The type specimen of Debeya (Dewalquea) haldemiana rediscovered. -Acta Mus

Abstract. The rediscovered specimen MGUWr 7536p, one of the syntypes of Dewalquea haldemiana, is described. It is selected as the lectotype of Debeya (Dewalquea) haldemiana (DEBEY ex SAPORTA et MARION 1873) HALAMSKI 2013 and of Debeya (Dewalquea) haldemiana var. haldemiana. It is the most complete and best preserved known specimen of the species. It allows supplementation of previous description of the following characters: brochidodromous secondary venation; acuminate leaflet apices; variation of the petiolule length from 0 (leaflets subsessile) to 30 mm; ramified tertiary venation. Other species considered up to now to belong to the same subgenus possess percurrent tertiary venation; a doubt is therefore expressed about the validity of the present circumscription of Debeya (Dewalquea). Additonally, validation of the previously published name Debeya (Dewalquea) haldemiana var. angustifolia (HOSIUS et VON DER MARCK 1880) HALAMSKI comb. nov. is presented. n Angiospermae, Debeya, palaeobotany, taxonomy

Eressella, a new uncinuloid brachiopod genus from the Middle Devonian of Europe and Africa

Eressella, a new genus of rhynchonellide brachiopods belonging to the superfamily Uncinuloidea Rzhonsnitskaya, 1956, is described with Rhynchonella coronata Kayser, 1871, as the type and only species. It is characterised by a dorsibiconvex profile with a resupinate ventral valve, costae rounded posteriorly, but acute and developing ventrally directed spur-like protuberances anteriorly, small dental cavities, cardinal process multilobed posteriorly and massive anteriorly, and the presence of a septalium and thick dorsal median septum. Given the present state of flux in the systematics of the superfamily, it is conventionally placed within the family Uncinulidae Rzhonsnitskaya, 1956, although similarities with the subfamily Betterbergiinae Savage, 1996 and the family Eucharitinidae Sartenaer, 2015 are also noted. Eressella coronata, hitherto the only representative of the genus, is known from the Eifel Hills (Eifelian, mainly middle Eifelian), from central Poland (especially from the Eifelian of the Holy Cross Mts.), and from the Moroccan Anti-Atlas (late Eifelian to early Givetian, details uncertain)

Oxfordian brachiopods from the Saïda and Frenda mountains (Tlemcenian Domain, north-western Algeria)

Five brachiopod species are reported from two middle to upper Oxfordian (Late Jurassic) outcrops, situated in the Saïda and Frenda mountains (Tlemcenian Domain, north-western Algeria) and belonging to the heterochronous (Callovian to Oxfordian, locally to the Kimmeridgian) Argiles de Saïda Formation (Saïda Clay Formation). The upper Oxfordian (probably Dichotomoceras bifurcatus Zone) outcrop A yielded Dictyothyris kurri and Loboidothyridoidea indet. The middle Oxfordian (Liosphinctes plicatilis Zone, Cardioceras vertebrale Subzone) outcrop B yielded Monticlarella rollieri, Karadagithyris boullierae sp. n., and Zittelina sp.; this is the first report of the last-mentioned genus from Africa. Karadagithyris boullierae sp. n. is a link between previously known Bajocian to Bathonian (Callovian?) Karadagithyris s.s. and Tithonian to Lower Cretaceous species, formerly segregated as Svaljavithyris; the latter is considered herein as synonym of Karadagithyris. It is characterised by a plano-uniplicate anterior commissure and a suberect beak. The adult loop of Zittelina is confirmed as bilacunar (kingeniform) and not diploform (campagiform). The bulk of the assemblages comprises small and either smooth or finely ornamented species and thus represents a low-energy environment. The lack of modern revisions of the reported species in their type areas is the reason why only Dictyothyris kurri may be used as an index species for the middle to late Oxfordian

Middle Devonian brachiopods from the southern Maїder (eastern Anti-Atlas, Morocco)

Sixty-two Middle Devonian brachiopod species are described on the basis of >1300 specimens from the Taboumakhlof Formation (and subordinately probably also from the top of the El Otfal Formation) at Madne el Mrakib (middle to late Eifelian or early Givetian?), Aferdou el Mrakib (late Eifelian to middle Givetian), and Guelb el Maharch (early Givetian) on the southern edge of the MaÎder Syncline (eastern Anti-Atlas, Mo- rocco). Representatives of Craniida (2 taxa), Strophomenida (5), Productida (2), Orthotetida (2), Orthida (5), Pentamerida (5), Rhynchonellida (9), Atrypida (14), Athyridida (7), Spiriferida (9), and Spiriferinida (2) are present. The fauna is dominated quantitatively by the Atrypida (24% of taxa, about the half of specimens, and the commonest species Atryparia dispersa making up about 13% of the material); the Rhynchonellida (16% of taxa, about one-sixth of specimens) are the second largest order. A new genus of the family Pugnacidae (order Rhynchonellida), Paulinaerhynchia, is proposed with the type species P. paulinae gen. et sp. nov. from Maharch; it is closest to Pugnax, from which it differs in distinct costation, lack of a dorsal septum and septalium, and rudimentary dental plates. Desquamatia (D.) deserti sp. nov., a large and finely costate representative of the genus, is described also from Maharch. Antirhynchonella and Glosshypothyridina are reported for the first time or con- firmed to be present in the Givetian. Forty (possibly up to 46) species (71 or possibly up to 82% of the taxa identified at the species level) are present also in either Eifel (Germany) or the Holy Cross Mountains (Poland). Such a high ratio of species in common attests to unconstrained faunal exchanges among benthic faunas between the northern and southern shores of the Variscan Sea during the Middle Devonian. This favours the palaeogeo- graphic hypothesis of a narrow Variscan Sea

Latest Famennian brachiopods from Kowala, Holy Cross Mountains, Poland

Latest Famennian (UD−VI, “Strunian”) brachiopod fauna from Kowala (Kielce Region, Holy Cross Mountains, Poland) consists of eighteen species within 6 orders, eleven of them reported in open nomenclature. Characteristic taxa include: Schellwienella pauli, Aulacella interlineata, Sphenospira julii, Novaplatirostrum sauerlandense, Hadyrhyncha sp., Cleiothyridina struniensis. New morphological details of Schellwienella pauli, Sphenospira julii, and Aulacella interlineata are provided. The described latest Famennian brachiopod fauna is distinctly richer than that from underlying upper Famennian deposits (11 species within 4 orders). Majority of species from Kowala seem to have been adapted to deep water settings and/or poor nutrient availability. The stratigraphic separation between Planovatirostrum in the UD−III to UD−V and Novaplatirostrum in the UD−VI observed in Sauerland and in Thuringia is valid also in the Holy Cross Mountains. This is the first comprehensive report of a relatively diversified latest Famennian brachiopod fauna from surface outcrops of Poland

Brachiopods and stratigraphy of the Upper Devonian (Frasnian) succession of the Radlin Syncline (Holy Cross Mountains, Poland)

The lower part of the Frasnian succession in the Radlin Syncline (Kielce–Łagów Synclinorium, southern region of the Holy Cross Mountains), in the two studied successions: Józefka at Górno and (for the first time) Radlin, consists of the rhythmic marly Szydłówek Beds, the fossil-rich limestones of the Wietrznia Beds (locally) and the atypically developed, calcareous Kostomłoty Beds. The carbon isotope chemostratigraphic pattern overall corresponds well to the global Early–Middle Frasnian biogeochemical perturbation, even if the major punctata positive excursion is only fragmentarily recorded in the Kostomłoty intrashelf basin. Two brachiopod assemblages are abundantly represented in both sections: the Phlogoiderhynchus polonicus Assemblage, typical of the Szydłówek Beds, and the Biernatella lentiformis Assemblage, limited to the middle part of the Wietrznia Beds. Both are highly dominated by the index species. Twenty nine lower Frasnian brachiopod species (Craniida – 1 species, Strophomenida – 1, Productida – 2, Protorthida – 1, Orthida – 5, Pentamerida – 1, Rhynchonellida – 4, Atrypida – 4, Athyridida – 3, Spiriferida – 4, Spiriferinida – 3) are described from the Szydłówek and Wietrznia Beds. Seven new species are introduced: Skenidioides cretus Halamski sp. nov., Biernatium minus Baliński sp. nov., Monelasmina montisjosephi Baliński sp. nov., Atryparia (Costatrypa) agricolae Halamski and Baliński sp. nov., Davidsonia enmerkaris Halamski sp. nov., Leptathyris gornensis Baliński sp. nov., and Echinocoelia parva Baliński sp. nov. Davidsonia enmerkaris Halamski sp. nov. is intermediate between Davidsonia Bouchard-Chantereaux, 1849 and Rugodavidsonia Copper, 1996 and is the youngest known representative of the suborder Davidsonioidea Copper, 1996. Skenidioides cretus Halamski sp. nov. is the last representative of the genus. Statistical investigation of a large sample of Spinatrypina (Exatrypa) explanata did not confirm the existence of two dimorphic forms, coarse- and fine-ribbed. The high-diversity Biernatella lentiformis Assemblage is quite dissimilar to coeval brachiopod assemblages described heretofore from the Holy Cross Mountains region. It is interpreted as consisting of mostly parautochthonous dwellers of deep-slope muddy habitats and a local, occasionally storm-agitated, intra-basin brachiopod-crinoid-coral shoal. The fauna was adapted probably to cooler and nutrient-poor waters during an initial phase of the severe carbon cycle perturbation