiating from clypeus and extending to 2/3 of malar space length, 0.4 times as long as height of eye. POL 1.2 times as long as OOL; OOL 2.3 times as long as length of lateral ocellus and 1.5 times as long as LOL. Transfacial distance 1.2 times as long as height of eye and 1.6 times as long as height of lower face (distance between antennal rim and ventral margin of clypeus); diameter of antennal torulus 1.9 times as long as distance between them, distance between torulus and eye margin slightly longer than diameter of torulus. Inner margins of eyes very slightly converning ventrally. Lower face coriaceous, with strongly elevated median area. Clypeus trapezoid, coriaceous, nearly as broad as high, with distinct deep anterior tentorial pits, deeply impressed along distinct epistomal sulcus and clypeo-pleurostomal line, ventrally broadly emarginated, medially not incised. Frons, vertex and occiput uniformly coriaceous. Antenna with 10 flagellomeres (in female paratype with 11 flagellomeres, an indistinct suture present between F11 and F10); nearly as long as body length; pedicel 2.0 times as long as broad; F1 1.6 times as long as pedicel and 1.5 times as long as F2; F2 very slightly longer than F3, F3–F9 nearly equal in length; F10 2.5 times as long as F9; placodeal sensilla distinct on F5–F10, absent on F1–F4. Mesosoma convex, 1.2 times as long as high in lateral view, with very few short white setae. Pronotum coriaceous, with some delicate subparallel interrupted wrinkles along mesopleuron. Mesoscutum delicately coriaceous, broader than long (width measured across the basis of tegulae); notauli complete, wellimpressed in all length; median mesoscutal line in a form of short triangle; parapsidal lines narrow but distinct; anterior parallel lines distinct, very short, extending to 1/4 of mesoscutum length. Mesoscutellum 1.5 times shorter than mesoscutum, slightly elongated, uniformly dull rugose, posteriorly rounded, strongly overhanging metanotum. Scutellar foveae subquadrate, slightly broader than high, well-delimited around, with shining, smooth bottom, separated by a narrow but distinct central median carina. Mesopleuron, including speculum, uniformly transversely delicately striate; mesopleural triangle shining, with some delicate wrinkles; acetabular carina delimiting a broad dull rugose area laterally. Metapleural sulcus reaching mesopleuron slightly above the half of its height; preaxilla and lateral axillar area coriaceous, with delicate wrinkles; axillar carina narrow, with some longitudinal delicate striae; axillula ovate, smooth, shining, with sparse white setae; subaxillular bar smooth, shining, in the most posterior end 1.5 times as high as height of metanotal trough. Metascutellum very delicately coriaceous, nearly as high as height of smooth, shining ventral impressed area; metanotal trough smooth, shining, with sparse white setae. Lateral propodeal carinae subparallel, very slightly curved outwards in the posterior 1/3, central propodeal area shining, smooth; lateral propodeal area uniformly coriaceous, with white setae; nucha very short, with delicate longitudinal sulci. Forewing with distinct brown veins, margin without cilia; radial cell 4.2 times as long as broad, Rs and R1 not reaching wing margin, areolet absent, Rs+M nearly reaching basalis in its lower half. Tarsal claws with strong basal lobe. Metasoma longer than head+mesosoma, nearly as long as high in lateral view, all tergites and hypopygium without micropunctures; prominent part of ventral spine of hypopygium slender, 5.6 times as long as broad in ventral view, with few short setae, which not extending beyond apex of spine. Body length 1.5–1.7 mm (holotype 1.6 mm). MALE. 1.4–1.7 mm. Similar to female but mesoscutum, mesoscutellum, central propodeal area, nucha and metasoma dorsally very dark brown to black (in all 42 examined males); frons and vertex with more delicate sculpture; compound eyes slightly larger, antenna with 12 flagellomeres, slightly longer than body length, F1 slightly curved, apically swollen, 1.3 times as long as F2, placodeal sensilla on all flagellomeres. Gall (Figs 14–17). An integral leaf gall, develops on both sides of the leaf (usually on terminal leaves of the twig), 10–20 mm long, irregularly shaped; multilocular, containing many cells. Young galls are fleshy and pale green to yellowish, and become brown and remain soft as they mature. Galls usually occur gregariously, resulting in a noticeable deformation of the leaf. Biology. Only the sexual generation is known from galls on Quercus brantii Lindl. Young galls become apparent from the beginning of May; adult wasps emerge from mid-May. Rare species. Distribution. Iran, Lorestan (Ghelaei) and Kermanshah provinces (Pave).Published as part of Tavakoli, M., Melika, G., Sadeghi, S. E., Pénzes, Z., Assareh, M. A., Atkinson, R., Bechtold, M., Mikó, I., Zargaran, M. R., Aligolizade, D., Barimani, H., Bihari, P., Pirozi, F., Fülöp, D., Somogyi, K., Challis, R., Preuss, S., Nicholls, J. & Stone, G. N., 2008, New species of oak gallwaps from Iran (Hymenoptera: Cynipidae: Cynipini), pp. 1-64 in Zootaxa 1699 (1) on pages 5-6, DOI: 10.11646/zootaxa.1699.1.1, http://zenodo.org/record/510717

Andricus megatruncicolus Tavakoli & Melika & Sadeghi & Pénzes & Assareh & Atkinson & Bechtold & Mikó & Zargaran & Aligolizade & Barimani & Bihari & Pirozi & Fülöp & Somogyi & Challis & Preuss & Nicholls & Stone 2008, new species

Andricus megatruncicolus Melika, new species Figs 87–104 Type material. HOLOTYPE female: IRAN, Lorestan, Ghelaei, Q. infectoria, em. XI-XII.2005. leg. M. Tavakoli. Lor. 157. PARATYPES: 24 females: 14 females with the same labels as the holotype; 4 females labelled as " IRAN, Kordestan, Marivan, Q. infectoria. 2003. leg. M. Tavakoli, Lor 123", and 6 females labelled as " IRAN, Lorestan, Ghelaei, Q. infectoria. mid-summer, 2003. leg. M. Tavakoli, Lor 77". The holotype and 5 female paratypes in HNHM; 5 female paratypes in RIFR; 10 female paratypes in SPL; 4 female paratypes in NHML. Other material examined. 36 females labelled as " IRAN, Kordestan, Marivan, Q. infectoria. 2003. leg. M. Tavakoli, Lor123" and 12 females labelled as " IRAN, Kermanshah, Gahvareh, Q. infectoria. mid-summer, leg. M. Tavakoli, Lor95". Etymology. The species name megatruncicolus reflects the close similarity of the adult and asexual generation gall of this species to Andricus truncicolus (Giraud), and the larger size of the gall. Diagnosis. Most closely resembles Andricus truncicolus in which the body is predominantly blackish brown; the head is more or less rounded in front view; OOL 2.5 times as long as length of lateral ocellus and 2.1 times as long as LOL; the diameter of the antennal torulus about 3.7 times as large as the distance between them; the clypeus only slightly broader or as broad as high; F1 1.25 times as long as F2, F2 only slightly longer than F3; the mesoscutum coriaceous anteriorly, reticulate posteriorly, especially in the internotauli area; the median mesoscutal line distinct, in a form of short triangle; the mesoscutellum rounded, very slightly longer than broad, with more delicate sculpture towards the centre of scutellar disk; scutellar foveae nearly rounded, as high as broad, separated by a very narrow, delicately longitudinally striate central carina; anteriorly scutellar foveae nearly reaching one another; the mesopleuron and metapleuron with some distinct punctures; lateral propodeal carinae slightly curved outwards posteriorly; the radial cell of the forewing 4.8–5.0 times as long as broad, while in A. megatruncicolus, new species, the body predominantly is reddish brown; the head is trapezoid in front view; OOL 2.8 times as long as length of lateral ocellus and 2.8 times as long as LOL; diameter of antennal torulus only 2.5 times as large as the distance between them; the clypeus at least 2.0 times as broad as high; F1=F2, F2 1.3 times as long as F3; the mesoscutum uniformly delicately coriaceous, with some distinct punctures in the anterior half; the median mesoscutal line absent; the mesoscutellum broader than long, with more dull rugose sculpture along sides of the mesoscutellum and more delicate ont he disk center; scutellar foveae transverse, much broader than high; separated by a broad delicately reticulate central carina; the mesopleuron and metapleuron without punctures; lateral propodeal carinae curved outwards in the mid-height; the radial cell of the forewing only 4.0 times as long as broad. Description. ASEXUAL FEMALE (holotype). Body predominantly reddish brown. Head entirely reddish brown; pronotum black, mesoscutum with extensive black marks; scutellar foveae, mesopleural triangle, metanotum black. Legs reddish brown, except black tibiae and tarsi. Wings with distinct brown veins. Body with dense white setae. Head coriaceous, trapezoid in front view, with dense white setae; 2.0 times as broad as long from above; 1.5 times as broad as high in front view. Gena coriaceous, strongly broadened behind eye, the most broadened part is on the mid-height of eye, well-visible in front view behind eye, gena nearly as broad as cross diameter of eye. Malar space coriaceous, 0.4 times as long as height of eye, with distinct striae radiating from clypeus and extending to half distance to eye. POL equal OOL; OOL 2.8 times as long as length of lateral ocellus and 2.8 times as long as LOL. Transfacial distance 1.2 times as long as height of eye; diameter of antennal torulus 2.5 times as large as distance between them, distance between torulus and eye margin slightly longer than diameter of torulus. Inner margins of eyes parallel. Lower face delicately coriaceous, with relatively strongly elevated delicately coriaceous median area, without striae radiating from clypeus. Clypeus rectangular, tip with white setae, with coriaceous elevated central area, 2.0 times as broad as high, with distinct anterior tentorial pits, impressed along distinct epistomal sulcus, clypeo-pleurostomal line distinct, ventrally widely emarginated and incised medially. Frons, vertex and occiput uniformly delicately coriaceous. Antenna with 12 flagellomeres, slightly longer than head+mesosoma; pedicel 1.6 times as long as broad; F1=F2, F2 1.3 times as long as F3; subsequent flagellomeres shorter; F12 slightly longer than F11; placodeal sensilla on F4–F12. Mesosoma convex, slightly longer than high in lateral view, with dense white setae. Pronotum uniformly coriaceous, with dense white setae along anterior edge. Mesoscutum uniformly delicately coriaceous, with some distinct punctures in the anterior half. Notauli complete, well-impressed in all length, slightly broader and converging posteriorly, smooth, shiny; median mesoscutal line absent; parapsidal lines and anterior parallel lines distinct, broad, extending to half of mesoscutum length. Mesoscutellum 1.8 times shorter than mesoscutum, broader than long, with dull rugose sculpture along sides and more delicate towards the disk center; slightly overhanging metanotum. Scutellar foveae transverse, much broader than high, with smooth, shiny deep bottom, posteriorly delimited by sculpture, without setae; separated by a broad delicately reticulate central carina. Mesopleuron including speculum smooth, without punctures; mesopleural triangle uniformly delicately coriaceous, with dense setae. Metapleural sulcus reaching mesopleuron slightly below half of its height; axillula ovate, delicately coriaceous, with dense white setae; subaxillular bar smooth, shining, in the most posterior end higher than height of metanotal trough. Metascutellum uniformly coriaceous, 3.0 times as high as height of ventral impressed area; metanotal trough delicately coriaceous, shiny, with dense white setae; ventral impressed area shiny, with some longitudinal weak wrinkles. Lateral propodeal carinae curved outwards in the mid-height, uniformly broad, with sparse setae prolong it length; central propodeal area with numerous weak wrinkles, without setae; lateral propodeal area coriaceous, with some wrinkles and dense white setae. Forewing much longer than body, margin with short cilia; radial cell 4.0 times as long as broad, areolet distinct; Rs+M extending to 2/3 of distance between areolet and basalis, projecting into its lower half. Metasoma nearly as long as head+mesosoma, longer than high in lateral view; all tergites without micropunctures, with dense white setae laterally, which absent dorsally; prominent part of ventral spine of hypopygium 6.6 times as long as broad, with sparse white setae, apical setae short, not extending beyond apex of spine. Body length 4.2–5.3 mm. Gall (Figs 99–104). Develops from accessory buds on branches. Solitary, sometimes in groups, 2–5 galls forming a cluster. The gall is spherical, 5–15 mm in diameter, broadly attached, surface patterned, resembling tortoiseshell. Apex covered in regular, radiating plates, giving a shield-like appearance. Initially whitish, greenish, later becoming brown and bark-coloured, but usually the greyish dusty layer staying on the surface of the gall even after the emerging of wasps. Inner chamber elliptical, monolocular, located at the base of the gall. Biology. Only asexual females are known from galls on Quercus infectoria Olivier. Comments. The extreme rarity of purely asexual lifecycles in oak gallwasps (Stone et al. 2002) suggests that A. megatruncicolus p robably has a sexual generation. The clade of Andricus species that includes A. megatruncicolus all have lifecycles involving alternation between oaks in the sections Quercus (for the asexual generation) and Cerris (for the sexual generation). The sexual generation galls in this clade are commonly small and inconspicuous bud or catkin galls (Melika et al. 2000; Cook et al. 2002; Rokas et al. 2003b), and it is probable that the same is true for this new species. Possible hosts from the Cerris oak section in the region include Quercus libani Olivier and Q. brantii Lindl. (= Q. persica Jaub. et Spach). Distribution. Iran, Kordestan (Marivan), Lorestan (Ghelaei) and Kermanshah (Gahvareh) provinces; Turkey, southern Anatolia (Beybesli) and north-eastern Anatolia (Erzurum).Published as part of Tavakoli, M., Melika, G., Sadeghi, S. E., Pénzes, Z., Assareh, M. A., Atkinson, R., Bechtold, M., Mikó, I., Zargaran, M. R., Aligolizade, D., Barimani, H., Bihari, P., Pirozi, F., Fülöp, D., Somogyi, K., Challis, R., Preuss, S., Nicholls, J. & Stone, G. N., 2008, New species of oak gallwaps from Iran (Hymenoptera: Cynipidae: Cynipini), pp. 1-64 in Zootaxa 1699 (1) on pages 14-16, DOI: 10.11646/zootaxa.1699.1.1, http://zenodo.org/record/510717

Andricus csokai Melika & Tavakoli 2008, new species

Andricus csokai Melika & Tavakoli, new species Figs 51–67 Type material. HOLOTYPE female: IRAN, Kermanshah, Gahvareh and Javanrod, X-XII.2005. Q. infectoria. leg. M. Tavakoli. Lor 149. PARATYPES: 39 females with the same labels as the holotype. The holotype and 5 female paratypes in HNHM; 5 female paratypes in RIFR; 24 female paratypes in SPL; 5 female paratypes in NHML. Etymology. In recognition of the continuing contribution of Dr. György Csóka (Forest Research Institute, Hungary) to research on oak gallwasps. Diagnosis. Most closely resembles Andricus quercusradicis (Fabricius) in which POL 1.5 times as long as OOL; OOL 1.4 times as long as the length of the lateral ocellus and 1.4 times as long as LOL; placodeal sensilla on F3–F12; the median mesoscutal line in a form of a small but distinct triangle; the mesoscutellum delicately coriaceous, posteriorly rounded and medially incised; scutellar foveae are well-delimited around; the mesopleuron including speculum, uniformly delicately coriaceous to alutaceous; the upper half of the mesopleural triangle nearly smooth, ventrally delicately coriaceous with some weak wrinkles and dense setae; lateral propodeal carinae subparallel, broader anteriorly, slightly curved outwards in the most posterior part; the central propodeal area with few delicate irregular wrinkles; metasomal tergite 2 without micropunctures; the prominent part of the ventral spine of the hypopygium 4.5–5.0 times as long as broad while in A. csokai, new species, POL 2.3 times as long as OOL; OOL only slightly longer than the length of the lateral ocellus and equal to LOL; placodeal sensilla on F4–F12; the median mesoscutal line extending to 1/4 of the mesoscutum length; the mesoscutellum rugose, posteriorly is rounded and not incised centrally; scutellar foveae posteriorly delimited by the rugulose sculpture only; the mesopleuron with some delicate longitudinal interrupted striae, while the speculum is reticulate, with distinct punctures; the mesopleural triangle uniformly shining, alutaceous; lateral propodeal carinae subparallel, uniformly broad, slightly curved inwards in the middle; the central propodeal area delicately uniformly coriaceous; metasomal tergite 2 with micropunctures in the posterior 1/3 of the tergite; the prominent part of the ventral spine of the hypopygium at least 6.0 times as long as broad. Description. ASEXUAL FEMALE (holotype). Body predominantly dark reddish brown, with some extensive dark patches. Head posteriorly, between antennal toruli, central triangular area below median ocellus, central elevated area of lower face, clypeus and narrow stripes of malar space along attachment of mandibles always black; mandibles and antennae reddish brown; compound eyes and ocelli silvery or light brown. Scutum between anterior parallel lines and along parapsidal lines black; scutellar fovea, axillar complex, mesopleuron (entirely or only speculum), mesosoma ventrally, metascutellum and central propodeal area black. Legs uniformly reddish brown, slightly lighter than body. Metasoma reddish brown, except metasomal tergite 2, which dorsally always black. Wings with distinct brown veins. Body with relatively dense white setae. Head coriaceous, with relatively sparse short white setae, more dense on lower face; 2.2 times as broad as long from above; 1.3 times as broad as high in front view and as broad as mesosoma. Gena delicately coriaceous, broadened behind eye, well-visible in front view behind eye, slightly narrower than cross diameter of eye, measuring behind eye. Malar space delicately coriaceous, 0.45 times as long as height of eye, with delicate striae radiating from clypeus and extending to half of its length. POL 2.3 times as long as OOL; OOL slightly longer than length of lateral ocellus and equal to LOL. Transfacial distance slightly longer than height of eye and 1.3 times as long as height of lower face (distance between antennal rim and ventral margin of clypeus); diameter of antennal torulus 2.4 times as large as distance between them; distance between torulus and eye margin equal to diameter of torulus. Inner margins of eyes slightly divirging ventrally. Lower face coriaceous, with very slightly elevated delicately coriaceous median area, without striae radiating from clypeus, with dense white setae. Clypeus coriaceous, impressed along sides and elevated centrally, slightly broader than high, with distinct anterior tentorial pits, deeply impressed along distinct epistomal sulcus, clypeo-pleurostomal line distinct, ventrally widely emarginated and only very slightly incised medially. Frons, especially the black triangular area below median ocellus reticulate, with distinct large punctures; vertex with some distinct punctures, interocellar area with some strong rugae; occiput uniformly delicately coriaceous. Antenna with 12 flagellomeres, slightly longer than head+mesosoma; pedicel 1.6 times as long as broad; F1 2.0 times as long as pedicel and only very slightly longer than F2, F2 1.3 times as long as F3; F4–F6 nearly equal in length, slightly shorter than F3; F7 and subsequent flagellomeres much shorter, nearly equal in length, F12 1.8 times as long as F11, placodeal sensilla on F4–F12. Mesosoma convex, slightly longer than high in lateral view, with relatively dense white setae. Pronotum delicately coriaceous, with some delicate wrinkles along antero-lateral edge only, with dense white setae, especially along anterior edge. Mesoscutum reticulate, punctures are dense, distance between punctures less than the diameter of a puncture. Notauli complete, well-impressed in all length, converging and not broadened posteriorly; median mesoscutal line extending to 1/4 of mesoscutum length; anterior parallel and parapsidal lines extending to 1/2 of mesoscutum length. Mesoscutellum coriaceous, posteriorly rounded and centrally not incised, 1.7 times shorter than mesoscutum, rounded, nearly as long as broad or slightly longer, slightly overhanging metanotum. Scutellar foveae transversely ovate, posteriorly delimited by rugulose sculpture, smooth, shining, without setae; separated by a distinct, delicately coriaceous central carina. Mesopleuron with some delicate longitudinal interrupted striae; speculum reticulate, with distinct punctures; mesopleural triangle uniformly shiny, alutaceous. Metapleural sulcus reaching mesopleuron slightly above the half of its height, lower part of metapleuron, delimited by metapleural sulcus shiny, smooth; preaxilla delicately coriaceous; axillula ovate, shining, coriaceous, with very dense white setae; subaxillular bar coriaceous, shining, in the most posterior end 2.0 times higher than height of metanotal trough; ventral bar of metanotal trough shiny, at least 2.0 times narrower than height of metanotal trough measuring above propodeal spiracle. Metascutellum uniformly delicately coriaceous, nearly as high as height of smooth, shiny delicately longitudinally wrinkled ventral impressed area; metanotal trough coriaceous, shiny, with relatively sparse white setae. Lateral propodeal carinae subparallel, uniformly broad, slightly curved inwards in the middle; central propodeal area delicately coriaceous, without setae; lateral propodeal area delicately uniformly coriaceous, with dense white setae; nucha short, with irregular wrinkles. Tarsal claws with distinct basal lobe. Forewing much longer than body, with distinct brown veins, margin with cilia; radial cell 3.7 times as long as broad, areolet distinct, Rs+M extending to 2/3 of distance between areolet and basalis. Metasoma nearly as long as head+mesosoma, higher than long in lateral view; metasomal tergite 2 with large patch of dense white setae antero-laterally, with micropunctures in the posterior 1/3, subsequent tergites and hypopygium without setae, with dense micropunctures; prominent part of ventral spine of hypopygium short, 6.0 times as long as broad, with few short white setae, apical setae short, not extending beyond apex of spine. Body length 4.1–4.8 mm. Gall (Figs 62–67). Galls are on roots which are close to the surface or on branches at ground level and up to 15–35 cm below the ground surface. Often found on the roots of fallen trees with many regrowth shoots. Commonly in groups. Varying in size from a walnut to an onion; thick walled; usually globular, multilocular, 2.5–5.0 cm in diameter. Initially fleshy, soon becomes woody. When young the gall is light brown or reddish, darkening (brownish) with age, after wasps emerged, the old gall turns black. Biology. Only the asexual generation is known from galls on Quercus infectoria Olivier. Rare species. The gall becomes apparent at the beginning of summer; adult wasps emerge at the beginning of November. Lack of information on the phylogenetic placement of this species precludes prediction of a likely host for the sexual generation. Distribution. Iran, known from Kermanshah province only (Gahvareh and Javanrod).Published as part of Tavakoli, M., Melika, G., Sadeghi, S. E., Pénzes, Z., Assareh, M. A., Atkinson, R., Bechtold, M., Mikó, I., Zargaran, M. R., Aligolizade, D., Barimani, H., Bihari, P., Pirozi, F., Fülöp, D., Somogyi, K., Challis, R., Preuss, S., Nicholls, J. & Stone, G. N., 2008, New species of oak gallwaps from Iran (Hymenoptera: Cynipidae: Cynipini), pp. 1-64 in Zootaxa 1699 (1) on pages 10-12, DOI: 10.11646/zootaxa.1699.1.1, http://zenodo.org/record/510717